Bivalvia | Unionida | Hyrioidea

family HYRIIDAE Swainson, 1840: cladomics

2 genera, 2 subfamilies

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 93 (132 total)
number of subtaxa (rank: subfamily) = 4
analyzed in 105 cladograms in 55 publications
maximum number of terminals in an analysis = 28 (mean = 4.07, median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 1.53, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 61
maximum number of included terminals = 28, (mean = 6.28 median = 5)
maximum number of subtaxa (subfamily) = 2 (mean = 1.92, median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyriidae cladogram distances
PCoA Ordination

NJ Tree of Hyriidae cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
28 28 34 2 604 0 0 0 
  +----------------------------------- Westralunio **           
  |                                                          
  |                             +----- Diplodon_suavidicus__2 **
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini **               
  |  |                          +--+                         
  |  |                             +-- Castaliini **            
  |  |                                                       
  |  |                             +-- Hyridella_depressa **    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta **     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1 **
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis **    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus **       
        |                                                    
        |                          +-- Cucumerunio **           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis **   
        |  |                                                 
        +--+  +----------------------- Echyridella **           
           |  |                                              
           |  |              +-------- Velesunio__1 **          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 ** 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus **    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga **     
                 |                                           
                 |  +----------------- Velesunio__3 **          
                 |  |                                        
                 +--+  +-------------- Microdontia **           
                    |  |                                     
                    +--+           +-- Lortiella **             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi **      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 ** 
                          +--+                               
                             |  +----- Velesunio__2 **          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 ** 
                                +--+                         
                                   +-- Alathyria_pertexta__1 **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
19 19 20 2 1 0 0 22.5 
                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio **           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus **    
        +--+                                  
           |  +-------- Microdontia **           
           +--+                               
              |  +----- Lortiella **             
              +--+                            
                 |  +-- Alathyria **             
                 +--+                         
                    +-- Velesunio_angasi **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
19 19 21 2 1 0 0 25.5 
                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio **           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus **    
           +--+                                  
              |  +-------- Alathyria **             
              +--+                               
                 |  +----- Velesunio_angasi **      
                 +--+                            
                    |  +-- Lortiella **             
                    +--+                         
                       +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
14 14 12 2 186 0 0 39 
           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae **          
  |                              
--+     +----- Alathyria_jacksoni **
  |  +--+                        
  |  |  |  +-- Alathyria_profuga ** 
  |  |  +--+                     
  +--+     +-- Velesunio **         
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
14 14 12 2 184 0 0 41 
           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae **          
  |                              
--+     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
14 14 9 2 186 0 0 38 
  +----------- Echyridella **       
  |                              
  +----------- Westralunio **       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
14 14 12 2 6 0 0 37 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
10 10 8 2 4 0 0 31.5 
  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
10 10 7 2 26 0 0 29 
  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
9 9 7 2 37 0 0 32.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
9 9 7 2 38 0 0 32.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
9 9 7 2 37 0 0 32.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
9 9 7 2 21 0 0 38 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
9 9 7 2 129 0 0 38 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 2 38 0 0 32.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
9 9 7 2 129 0 0 38 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
9 9 7 2 21 0 0 38 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
8 8 9 2 1 0 0 35 
     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae **     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
7 7 5 2 1 0 0 37 
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae **     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iTtTbiriSoTroTroSiD
7 7 6 1 0 0 0 29 
1

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
6 6 5 2 5 0 0 36.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 2 36 0 0 33 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
5 5 2 2 26 0 0 36 
     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio **     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella **     
     |                 
     +-- Rhipidodontini **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 2 29 0 0 33 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5 5 3 2 136 0 0 33 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
5 5 3 2 26 0 0 33 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iTtTbiriSoTroTroSiD
5 5 3 1 0 0 0 31 
1

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
5 5 3 2 26 0 0 33 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
5 5 1 2 26 0 0 35 
  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella **     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 2 26 0 0 33.5 
     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 2 36 0 0 35 
        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae **
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 2 0 0 32 
  +----- Westralunio **
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyriinae **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 36 0 0 34 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
4 4 2 2 32 0 0 34 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 14 0 0 31.5 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 2 0 0 0 34 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
4 4 2 2 32 0 0 34 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 2 13 0 0 34.5 
  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 13 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
3 3 0 2 12 0 0 34 
  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
3 3 0 2 12 0 0 34 
  +-- Hyriini **       
  |                 
--+-- Velesunioninae **
  |                 
  +-- Hyridellini **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 1 28 0 0 33 
1

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 2 65 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iTtTbiriSoTroTroSiD
3 3 1 1 0 0 0 33 
1

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 25 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 2 12 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 2 25 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
3 3 1 2 26 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 12 0 0 35 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 2 31 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 2 88 0 0 35 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 31 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 2 65 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 2 88 0 0 35 
  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 2 65 0 0 33 
  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 0 0 -- 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 2 0 2 120 0 0 -- 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 24 0 0 -- 
1

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 45 0 0 -- 
  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Velesunio_ambiguus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 0 0 -- 
-- Velesunio_ambiguus **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Echyridella_menziesii **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Alathyria_jacksoni **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Echyridella_menziesii **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Echyridella_menziesii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Echyridella_menziesii **

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Velesunio_ambiguus **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Echyridella_menziesii **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Echyridella_menziesii **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Echyridella_menziesii **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Velesunio_ambiguus **