MUSSELpdb | tribe Pseudodontini Frierson, 1927 cladomics

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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe PSEUDODONTINI Frierson, 1927: cladomics
2 subtribes

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 51 (52 total)
number of subtaxa (rank: subtribe) = 2
analyzed in 97 cladograms in 59 publications
maximum number of terminals in an analysis = 44 (mean = 8.74, median = 4)
maximum number of subtaxa (subtribe) = 2 (mean = 1.48, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 87
maximum number of included terminals = 44, (mean = 9.63 median = 5)
maximum number of subtaxa (subtribe) = 2 (mean = 1.54, median = 2)
proportion of cladograms that support basic monophyly = 0.99

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 37
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 43 (mean = 13.84, median = 11)
maximum number of subtaxa (subtribe) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
10	10	8	2	36	25	8	0

        +----- Ctenodesmini     
  +-----+                       
  |     |  +-- Rectidentini     
  |     +--+                    
  |        +-- Contradentini    
--+                             
  |     +----- Caudiculatus     
  |  +--+                       
  |  |  |  +-- Lamprotulini     
  |  |  +--+                    
  +--+     +-- Gonideini        
     |                          
     |  +----- Schepmaniini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2025 [Publ 4446], fig. 4

tT	bi	riS	oT	roT	roS	iD
44	44	43	2	0	0	0	41

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

tT	bi	riS	oT	roT	roS	iD
43	43	42	2	7	7	3	40

        +-- Lamprotulini     
  +-----+                    
  |     +-- Gonideini        
--+                          
  |  +----- Schepmaniini     
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S2

tT	bi	riS	oT	roT	roS	iD
41	41	39	2	0	0	0	36

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
40	40	41	2	18	18	7	37

        +-------- Chamberlainiini  
  +-----+                          
  |     |  +----- Ctenodesmini     
  |     +--+                       
  |        |  +-- Rectidentini     
--+        +--+                    
  |           +-- Contradentini    
  |                                
  |  +----------- Lamprotulini     
  +--+                             
     |  +-------- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. S1

tT	bi	riS	oT	roT	roS	iD
39	39	37	2	0	0	0	35

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2023 [Publ 4312], fig. 2

tT	bi	riS	oT	roT	roS	iD
39	39	38	2	0	0	0	34

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
30	30	29	2	52	38	4	25

        +-- Indopseudodontina **
  +-----+                    
  |     +-- Pseudodontina **    
  |                          
--+     +-- Gonideini        
  |  +--+                    
  |  |  +-- Lamprotulini     
  +--+                       
     |  +-- Rectidentini     
     +--+                    
        +-- Contradentini

Konopleva et al., 2021 [Publ 4213], fig. 1

tT	bi	riS	oT	roT	roS	iD
23	23	21	2	4	4	2	20

  +----- Lamprotula       
  |                       
  +----- Leguminaia       
  |                       
--+----- Potomida         
  |                       
  +----- Gonidea          
  |                       
  |  +-- Indopseudodontina **
  +--+                    
     +-- Pseudodontina **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
22	22	19	2	14	14	5	22

           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
22	22	20	2	14	14	5	22

           +-- Contradentini    
  +--------+                    
  |        +-- Rectidentini     
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
21	21	20	2	97	51	5	22

        +-- Rectidentini             
  +-----+                            
  |     +-- Contradentini            
  |                                  
  |     +-- Indopseudodontina **        
  |  +--+                            
  |  |  +-- Pseudodontina **            
--+  |                               
  |  |  +-- Sinosolenaia_iridinea    
  |  |  |                            
  |  |  +-- Sinosolenaia_recognita__1
  |  |  |                            
  |  |  +-- Sinosolenaia_carinata    
  |  |  |                            
  +--+  +-- Leguminaia               
     |  |                            
     |  +-- Microcondylaea           
     |  |                            
     |  +-- Gonidea                  
     |  |                            
     |  +-- Ptychorhynchus           
     |  |                            
     |  +-- Parvasolenaia            
     |  |                            
     +--+-- Lamprotula_leaii         
        |                            
        +-- Lamprotula               
        |                            
        +-- Lamprotula_caveata       
        |                            
        +-- Potomida                 
        |                            
        +-- Pronodularia             
        |                            
        +-- Sinohyriopsis            
        |                            
        +-- Chamberlainia            
        |                            
        +-- Sinosolenaia_recognita__2
        |                            
        +-- Sinosolenaia_oleivora

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
21	21	20	2	91	51	5	22

           +-- Rectidentini     
  +--------+                    
  |        +-- Contradentini    
  |                             
--+        +-- Indopseudodontina **
  |  +-----+                    
  |  |     +-- Pseudodontina **    
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Chamberlainiini  
        +--+                    
           +-- Lamprotulini

