The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina genus Pseudodon Gould, 1844: cladomics
9 Recent valid species (9 total species) Cladomics Summarynumber of species = 9 (9 total)
analyzed in 98 cladograms in 58 publications
maximum number of terminals in an analysis =
7 (mean = 1.79, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 32
maximum number of included terminals = 7, (mean =
3.41 median = 3)
proportion of cladograms that support basic monophyly
= 0.88
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Zieritz et al., 2026 [Publ 4812], fig. 5a ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 7 | 9 | 1 | 1 | 1 | 0 | +-------------------- Namkongnaia
|
--+ +----------------- Pseudodon_mekongi **
| |
| | +----- Pseudodon_lenyanensis__2 **
+--+ +--------+
| | | +-- Pseudodon_inoscularis **
| | +--+
+--+ +-- Pseudodon_lenyanensis__1 **
|
| +----------- Pseudodon_vondembuschianus **
+--+
| +-------- Pseudodon_vagulus **
+--+
| +----- Pseudodon_cokelatus__2 **
+--+
| +-- Pseudodon_walpolei **
+--+
+-- Pseudodon_cokelatus__1 **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 6 | 7 | 626 | 34 | 9 | 10 | +-- Lannanaia
+-----------------------+
| +-- Pilsbryoconcha
|
--+ +----------------------- Songkhlanaia
| |
| | +-- Nyeinchanconcha
+--+ +-----------------+
| | +-- Sundadontina
| |
| | +-- Isannaia
+--+ +--+
| | +-- Namkongnaia
| +-----------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +-------------- Pseudodon_walpolei **
| |
+--+ +----------- Pseudodon_vagulus **
| |
| | +----- Pseudodon_vondembuschianus **
+--+ +--+
| | | +-- Pseudodon_mekongi **
| | +--+
+--+ +-- Schepmania
|
| +----- Pseudodon_lenyanensis__2 **
+--+
| +-- Pseudodon_inoscularis **
+--+
+-- Pseudodon_lenyanensis__1 **
Jeratthitikul et al., 2025 [Publ 4476], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 8 | 8 | 4 | 10 | +-------------------- Pilsbryoconcha
|
| +----- Namkongnaia
--+ +-----------+
| | | +-- Isannaia
| | +--+
+--+ +-- Sundadontina
|
| +-------------- Pseudodon_walpolei **
| |
+--+ +-- Pseudodon_inoscularis **
| +--------+
| | +-- Pseudodon_lenyanensis **
+--+
| +-------- Pseudodon_mekongi **
+--+
| +----- Pseudodon_vondembuschianus **
+--+
| +-- Pseudodon_vagulus **
+--+
+-- Pseudodon_septuaginta **
Zieritz et al., 2024 [Publ 4381], fig. 8A iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 1 | 1 | 1 | 3 | +-------------- Namkongnaia
|
--+ +----------- Pseudodon_mekongi **
| |
+--+ +-- Pseudodon_lenyanensis **
| +-----+
| | +-- Pseudodon_inoscularis **
+--+
| +----- Pseudodon_vondembuschianus **
+--+
| +-- Pseudodon_walpolei **
+--+
+-- Pseudodon_vagulus **
Bolotov et al., 2023 [Publ 4312], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 35 | 30 | 7 | 8 | +-------------------- Songkhlanaia
|
| +-- Nyeinchanconcha
--+ +--------------+
| | +-- Sundadontina
| |
+--+ +----- Namkongnaia
| +--------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----------- Pilsbryoconcha
| |
+--+ +-------- Pseudodon_vondembuschianus **
| |
+--+ +-- Pseudodon_mekongi **
| +--+
| | +-- Pseudodon_vagulus **
+--+
| +-- Pseudodon_lenyanensis **
+--+
+-- Pseudodon_inoscularis **
Bolotov et al., 2023 [Publ 4312], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 33 | 28 | 7 | 6 | +----------------------- Pilsbryoconcha
|
| +----- Nyeinchanconcha
--+ +--------------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----------------- Songkhlanaia
| |
+--+ +-------------- Sundadontina
| |
+--+ +----------- Namkongnaia
| |
+--+ +-- Pseudodon_inoscularis **
| +-----+
| | +-- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_mekongi **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_vagulus **
Bolotov et al., 2023 [Publ 4312], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 33 | 28 | 7 | 4 | +-------------------- Pilsbryoconcha
|
--+ +----------------- Songkhlanaia
| |
| | +-------- Sundadontina
+--+ +-----+
| | | +----- Nyeinchanconcha
| | +--+
| | | +-- Thaiconcha
+--+ +--+
| +-- Bineurus
|
| +----------- Namkongnaia
| |
+--+ +-------- Pseudodon_mekongi **
| |
+--+ +-- Pseudodon_vondembuschianus **
| +--+
| | +-- Pseudodon_vagulus **
+--+
| +-- Pseudodon_lenyanensis **
+--+
+-- Pseudodon_inoscularis **
Konopleva et al., 2025 [Publ 4446], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 39 | 34 | 9 | 6 | +-------------------------- Lannanaia
|
--+ +----------------------- Pilsbryoconcha
| |
