The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina genus Inversiunio Habe, 1991: cladomics
4 Recent valid species (4 total species) Cladomics Summarynumber of species = 4 (4 total)
analyzed in 27 cladograms in 15 publications
maximum number of terminals in an analysis =
4 (mean = 2.19, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 18
maximum number of included terminals = 4, (mean =
2.78 median = 3)
proportion of cladograms that support basic monophyly
= 1
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Sano et al., 2020 [Publ 4138], fig. 1 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 6 | 32 | 4 | 3 | -- | +----- Schistodesmus
|
| +-- Inversiunio_reinianus__1 **
--+ |
| +-- Inversiunio_reinianus__2 **
| |
| +-- Inversiunio_jokohamensis__1 **
+--+
+-- Inversiunio_jokohamensis__2 **
|
+-- Inversiunio_jokohamensis__3 **
|
+-- Inversiunio_jokohamensis__4 **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 5 | 627 | 28 | 6 | -- | +-- Schistodesmus
+-----------------+
| +-- Nodularia
|
--+ +----- Pseudocuneopsis
| +-----------+
| | | +-- Diaurora
| | +--+
+--+ +-- Cuneopsis
|
| +-------------- Tchangsinaia
+--+
| +----------- Inversiunio_yanagawensis__2 **
+--+
| +-------- Inversiunio_reinianus **
+--+
| +----- Inversiunio_verrucosus **
+--+
| +-- Inversiunio_jokohamensis **
+--+
+-- Inversiunio_yanagawensis__1 **
Lopes-Lima et al., 2020 [Publ 4080], fig. 12 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 5 | 4 | 2 | 1 | -- | +----------- Nodularia
|
--+ +-------- Inversiunio_jokohamensis **
| |
+--+ +----- Inversiunio_verrucosus__1 **
| |
+--+----- Inversiunio_verrucosus__2 **
|
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 5 | 9 | 6 | 1 | -- | +-------- Inversiunio_jokohamensis **
|
--+ +----- Inversiunio_yanagawensis__2 **
| |
+--+----- Inversiunio_verrucosus **
|
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis__1 **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 118 | 14 | 4 | -- | +----------------- Nodularia
|
--+ +-------------- Schistodesmus
| |
+--+-------------- Cuneopsis
|
| +----------- Tchangsinaia
+--+
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_verrucosus **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 5 | 30 | 5 | 3 | -- | +-------- Inversiunio_jokohamensis **
|
--+ +----- Inversiunio_yanagawensis__3 **
| |
+--+ +-- Inversiunio_yanagawensis__1 **
| |
+--+-- Inversiunio_yanagawensis__2 **
|
+-- Inversiunio_reinianus **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 4 | 65 | 4 | 3 | -- | +----- Nodularia
+-----+
| | +-- Schistodesmus
| +--+
--+ +-- Tchangsinaia
|
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis__2 **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis__1 **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 4 | 65 | 4 | 3 | -- | +-------------- Nodularia
|
--+ +-- Schistodesmus
| +--------+
| | +-- Tchangsinaia
+--+
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis__2 **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis__1 **
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 4 | 6 | 18 | 1 | 1 | -- | +----------------- Nodularia
--+
| +-------------- Inversiunio_jokohamensis **
+--+
| +----------- Inversiunio_yanagawensis__1 **
+--+
| +-------- Inversiunio_yanagawensis__2 **
+--+
| +----- Inversiunio_yanagawensis__3 **
+--+
| +-- Inversiunio_yanagawensis__4 **
+--+
+-- Inversiunio_reinianus **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 36 | 12 | 5 | -- | +-------------------- Pseudocuneopsis
--+
| +----------------- Nodularia
+--+
| +-------------- Schistodesmus
+--+
| +----------- Cuneopsis
+--+
| +-------- Tchangsinaia
+--+
| +----- Inversiunio_jokohamensis **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 4 | 65 | 4 | 3 | -- | +----- Nodularia
+-----+
| | +-- Tchangsinaia
| +--+
--+ +-- Schistodesmus
|
| +-------- Inversiunio_jokohamensis **
+--+
| +----- Inversiunio_yanagawensis__2 **
+--+
| +-- Inversiunio_reinianus **
+--+
+-- Inversiunio_yanagawensis__1 **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 12 | 6 | -- | +-- Inversiunio_yanagawensis **
--+
+-- Inversiunio_reinianus **
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 11 | 6 | -- | +----- Tchangsinaia
+-----+
| | +-- Inversiunio_yanagawensis **
| +--+
| +-- Inversiunio_reinianus **
--+
| +-- Cuneopsis
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Nodularia
+--+
| +-- Diaurora
+--+
+-- Schistodesmus
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 12 | 6 | -- | +-- Inversiunio_yanagawensis **
--+
+-- Inversiunio_reinianus **
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 22 | 1 | 1 | -- | +----- Nodularia
--+
| +-- Inversiunio_jokohamensis **
+--+
+-- Inversiunio_reinianus **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 11 | 6 | -- | +----- Tchangsinaia
--+
| +-- Inversiunio_yanagawensis **
+--+
+-- Inversiunio_reinianus **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 11 | 6 | -- | +-- Inversiunio_yanagawensis **
--+
+-- Inversiunio_reinianus **
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 11 | 6 | -- | +----- Tchangsinaia
+--------+
| | +-- Inversiunio_yanagawensis **
| +--+
--+ +-- Inversiunio_reinianus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Cuneopsis
+--+
| +----- Nodularia
+--+
| +-- Diaurora
+--+
+-- Schistodesmus
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 8 | 3 | -- | +-------- Nodularia
--+
| +----- Cuneopsis
+--+
| +-- Inversiunio_jokohamensis **
+--+
+-- Schistodesmus
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 8 | 3 | -- | +----- Cuneopsis
|
--+----- Nodularia
|
| +-- Inversiunio_jokohamensis **
+--+
+-- Schistodesmus
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 10 | 5 | -- | +----- Cuneopsis
|
| +-- Nodularia
+--+
--+ +-- Schistodesmus
|
| +-- Tchangsinaia
| |
+--+-- Inversiunio_yanagawensis **
|
+-- Pseudocuneopsis
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 10 | 5 | -- | +-------------- Nodularia
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 10 | 5 | -- | +----------- Cuneopsis
|
--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio_yanagawensis **
+--+
+-- Pseudocuneopsis
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 7 | 2 | -- | +----- Inversiunio_jokohamensis **
--+
| +-- Cuneopsis
+--+
+-- Nodularia
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 9 | 5 | -- | +----------- Cuneopsis
|
--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio_yanagawensis **
+--+
+-- Pseudocuneopsis
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Inversiunio_yanagawensis **
--+
+-- Nodularia
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | -- Inversiunio_reinianus **
|