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	28	26	5	19

                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
  |                    +-- Rectidentini     
--+  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  |  |              +----- Inversidens      
  +--+  +-----------+                       
     |  |           |  +-- Chamberlainiini  
     |  |           +--+                    
     |  |              +-- Lamprotulini     
     +--+                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	59	14	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
17	17	15	2	59	14	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
17	17	14	2	28	26	5	18

                       +-- Indopseudodontina **
  +--------------------+                    
  |                    +-- Pseudodontina **    
  |                                         
--+                    +-- Rectidentini     
  |  +-----------------+                    
  |  |                 +-- Contradentini    
  |  |                                      
  +--+  +----------------- Inversidens      
     |  |                                   
     |  |              +-- Chamberlainiini  
     |  +--------------+                    
     +--+              +-- Lamprotulini     
        |                                   
        |  +-------------- Sinosolenaia     
        |  |                                
        |  |        +----- Gonidea          
        +--+  +-----+                       
           |  |     |  +-- Leguminaia       
           |  |     +--+                    
           |  |        +-- Microcondylaea   
           +--+                             
              |        +-- Koreosolenaia    
              |  +-----+                    
              |  |     +-- Parvasolenaia    
              +--+                          
                 |  +----- Postolata        
                 +--+                       
                    |  +-- Ptychorhynchus   
                    +--+                    
                       +-- Obovalis

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

tT	bi	riS	oT	roT	roS	iD
16	16	14	1	0	0	0	19

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
15	15	15	2	186	41	5	19

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
15	15	13	2	110	36	5	18

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
15	15	16	2	184	40	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
15	15	9	2	186	41	5	17

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
14	14	13	2	125	35	5	19

              +----- Gonideini                 
  +-----------+                                
  |           |  +-- Chamberlainiini           
  |           +--+                             
  |              +-- Lamprotulini              
--+                                            
  |  +-------------- Pilsbryoconcha **            
  |  |                                         
  |  |  +----------- Indopseudodontina **         
  +--+  |                                      
     |  |        +-- Thaiconcha **                
     |  +--------+                             
     |  |        +-- Bineurus **                  
     +--+                                      
        |     +----- Pseudodon_cambodjensis **    
        |  +--+                                
        |  |  |  +-- Pseudodon_mekongi **         
        |  |  +--+                             
        |  |     +-- Pseudodon_vondembuschianus **
        +--+                                   
           |     +-- Pseudodon_inoscularis **     
           |  +--+                             
           |  |  +-- Sundadontina_tumida **       
           +--+                                
              |  +-- Nyeinchanconcha **           
              +--+                             
                 +-- Sundadontina_cumingii **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
13	13	13	2	126	36	5	15

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

tT	bi	riS	oT	roT	roS	iD
13	13	11	2	0	0	0	16

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
11	11	9	1	36	31	7	9

           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	2	2	41	27	2	13

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
11	11	9	1	37	32	7	11

           +----- Ctenodesmini   
  +--------+                     
  |        |  +-- Rectidentini   
  |        +--+                  
--+           +-- Contradentini  
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontina **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	2	2	72	24	4	13

  +-- Indopseudodontina **
--+                    
  +-- Pseudodontina **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	23	19	7	3

  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  |  |        +----- Ctenodesmini     
  +--+  +-----+                       
     |  |     |  +-- Rectidentini     
     |  |     +--+                    
     +--+        +-- Contradentini    
        |                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
8	8	5	2	114	25	5	13

        +-- Chamberlainiini  
  +-----+                    
  |     +-- Lamprotulini     
--+                          
  |  +----- Gonideini        
  +--+                       
     |  +-- Indopseudodontina **
     +--+                    
        +-- Pseudodontina **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	30	28	7	4

  +----------------- Chamberlainiini  
  |                                   
  |           +----- Ctenodesmini     
--+  +--------+                       
  |  |        |  +-- Rectidentini     
  |  |        +--+                    
  +--+           +-- Contradentini    
     |                                
     |  +----------- Lamprotulini     
     +--+                             
        |  +-------- Gonideini        
        +--+                          
           |  +----- Schepmaniini     
           +--+                       
              |  +-- Indopseudodontina **
              +--+                    
                 +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
8	8	5	2	29	27	7	7

