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Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Lea (1852): attributed nominal species
     publication summary

Complete Citation

Lea, I. 1852. Descriptions of new species of the family Unionidæ [New fresh water and land shells]. Transactions of the American Philosophical Society 10 [O. 5]: 253-294, pls. 12-29 (Obs. 5: 9-50). Available electronically.

Attributed Nominal Species

Anodonta arkansensis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 293 [49], pl. 29, fig. 56.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86603, Lit. Ark. R., at Santa Fe...

Anodonta californiensis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 286 [42], pl. 25, fig. 47.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): LECTOTYPE [AS HOLOTYPE] USNM_86393 (by R.I. Johnson, 1974: 26), Colo. R., Calif.

Anodonta denigrata Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 285, pl. 25, fig. 45.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86535, Campbell Co., Tenn.

Anodonta linnaeana Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 289 [45], pl. 27, fig. 51 [as ‘linnæana’].
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘linnæana’]. [nomen nudum]
(correct name for Anodonta linnaena Lea, 1852)
Type(s): FIGURED SYNTYPE USNM_86702, Lake Concordia, Louisiana.

Anodonta linnaena Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 289 [45] [as ‘linnæna’].
(incorrect original spelling of Anodonta linnaeana Lea, 1852)

Anodonta oblita Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 290, pl. 28. fig. 52.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86536, Campbell Co., Tenn.

Anodonta opaca Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 285, pl. 25, fig. 46.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_128835, New Orleans.

Anodonta schroeteriana Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 292, pl. 29, fig. 55 [as ‘schröteriana’].
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘schröteriana’]. [nomen nudum]
Type(s): HOLOTYPE USNM_86727, Rio Negro, Brazil.

Anodonta shaefferiana Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 288 [44], pl. 26, fig. 50.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86685, Horn Lake Creek, Tenn.

Anodonta tortilis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 291, pl. 28, fig. 54.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE (MONOTYPY) USNM_86405, Cartagena, S.A.

Anodonta trautwiniana Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 287, pl. 26, fig. 48.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘troutwiniana’]. [nomen nudum]
Type(s): LECTOTYPE [AS HOLOTYPE] USNM_86684 (by Johnson & Baker, 1973: 173), Carthegena, U.S. Colombia.

Anodonta virens Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 290 [46], pl. 28, fgi. 53.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86666, Red R., Alexandria, Louisiana.

Anodonta wheatleyi Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 287, pl. 26, fig. 49.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86723, Amazon River, Pará.

Unio affinis Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 271 [27], pl. 19, fig. 26.
Type(s): HOLOTYPE USNM_84910, Louisiana.

Unio barrattii Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 256 [12], pl. 13, fig. 5.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_86010, Abbeville District, S.C.

Unio buxeus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 261 [17], pl. 15, fig. 13.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85153, Abbeville, S Carolina.

Unio clarkianus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 273, pl. 21, fig. 30.
Type(s): SYNTYPE ANSP_126052, Tenn.

Unio concavus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 260, pl. 15, fig. 11.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85154, Abbeville District, S Carolina.

Unio cuvierianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 263 [19], pl. 16, fig. 16.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85401, Washington Co., Ga.

Unio decoratus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 257, pl. 13, fig. 6.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): LECTOTYPE USNM_83972, Abbeville, S. Carolina.

Unio floridensis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 274 [30], pl. 21, fig. 31.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE ANSP_42081, Chattahoochee R.

Unio forbesianus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 264 [20], pl. 16, fig. 17.
Type(s): HOLOTYPE USNM_84542, Savannah R., Ga.

Unio fraternus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 263 [19], pl. 16, fig. 15.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): LECTOTYPE USNM_85396, Abbeville, S.C.

Unio gibbesianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 254 [10], pl. 12, fig. 2.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85685, Abbeville, S.C.

Unio hanleyianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 279 [35], pl. 23, fig. 37.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251 [as ‘henleyianus’]. [nomen nudum]
Type(s): HOLOTYPE USNM_84717, Coosawattee R., Ga.

Unio hebes Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 267 [23], pl. 18, fig. 21.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85383, Oconee R., near Athens, Georgia.

Unio ineptus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 261 [17], pl. 15, fig. 12.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251 [as ‘irreptus’]. [nomen nudum]
Type(s): HOLOTYPE USNM_85326, Abbeville District, SC.

