MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Wed 04 February 2026
Publication Lopes-Lima et al. (2024): summary
Full Citation
Lopes-Lima, M., J. Geist, S. Egg, L. Beran, A. Bikashvili et al. 2024. Integrative phylogenetic, phylogeographic and morphological characterisation of the Unio crassus species complex reveals cryptic diversity with important conservation implications. Molecular Phylogenetics and Evolution 195. 108046 (27 pp.). Available electronically.
Attributed Nominal Species
Unio courtilieri ‘Hattemann’ of Authors = Unio crassus
‘Hattemann’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 132 [26].
‘Hattemann’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 256 [18].
‘Hattmann’ Germain, 1931. Faune de France 22: 724, 727.
Lopes-Lima et al., 2024. MPE 195: suppl. append. 2, p. 3.
(error for Unio courtillieri Hattemann, 1859)
Nodularia douglasiae ‘Griffith & Pidgeon’ of Authors = Nodularia dualobtusa
‘Griffith & Pidgeon’ of Authors.
Cha et al., 2018. mtDNA B 3(2): 946. [in part]
Fukata & Iigo, 2019. mtDNA B 4(2): 3488. [in part]
Choi et al., 2020. Sci. Repts. 10: 5 [as ‘Nodularia douglasiae Clade A’]. [in part]
Sayenko et al., 2021. Systemat. Biodiv. 19: 57, Table S2. [in part]
Mabuchi et al., 2021. mtDNA B 6(4): 1536. [in part]
Kim et al., 2021. mtDNA B 6(1): 80. [in part]
Lopes-Lima et al., 2024. MPE 195: Table S2. [in part]
Dai et al., 2024. Invert. Syst. 38(6): fig. 3, Table S1. [in part]
Wu et al., 2024. Invert. Syst. 38, IS24014: Table S1. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S2. [in part]
(misidentified reference to Unio douglasiae Griffith & Pidgeon, 1833)
Taxonomic Opinions
There are 257 species taxonomic opinions attributed to this publication.
Unio bruguierianus Bourguignat, 1853
+ Unio orientalis Bourguignat, 1852
+ Unio prusii Bourguignat, 1856
+ Unio schwerzenbachi Bourguignat, 1856
+ Unio vescoi Bourguignat, 1856
+ Unio kotschyi Küster, 1862
+ Unio turcicus ‘Parreiss in sched.’ Küster, 1862
+ Unio desectus f. pursacensis Sturany, 1902
+ Unio ancyrensis Kobelt, 1913
+ Unio ancyrensis var. louisei Kobelt, 1913
+ Unio crassus ‘Philipsson’ of Authors
Unio crassus Philipsson in Retzius, 1788
+ Unio musivus Spengler, 1793
+ Unio ater Nilsson, 1822
+ Unio rubens Menke, 1828
+ Unio rugatus Menke, 1828
+ Unio drouetii Dupuy, 1849
+ Unio courtillieri Hattemann, 1859
+ Unio batavus var. minimus de Joannis, 1859
+ Unio batavus var. baraceus de Joannis, 1859
+ Unio lambottei Malzine, 1867
+ Unio crassus var. maximus Kobelt, 1872
64 more...
Unio desectus Westerlund in Westerlund & Blanc, 1879
+ Unio bruguierianus ‘Bourguignat’ of Authors
Unio gontierii Bourguignat, 1856
+ Unio colchicus Drouët, 1881
+ Unio sieversi Drouët, 1881
+ Unio mingrelicus Drouët, 1881
+ Crassiana mengrelica Kantor et al., 2010
+ Unio raddei Drouët, 1881
+ Unio stevenianus Drouët, 1881
+ Unio stepanoffi Drouët, 1881
+ Unio raddei var. koutaisiana Kobelt, 1886
+ Unio gregorii Kobelt, 1912
+ Unio mingrelicus var. sobriewskii Rosén, 1925
2 more...
Unio mardinensis Lea, 1865
+ Unio kullethensis Lea, 1865
+ Unio orphaensis Lea, 1865
+ Unio rothi var. komarowi O. Boettger, 1880
+ Unio araxenus Drouët, 1881
+ Unio subbatavus Kobelt, 1913
+ Unio (sec. Lymnium) pseudonymus Simpson, 1900
+ Unio armeniacus Kobelt, 1912
+ Unio modiola Preston, 1912
+ Unio specialis Kobelt, 1913
+ Unio (orfaensis var. ?) ehrmanni Kobelt, 1913
7 more...
Unio nanus Lamarck, 1819
+ Mya ovata Studer in Coxe, 1789
+ Mya pictorum Coxe, 1789
+ Unio inflatus Studer, 1820
+ Unio rostratus Studer, 1820
+ Unio sinuatus Studer, 1820
+ Unio planus Studer, 1820
+ Unio dilatatus Studer, 1820
+ Unio ovatus Studer, 1820
+ Unio riparia Pfeiffer, 1821
+ Unio cytherea Küster, 1833
92 more...
Unio sesirmensis Kobelt, 1913
Unio tumidiformis da Silva e Castro, 1885
+ Unio batavus Morelet, 1845
+ Unio eupygus da Silva e Castro, 1885
+ Unio macropygus da Silva e Castro, 1885
+ Unio sadoicus da Silva e Castro, 1885
+ Unio baeticus Kobelt, 1888
+ Unio callipygus Drouët, 1893
+ Unio conimbricus Kobelt, 1893
Unio vicarius Westerlund in Westerlund & Blanc, 1879
+ Unio byzantinus ‘Parreyss, in sched.’ Drouët, 1879
+ Unio heldreichi O. Boettger, 1886
+ Unio thiesseae Drouët, 1892
+ Unio crassus ‘Philipsson’ of Authors
+ Unio bruguierianus ‘Bourguignat’ of Authors