  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |                          +-- Indopseudodontina **         
--+  +-----------------------+                             
  |  |                       +-- Pseudodontina **             
  |  |                                                     
  |  |                       +-- Ensidens                  
  |  |                       |                             
  |  |                       +-- Hyriopsis_desowitzi       
  +--+                       |                             
     |                       +-- Rectidens                 
     |  +--------------------+                             
     |  |                    +-- Hyriopsis_bialata         
     |  |                    |                             
     |  |                    +-- Ctenodesmini              
     |  |                    |                             
     +--+                    +-- Contradentini             
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Zieritz et al., 2024 [Publ 4381], fig. 8A

tT	bi	riS	oT	roT	roS	iD
8	8	6	1	0	0	0	12

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

           +----- Schepmaniini     
  +--------+                       
  |        |  +-- Indopseudodontina **
  |        +--+                    
--+           +-- Pseudodontina **    
  |                                
  |  +----------- Gonideini        
  |  |                             
  +--+        +-- Chamberlainiini  
     |  +-----+                    
     |  |     +-- Lamprotulini     
     +--+                          
        |  +----- Ctenodesmini     
        +--+                       
           |  +-- Rectidentini     
           +--+                    
              +-- Contradentini

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

  +-------------- Chamberlainiini  
  |                                
  |        +----- Ctenodesmini     
--+  +-----+                       
  |  |     |  +-- Rectidentini     
  |  |     +--+                    
  |  |        +-- Contradentini    
  +--+                             
     |        +-- Lamprotulini     
     |  +-----+                    
     |  |     +-- Gonideini        
     +--+                          
        |  +----- Schepmaniini     
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	4

  +-------------------------- Chamberlainiini  
  |                                            
  |                    +----- Ctenodesmini     
  |  +-----------------+                       
--+  |                 |  +-- Rectidentini     
  |  |                 +--+                    
  |  |                    +-- Contradentini    
  |  |                                         
  +--+                 +----- Schepmaniini     
     |  +--------------+                       
     |  |              |  +-- Indopseudodontina **
     |  |              +--+                    
     |  |                 +-- Pseudodontina **    
     +--+                                      
        |                 +-- Inversidens      
        |  +--------------+                    
        |  |              +-- Lamprotulini     
        +--+                                   
           |  +-------------- Sinosolenaia     
           |  |                                
           +--+           +-- Gonidea          
              |  +--------+                    
              |  |        +-- Microcondylaea   
              +--+                             
                 |  +-------- Parvasolenaia    
                 +--+                          
                    |  +----- Koreosolenaia    
                    +--+                       
                       |  +-- Postolata        
                       +--+                    
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
8	8	6	2	29	27	7	6

  +----------------------------- Schepmaniini       
  |                                                 
  |                          +-- Indopseudodontina **  
  |  +-----------------------+                      
--+  |                       +-- Pseudodontina **      
  |  |                                              
  |  |              +----------- Hyriopsis_bialata  
  |  |              |                               
  +--+  +-----------+        +-- Rectidens          
     |  |           |  +-----+                      
     |  |           |  |     +-- Ctenodesmini       
     |  |           +--+                            
     |  |              |  +----- Ensidens           
     |  |              +--+                         
     +--+                 |  +-- Hyriopsis_desowitzi
        |                 +--+                      
        |                    +-- Contradentini      
        |                                           
        |  +-------------------- Inversidens        
        |  |                                        
        |  |              +----- Lamprotula         
        +--+  +-----------+                         
           |  |           |  +-- Chamberlainiini    
           |  |           +--+                      
           +--+              +-- Potomida           
              |                                     
              |  +-------------- Sinosolenaia       
              |  |                                  
              +--+           +-- Gonidea            
                 |  +--------+                      
                 |  |        +-- Microcondylaea     
                 +--+                               
                    |  +-------- Koreosolenaia      
                    +--+                            
                       |  +----- Parvasolenaia      
                       +--+                         
                          |  +-- Postolata          
                          +--+                      
                             +-- Ptychorhynchus

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	43	17	3	9

  +----- Contradentini    
--+                       
  |  +-- Pseudodontina **    
  +--+                    
     +-- Indopseudodontina **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	1	34	8	4	13

        +-- Rectidentini 
  +-----+                
  |     +-- Contradentini
--+                      
  |  +----- Pseudodontina **
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	85	23	5	11

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	85	23	5	11

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
6	6	4	1	25	4	2	13

  +----- Pseudodontina **
--+                   
  |  +-- Rectidentini 
  +--+                
     +-- Contradentini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	75	22	5	13

              +-- Chamberlainiini
  +-----------+                  
  |           +-- Lamprotulini   
  |                              
--+           +-- Parvasolenaia  
  |  +--------+                  
  |  |        +-- Pseudodontina **  
  +--+                           
     |  +-------- Sinosolenaia   
     +--+                        
        |  +----- Gonidea        
        +--+                     
           |  +-- Microcondylaea 
           +--+                  
              +-- Leguminaia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	52	12	5	9

  +-------------- Chamberlainiini  
  |                                
--+           +-- Rectidentini     
  |  +--------+                    
  |  |        +-- Contradentini    
  +--+                             
     |  +-------- Lamprotulini     
     +--+                          
        |  +----- Gonideini        
        +--+                       
           |  +-- Indopseudodontina **
           +--+                    
              +-- Pseudodontina **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	52	12	5	9