Unio ingallsianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 282, pl. 24, fig. 41.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85933, Siam.

Unio keinerianus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40.
(incorrect original spelling of Unio kienerianus Lea, 1852)
Type(s): HOLOTYPE USNM_84273, Coosawattee R., Ga.

Unio kienerianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40 [as ‘keineriana’].
(correct name for Unio keinerianus Lea, 1852)

Unio kleinianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 265 [21], pl. 17, fig. 18.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_84041, Suwanee R., Fla.

Unio lamarckianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 266 [22], pl. 17, fig. 20.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_84306, White R., Arkansas.

Unio lazarus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 259 [15], pl. 14, fig. 9.
Type(s): HOLOTYPE USNM_86155, Abbeville District, S.C.

Unio luridus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 273 [29], pl. 20, fig. 29.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85253, Coosawattee R., Murray Co., Ga.

Unio merus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 260 [16], pl. 15, fig. 10.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85698, Abbeville District, S.C.

Unio moussonianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 268 [24], pl. 118, fig. 22.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85168, Georgia.

Unio nigellus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 283 [39], pl. 24, fig. 42.
Type(s): LECTOTYPE USNM_85567, Chattahoochee, Columbus [Columbia], Ga.

Unio nigerrimus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 268, pl. 18, fig. 23.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85307, Alexandria, Louisiana.

Unio nigrinus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 284 [40], pl. 24, fig. 44.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_86132, W Florida.

Unio nux Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 283 [39], pl. 24, fig. 43.
Type(s): HOLOTYPE USNM_85316, Alabama R., Ala.

Unio oregonensis Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 275 [31], pl. 22, fig. 33.
Type(s): HOLOTYPE USNM_85073, Columbia R., Oregon.

Unio perstriatus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 255 [11], pl. 12, fig. 3.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85892, Abbeville, S.C.

Unio placitus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 279 [35], pl. 23, fig. 38.
Type(s): HOLOTYPE USNM_85152, Alabama.

Unio powellii Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 270 [26], pl. 19, fig. 25.
Type(s): HOLOTYPE USNM_85024, Saline R., Arkansas.

Unio prevostianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 269 [25], pl. 19, fig. 24.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]

Unio proximus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 271 [27], pl. 20, fig. 27.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85131, Georgia.

Unio pygmaeus Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘pygmæus’]. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 262, pl. 15, fig. 14 [as ‘pygmæus’].
Type(s): HOLOTYPE USNM_85240, Abbeville District, SC.

Unio reeveianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 272, pl. 20, fig. 28.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘reevesianus’]. [nomen nudum]
Type(s): HOLOTYPE USNM_85025, Alexandria, Louisiana.

Unio rufusculus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 258, pl. 14, fig. 7.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85683, Abbeville, S.C.

Unio rumphianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 276 [32], pl. 22, fig. 34.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_84189, West Georgia.

Unio sagittiformis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 277 [33], pl. 22, fig. 35.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252 [as ‘sagittaformis’]. [nomen nudum]
Type(s): HOLOTYPE USNM_85970, Oconee R., near Athens, Ga.

Unio satur Lea, 1852
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 265 [21], pl. 17, fig. 19.
Type(s): HOLOTYPE USNM_84505, L. Calcasieu, La.

Unio sordidis Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 254 [10].
(incorrect original spelling of Unio sordidus Lea, 1852)

Unio sordidus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 254 [10], pl. 12, fig. 1 [as ‘sordidis’].
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
(correct name for Unio sordidis Lea, 1852)
Type(s): HOLOTYPE USNM_85688, Abbeville, S.C.

Unio stewardsonii Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 278, pl. 23, fig. 36.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE ANSP_56572, Chattanooga River, Ga.

Unio succissus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 275 [31], pl. 21, fig. 32.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_84574, W Florida.

Unio troschelianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 280 [36], pl. 23, fig. 39.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_84790, Coosawattee R., Ga.

Unio tuomeyi Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 256 [12], pl. 13, fig. 4.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85669, Abbeville District, S. Carolina.

Unio whiteianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 258 [14], pl. 14, fig. 8.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85657, near Savannah, Georgia.

 
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