+--+ +-------------------- Songkhlanaia
| |
| | +----- Nyeinchanconcha
+--+ +-----------+
| | | +-- Thaiconcha
| | +--+
| | +-- Bineurus
+--+
| +-------------- Sundadontina
| |
| | +-- Namkongnaia
+--+ +--------+
| | +-- Isannaia
| |
+--+ +-- Pseudodon_inoscularis **
| +-----+
| | +-- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_mekongi **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_vagulus **
Jeratthitikul et al., 2024 [Publ 4387], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 54 | 31 | 9 | 6 | +-- Pseudodon_inoscularis **
+-----+
| +-- Pseudodon_lenyanensis **
+-----+
| | +----- Pseudodon_mekongi **
| +--+
| | +-- Pseudodon_vagulus **
| +--+
| +-- Pseudodon_vondembuschianus **
--+
| +-------- Sundadontina
| +--+
| | | +----- Nyeinchanconcha
| | +--+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +-- Isannaia
| +-----+
| | +-- Pilsbryoconcha
+--+
| +----- Namkongnaia
+--+
| +-- Lannanaia
+--+
+-- Songkhlanaia
Jeratthitikul et al., 2025 [Publ 4423], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 45 | 33 | 9 | 6 | +-- Lannanaia
+-----------------------+
| +-- Pilsbryoconcha
|
--+ +----------------------- Songkhlanaia
| |
| | +----- Nyeinchanconcha
+--+ +--------------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----------------- Sundadontina
| |
+--+ +-------------- Isannaia
| |
+--+ +----------- Namkongnaia
| |
+--+ +-- Pseudodon_inoscularis **
| +-----+
| | +-- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_mekongi **
+--+
| +-- Pseudodon_vagulus **
+--+
+-- Pseudodon_vondembuschianus **
Konopleva et al., 2021 [Publ 4213], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 23 | 15 | 6 | 8 | +----- Pilsbryoconcha_exilis__2
+--------+
| | +-- Pilsbryoconcha_exilis__1
| +--+
| +-- Namkongnaia
--+
| +----- Nyeinchanconcha
| +-----+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +-------- Sundadontina
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Konopleva et al., 2023 [Publ 4262], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 79 | 22 | 7 | 8 | +-------------------- Pilsbryoconcha
|
--+ +----------------- Songkhlanaia
| |
+--+ +-------------- Namkongnaia
| |
| | +----- Pseudodon_lenyanensis **
+--+ +-----+
| | | +-- Pseudodon_mekongi **
| | +--+
+--+ +-- Pseudodon_vondembuschianus **
|
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 4 | 3 | 135 | 9 | 5 | 10 | +-- Pseudodon_mekongi **
+-----+
| +-- Pseudodon_vondembuschianus **
|
--+ +-- Pseudodon_inoscularis **
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha
+--+
+-- Sundadontina_cumingii
Jeratthitikul et al., 2021 [Publ 4219], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 32 | 14 | 6 | 8 | +----------------- Pilsbryoconcha
|
| +-- Nyeinchanconcha
| +--+
--+ | +-- Sundadontina_cumingii
| +--------+
| | | +-- Thaiconcha
| | +--+
| | +-- Bineurus
+--+
| +----- Sundadontina_tumida
| +-----+
| | | +-- Sundadontina_brandti
| | +--+
+--+ +-- Sundadontina_taskaevi
|
| +-------- Namkongnaia
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Dai et al., 2023 [Publ 4285], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 42 | 11 | 5 | 8 | +-- Nyeinchanconcha
+--------+
| +-- Bineurus
|
| +-- Thaiconcha
--+--------+
| +-- Sundadontina
|
| +-------- Pilsbryoconcha
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 108 | 13 | 5 | 8 | +----------------- Bineurus
|
--+ +-------------- Nyeinchanconcha
| |
+--+ +-- Thaiconcha
| +--------+
| | +-- Pilsbryoconcha
+--+
| +-------- Sundadontina
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Jeratthitikul et al., 2021 [Publ 4219], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 31 | 13 | 6 | 8 | +----- Pilsbryoconcha_exilis
+-----------+
| | +-- Pilsbryoconcha_linguaeformis
| +--+
| +-- Namkongnaia_lemeslei__2
--+
| +----- Nyeinchanconcha
| +--------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----------- Sundadontina
| |
+--+ +-- Namkongnaia_lemeslei__1
| +-----+
| | +-- Namkongnaia_inkhavilayi
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Dai et al., 2023 [Publ 4285], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 42 | 11 | 5 | 8 | +-- Nyeinchanconcha
+-----------+
| +-- Bineurus
|
--+ +-- Thaiconcha
| +--------+
| | +-- Sundadontina
+--+
| +-------- Pilsbryoconcha
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 114 | 13 | 5 | 8 | +----------------- Bineurus
|
--+ +-------------- Nyeinchanconcha
| |
+--+ +-- Thaiconcha
| +--------+