  +----------- Chamberlainiini  
  |                             
  |        +-- Rectidentini     
--+  +-----+                    
  |  |     +-- Contradentini    
  |  |                          
  +--+     +-- Gonideini        
     |  +--+                    
     |  |  +-- Lamprotulini     
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
5	19	17	2	19	14	3	26

                 +----- Chamberlainia    
  +--------------+                       
  |              |  +-- Sundadontina **     
  |              +--+                    
  |                 +-- Sinohyriopsis    
  |                                      
--+              +----- Indopseudodontina **
  |  +-----------+                       
  |  |           |  +-- Pseudodon **        
  |  |           +--+                    
  |  |              +-- Pilsbryoconcha **   
  +--+                                   
     |              +-- Sinosolenaia     
     |  +-----------+                    
     |  |           +-- Lamprotulini     
     |  |                                
     +--+           +-- Ptychorhynchus   
        |  +--------+                    
        |  |        +-- Parvasolenaia    
        +--+                             
           |  +-------- Inversidens      
           +--+                          
              |  +----- Gonidea          
              +--+                       
                 |  +-- Microcondylaea   
                 +--+                    
                    +-- Leguminaia

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	57	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	57	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	0	2	54	0	0	10

  +-- Pseudodon **        
  |                    
  +-- Nyeinchanconcha **  
--+                    
  +-- Indopseudodontina **
  |                    
  +-- Sundadontina **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	0	1	31	31	2	11

  +----- Indopseudodontina **
--+                       
  |  +-- Rectidentini     
  +--+                    
     +-- Contradentini

Kongim et al., 2023 [Publ 4290], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	12	12	3	10

  +----------- Chamberlainiini  
--+                             
  |  +-------- Lamprotulini     
  +--+                          
     |  +----- Gonideini        
     +--+                       
        |  +-- Indopseudodontina **
        +--+                    
           +-- Pseudodontina **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	46	15	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	44	2	2	--

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	84	16	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Chen et al., 2020 [Publ 4170], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	32	9	3	--

  +----- Pseudodontina **  
--+                     
  |  +-- Lamprotulini   
  +--+                  
     +-- Chamberlainiini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	51	17	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	72	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	51	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	72	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	18	16	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	9	4	--

  +-------- Chamberlainiini
  |                        
--+     +-- Gonideini      
  |  +--+                  
  |  |  +-- Pseudodontina **  
  +--+                     
     |  +-- Contradentini  
     +--+                  
        +-- Lamprotulini

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	45	2	2	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	72	12	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	94	18	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	35	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	90	19	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	45	2	2	--

Froufe et al., 2019 [Publ 3936], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	17	6	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	26	24	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	21	10	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Pseudodontina **  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	29	6	--

  +----------------- Chamberlainiini  
  |                                   
--+  +-------------- Lamprotulini     
  |  |                                
  +--+  +----------- Gonideini        
     |  |                             
     +--+        +-- Pseudodontina **    
        |  +-----+                    
        |  |     +-- Indopseudodontina **
        +--+                          
           |  +----- Contradentini    
           +--+                       
              |  +-- Ctenodesmini     
              +--+                    
                 +-- Rectidentini

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	86	17	5	--

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	16	2	2	--

Froufe et al., 2019 [Publ 3936], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	17	6	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Gonideini      
     +--+                  
        +-- Pseudodontina **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	94	18	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	18	16	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Contradentini  
  |  +-----+                  
  |  |     +-- Rectidentini   
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	52	10	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Pseudodontina **  
  +--+                     
     |  +-- Lamprotulini   
     +--+                  
        +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	44	2	2	--

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	37	10	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Gonideini      
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Lamprotulini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	90	19	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	52	21	5	--

  +----------- Chamberlainiini
  |                           
--+        +-- Rectidentini   
  |  +-----+                  
  |  |     +-- Contradentini  
  +--+                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontina **  
        +--+                  
           +-- Gonideini

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	67	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	42	11	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

  +-- Pseudodontina **
--+                
  +-- Contradentini

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	1	1	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	1	1	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	62	0	0	--

-- Indopseudodon_kayinensis **

Walker et al., 2006 [Publ 2666], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	2	--

  +-- Gonideini    
--+                
  +-- Pseudodontina **

Walker et al., 2006 [Publ 2666], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

-- Pseudodon_vondembuschianus **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

-- Pseudodon_vondembuschianus **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	9	3	--

  +-------- Chamberlainiini
--+                        
  |  +----- Lamprotulini   
  +--+                     
     |  +-- Pseudodontina **  
     +--+                  
        +-- Gonideini

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	7	2	--

-- Pilsbryoconcha_exilis **

Walker et al., 2006 [Publ 2666], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	2	1	--

-- Pseudodon_vondembuschianus **

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