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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Pseudanodonta Bourguignat, 1876: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 72 cladograms in 46 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110163192--
  +----------------------- Anodonta_anatina           
  |                                                   
--+  +-------------------- Pseudanodonta_complanata **   
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2
     |  |                                             
     |  |           +----- Anodonta_cygnea            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata        
           |                                          
           |  +----------- Brachyanodon_impura        
           |  |                                       
           +--+  +-------- Brachyanodon               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2
                    |                                 
                    |  +-- Brachyanodon_californiensis
                    +--+                              
                       +-- Brachyanodon_oregonensis__1

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015242--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113832--
  +----- Pseudanodonta_complanata **
--+                              
  |  +-- Anodonta                
  +--+                           
     +-- Brachyanodon

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
11015142--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014311--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Wu et al., 2026 [Publ 4825], fig. 6B

iTtTbiriSoTroTroSiD
11016431--
-- Pseudanodonta_complanata **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
1101621--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Pseudanodonta_complanata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112132--
-- Pseudanodonta_complanata **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012300--
-- Pseudanodonta_complanata **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
1101321--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017442--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
1101821--
-- Pseudanodonta_complanata **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
11011821--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
1101621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
11016331--
-- Pseudanodonta_complanata **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014311--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015521--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019442--
-- Pseudanodonta_complanata **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011721--
-- Pseudanodonta_complanata **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
1101821--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
11014011--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019042--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119832--
-- Pseudanodonta_complanata **

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
1101221--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
11019431--
-- Pseudanodonta_complanata **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
1101531--
     +-- Anodonta_anatina        
  +--+                           
  |  +-- Pseudanodonta_complanata **
--+                              
  |  +-- Anodonta_exulcerata     
  +--+                           
     +-- Anodonta_cygnea

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110120032--
-- Pseudanodonta_complanata **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012300--
-- Pseudanodonta_complanata **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012200--
-- Pseudanodonta_complanata **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
1101721--
  +-- Pseudanodonta_complanata **
  |                           
--+-- Anodonta_anatina        
  |                           
  +-- Anodonta_cygnea

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
1101400--
-- Pseudanodonta_complanata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11018032--
  +----- Pseudanodonta_complanata **
  |                              
--+  +-- Anodonta_anatina        
  |  |                           
  +--+-- Anodonta_cygnea         
     |                           
     +-- Brachyanodon

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
11013041--
  +----------- Anodonta_anatina        
--+                                    
  |  +-------- Pseudanodonta_complanata **
  +--+                                 
     |  +----- Anodonta_exulcerata     
     +--+                              
        |  +-- Anodonta_seddonae       
        +--+                           
           +-- Anodonta_cygnea

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112432--
-- Pseudanodonta_complanata **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011321--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Pseudanodonta_complanata **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
11013041--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
1101431--
-- Pseudanodonta_complanata **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
11013521--
-- Pseudanodonta_complanata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110113942--
  +-- Brachyanodon            
--+                           
  +-- Pseudanodonta_complanata **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011842--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019021--
-- Pseudanodonta_complanata **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1101511--
  +-- Anodonta     
--+                
  +-- Pseudanodonta **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014011--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
1101821--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Pseudanodonta_complanata **

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
1101731--
  +-------- Anodonta_anatina        
--+                                 
  |  +----- Pseudanodonta_complanata **
  +--+                              
     |  +-- Anodonta_cygnea         
     +--+                           
        +-- Anodonta_exulcerata

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019021--
-- Pseudanodonta_complanata **

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
1101221--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019042--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
1101221--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110120032--
-- Pseudanodonta_complanata **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
1101321--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Pseudanodonta_complanata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113832--
-- Pseudanodonta_complanata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017532--
  +----- Pseudanodonta_complanata **
--+                              
  |  +-- Brachyanodon            
  +--+                           
     +-- Anodonta

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
11011221--
-- Pseudanodonta_complanata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013521--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
11011621--
  +----- Anodonta_anatina        
--+                              
  |  +-- Pseudanodonta_complanata **
  +--+                           
     +-- Anodonta_cygnea

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014011--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta_anatina        
  |                           
--+-- Anodonta_cygnea         
  |                           
  +-- Pseudanodonta_complanata **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta_anatina        
  |                           
--+-- Pseudanodonta_complanata **
  |                           
  +-- Anodonta_cygnea

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
11014032--
  +-- Brachyanodon            
--+                           
  +-- Pseudanodonta_complanata **

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
1101411--
  +-- Pseudanodonta_complanata **
--+                           
  +-- Anodonta

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
11014011--
  +-- Anodonta                
--+                           
  +-- Pseudanodonta_complanata **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
1101431--
-- Pseudanodonta_complanata **

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
11011762--
-- Pseudanodonta_complanata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019442--
  +----- Anodonta                
--+                              
  |  +-- Brachyanodon            
  +--+                           
     +-- Pseudanodonta_complanata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017532--
  +-------- Pseudanodonta_complanata **
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Brachyanodon            
     +--+                           
        +-- Anodonta_cygnea
 
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