Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina
8 Recent freshwater species (9 total species)
(homonym of Jousseaume, 1875 [Gastr.]; intro'd as sub-gen. of Quadrula Rafinesque, 1820; renamed Pustulosa Frierson, 1927)
Frierson, 1927. Check List N. Amer. Naiades: 48.
Type species: Obliquaria (Quadrula) bullata Rafinesque, 1820 (original designation) = Pustulosa pustulosa
(intro'd as sub-gen. of Quadrula Rafinesque, 1820; new name for Bullata Frierson, 1927)
Frierson, 1927. Check List N. Amer. Naiades: errata et corrigenda.
Thiele, 1934. Handb. Syst. Weichtierk. 3: 827 [1260].
Type species: Obliquaria (Quadrula) bullata Rafinesque, 1820 = Pustulosa pustulosa
The Taxonomic History lists the ways authors have classified the species currently assigned to Pustulosa, since 1900.
Quadrula (sec. Theliderma) — Simpson (1900), Simpson (1914)
Quadrula (sec. Fusconaia) — Simpson (1900), Simpson (1914)
Unio (s.s.) — Call (1900)
Quadrula — Ihering (1901), Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ... Chase et al. (2018), Inoue et al. (2018), Marshall et al. (2018), Klymus et al. (2020), Gasparini et al. (2020)
Obovaria — Simpson (1914)
Unio — de Gregorio (1914), Henderson (1935)
Fusconaia — Ortmann (1923), Ortmann (1925), Clench & Turner (1956), Haas (1969), Burch (1975), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Quadrula (Bullata) — Frierson (1927)
Quadrula (Quincuncina) — Frierson (1927)
Quadrula (Striata) — Frierson (1927)
Quadrula (Fusconia [sic]) — Frierson (1927)
Quadrula (s.s.) — F.C. Baker (1928)
Quincuncina — Clench & Turner (1956), Haas (1969), Burch (1975), Davis & Fuller (1981), Davis (1983), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Pustulosa — Modell (1957), Neemuchwala et al. (2023), FMCS (2025), Campbell & Lydeard (2026)
Quadrula (Pustulosa) — Haas (1969)
Pleurobema — Morrison (1969), Vidrine (1993)
Obliquaria — Burch (1975)
Amphinaias — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011), ITIS (2013), Roe & Boyer (2015)
Rotundaria — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias — Williams et al. (2017), Burlakova et al. (2018), Johnson et al. (2018), Pieri et al. (2018), Lopes-Lima et al. (2019), ... FMCS (2021), de Moulpied et al. (2022), Hopper et al. (2023), Hauck et al. (2023), Freeman et al. (2025)
Cyclonaias (Quadrula) — Porto-Hannes et al. (2021)