| | +-- Pilsbryoconcha
+--+
| +-------- Sundadontina
+--+
| +----- Pseudodon_lenyanensis **
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 7 | 0 | 3 | 80 | 5 | 2 | 12 | +-- Bineurus_mouhotii
|
+-- Pseudodon_vondembuschianus **
|
+-- Pseudodon_mekongi__1 **
|
+-- Pilsbryoconcha_exilis
--+
+-- Pilsbryoconcha_carinifera
|
+-- Bineurus_loeiensis
|
+-- Bineurus_exilis
|
+-- Pseudodon_mekongi__2 **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 5 | 4 | -- | +----- Pilsbryoconcha
+--+
| | +-- Pseudodon_walpolei **
| +--+
--+ +-- Pseudodon_mekongi **
|
| +----- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 198 | 9 | 5 | -- | +----------- Pilsbryoconcha
|
| +----- Nyeinchanconcha
--+ +--+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----- Sundadontina
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 45 | 9 | 5 | -- | +-------------- Pilsbryoconcha
|
| +----- Pseudodon_vondembuschianus__2 **
--+ +-----+
| | | +-- Pseudodon_mekongi **
| | +--+
+--+ +-- Pseudodon_vondembuschianus__1 **
|
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Bineurus
+--+
+-- Thaiconcha
Konopleva et al., 2017 [Publ 3807], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 4 | 4 | -- | +----------- Sundadontina
|
--+ +-- Nyeinchanconcha
| +-----+
| | +-- Bineurus
+--+
| +----- Pilsbryoconcha
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_mekongi **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 196 | 9 | 5 | -- | +----------- Pilsbryoconcha
|
| +----- Sundadontina_tumida
--+ +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
+--+
| +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina_cumingii
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 198 | 9 | 5 | -- | +----- Nyeinchanconcha
|
+----- Thaiconcha
|
+----- Sundadontina_cumingii
|
+----- Sundadontina_tumida__1
--+
+----- Sundadontina_tumida__2
|
+----- Pilsbryoconcha
|
+----- Bineurus
|
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 38 | 4 | 4 | -- | +----------- Pilsbryoconcha
|
--+ +-- Pseudodon_mekongi **
| +-----+
| | +-- Pseudodon_vondembuschianus **
+--+
| +----- Sundadontina
+--+
| +-- Nyeinchanconcha
+--+
+-- Bineurus
Chen et al., 2025 [Publ 4663], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 46 | 9 | 5 | -- | +-------------- Pilsbryoconcha
|
| +----- Pseudodon_vondembuschianus__2 **
--+ +-----+
| | | +-- Pseudodon_mekongi **
| | +--+
+--+ +-- Pseudodon_vondembuschianus__1 **
|
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Bineurus
+--+
+-- Thaiconcha
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 136 | 9 | 5 | -- | +----------- Pilsbryoconcha
|
| +----- Sundadontina_tumida
--+ +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
+--+
| +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina_cumingii
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 122 | 9 | 5 | -- | +-------- Bineurus
|
| +-- Nyeinchanconcha
+-----+
| +-- Sundadontina_cumingii
--+
| +-- Thaiconcha
+-----+
| +-- Pilsbryoconcha
|
| +----- Sundadontina_tumida
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Pseudodon_vondembuschianus **
Konopleva et al., 2019 [Publ 3900], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 7 | 0 | 3 | 49 | 5 | 2 | 12 | +-- Bineurus_mouhotii
|
+-- Pseudodon_vondembuschianus **
|
+-- Pseudodon_mekongi__1 **
|
+-- Pilsbryoconcha_exilis
--+
+-- Pilsbryoconcha_carinifera
|
+-- Bineurus_loeiensis
|
+-- Bineurus_exilis
|
+-- Pseudodon_mekongi__2 **
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 9 | 5 | -- | +-------------- Pilsbryoconcha
|
--+ +-- Pseudodon_mekongi **
| +--------+
| | +-- Pseudodon_vondembuschianus **
+--+
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Bineurus
+--+
+-- Thaiconcha
Bolotov et al., 2022 [Publ 4239], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 2 | 2 | -- | -- Pseudodon_lenyanensis **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Matos et al., 2024 [Publ 4383], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Wu et al., 2024 [Publ 4350], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Bineurus
+--+
| +-- Pseudodon_mekongi **
+--+
+-- Thaiconcha
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Wu et al., 2024 [Publ 4350], fig. 5c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
|
--+ +----- Bineurus_anodontinum
| |
+--+----- Bineurus_loeiensis
|
| +-- Pseudodon_mekongi **
+--+
+-- Thaiconcha
Zieritz et al., 2026 [Publ 4812], fig. 4a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 8 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Bolotov et al., 2022 [Publ 4252], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 60 | 2 | 2 | -- | -- Pseudodon_lenyanensis **
Wu et al., 2024 [Publ 4388], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Pseudodon_mekongi **
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Bineurus
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Sundadontina
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Pseudodon_vondembuschianus **
+--+
| +-- Bineurus
+--+
+-- Nyeinchanconcha
Klymus et al., 2025 [Publ 4814], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Klymus et al., 2025 [Publ 4814], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Bolotov et al., 2022 [Publ 4252], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 60 | 2 | 2 | -- | -- Pseudodon_lenyanensis **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 140 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Bolotov et al., 2022 [Publ 4239], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 2 | 2 | -- | -- Pseudodon_lenyanensis **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- | +-- Pseudodon_vondembuschianus **
--+
+-- Sundadontina
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2024 [Publ 4350], fig. 6-7 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Pseudodon_mekongi **
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Bolotov et al., 2022 [Publ 4239], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 2 | 2 | -- | -- Pseudodon_lenyanensis **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- | Zieritz et al., 2021 [Publ 4218], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 4 | -- | +----------- Pilsbryoconcha
--+
| +-------- Pseudodon_mekongi **
+--+
| +----- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Matos et al., 2024 [Publ 4384], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Bolotov et al., 2022 [Publ 4250], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 1 | 1 | -- | +-- Pseudodon_lenyanensis **
--+
+-- Sundadontina
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 5 | 4 | -- | +-------- Pilsbryoconcha
|
--+ +----- Pseudodon_mekongi **
| |
+--+ +-- Nyeinchanconcha
| |
+--+-- Bineurus
|
+-- Sundadontina
Liu et al., 2024 [Publ 4380], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Pseudodon_mekongi **
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Pseudodon_vondembuschianus **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Zhang et al., 2024 [Publ 4397], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Pseudodon_vondembuschianus **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 2 | 1 | -- | +----- Pilsbryoconcha_exilis
--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pilsbryoconcha_carinifera
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 85 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 1 | 1 | -- | +-- Pseudodon_mekongi **
--+
+-- Pilsbryoconcha
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- | +-------- Bineurus
--+
| +----- Pilsbryoconcha
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Sundadontina
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 14 | 6 | -- | +----- Pilsbryoconcha
+--+
| | +-- Namkongnaia
| +--+
| +-- Pseudodon_mekongi **
--+
| +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Pseudodon_vondembuschianus **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 1 | 1 | -- | +-- Pilsbryoconcha
--+
+-- Pseudodon_mekongi **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 2 | 1 | -- | +----- Pilsbryoconcha_exilis
--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pilsbryoconcha_carinifera
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 1 | 1 | -- | +-- Pseudodon_vondembuschianus **
--+
+-- Pilsbryoconcha
Wu et al., 2024 [Publ 4350], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- | +-------- Pilsbryoconcha
--+
| +----- Pseudodon_mekongi **
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 14 | 6 | -- | +----- Pilsbryoconcha
+--+
| | +-- Namkongnaia
| +--+
| +-- Pseudodon_mekongi **
--+
| +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 10 | 6 | -- | +----------- Bineurus
|
| +----- Namkongnaia
--+ +--+
| | | +-- Sundadontina
| | +--+
+--+ +-- Nyeinchanconcha
|
| +----- Pseudodon_mekongi **
+--+
| +-- Pilsbryoconcha
+--+
+-- Thaiconcha
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- |
|
