The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Ptychobranchus Simpson, 1900: cladomics
6 Recent valid species (8 total species) Cladomics Summarynumber of species = 6 (8 total)
analyzed in 31 cladograms in 18 publications
maximum number of terminals in an analysis =
5 (mean = 1.13, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 1
maximum number of included terminals = 5, (mean =
5 median = 5)
proportion of cladograms that support basic monophyly
= 1
Roe, 2013 [Publ 3571], fig. 4 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 6 | 6 | 5 | 0 | +----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Medionidus
| |
+--+ +-- Cyprogenia
| +--------------+
| | +-- Dromus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Ptychobranchus_jonesi **
+--+
| +-------- Ptychobranchus_greenii **
+--+
| +----- Ptychobranchus_occidentalis__2 **
+--+
| +-- Ptychobranchus_occidentalis__1 **
+--+
+-- Ptychobranchus_fasciolaris **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- | +-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus_fasciolaris **
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 13 | 11 | -- | +-------------- Potamilus_leptodon
|
| +----------- Potamilus_alatus
--+ |
| | +-------- Venustaconcha
| | |
+--+ +-------- Leaunio
| |
| | +-- Ellipsaria
| +-----+
+--+ +-- Truncilla
|
| +-- Medionidus
+-----+
| +-- Ptychobranchus_subtentus **
|
| +----- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +-- Villosa
|
+-- Ellipsaria
|
--+-- Lampsilis
|
+-- Potamilus
|
+-- Ptychobranchus_subtentus **
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 8 | 6 | -- | +-------------- Potamilus_fragilis
|
--+ +----------- Ptychobranchus_subtentus **
| |
+--+ +-------- Medionidus
| |
+--+ +----- Potamilus_purpuratus
| |
+--+----- Lampsilis
|
| +-- Ellipsaria
+--+
+-- Villosa
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +-------------- Obliquaria
|
+-------------- Truncilla
--+
+-------------- Ptychobranchus_fasciolaris **
|
| +----------- Ligumia
+--+
| +-------- Cambarunio
+--+
| +----- Epioblasma
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 8 | 6 | -- | +-------------------- Obliquaria
--+
| +----------------- Medionidus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Potamilus_fragilis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ellipsaria
+--+
| +-- Villosa
+--+
+-- Lampsilis
Serb, 2006 [Publ 2678], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Serb, 2006 [Publ 2678], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 10 | 7 | -- | +----- Ptychobranchus_subtentus **
+--------+
| | +-- Potamilus_fragilis
| +--+
| +-- Medionidus
--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- | +-------------------- Truncilla
|
--+ +----------------- Ligumia
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----------+
| | +-- Sagittunio
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_fasciola
|
| +----- Epioblasma
+--+
| +-- Cambarunio
+--+
+-- Leaunio
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus_fasciolaris **
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- | +-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus_fasciolaris **
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- | +----------------------- Cyrtonaias
|
--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +----------------- Ligumia
| |
+--+ +-------------- Potamilus
| |
+--+ +----------- Pachynaias
| |
| | +-- Epioblasma
+--+ +-----+
| | +-- Truncilla
| |
+--+ +-- Mesonaias
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- | +-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- | +----- Truncilla
|
+----- Ptychobranchus_fasciolaris **
--+
| +-- Ortmanniana
| |
| +-- Cambarunio
+--+
+-- Ligumia
|
+-- Epioblasma
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +----------------- Medionidus
--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus_subtentus **
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis
+--+
| +-- Villosa
+--+
+-- Ellipsaria
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 13 | 11 | -- | +----- Ellipsaria
|
+----- Potamilus_leptodon
|
+----- Medionidus
|
+----- Potamilus_alatus
|
+----- Ptychobranchus_subtentus **
--+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Serb, 2006 [Publ 2678], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------- Obliquaria
|
| +-- Ptychobranchus_fasciolaris **
+-----------+
--+ +-- Lampsilis
|
| +-- Cyprogenia_aberti__6
| +--------+
| | +-- Dromus
+--+
| +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_aberti__4
| |
| | +-- Cyprogenia_stegaria__1
+--+ |
| +-- Cyprogenia_stegaria__2
| |
+--+-- Cyprogenia_aberti__1
|
+-- Cyprogenia_aberti__2
|
+-- Cyprogenia_aberti__3
Chapman et al., 2008 [Publ 2744], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- | +-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus_fasciolaris **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- | +-- Cambarunio
--+
+-- Ptychobranchus_fasciolaris **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 13 | 11 | -- | +-------------------- Potamilus_leptodon
|
--+ +----------------- Potamilus_alatus
| |
| | +-- Medionidus
+--+ +-----------+
| | +-- Ptychobranchus_subtentus **
| |
+--+ +-- Ellipsaria
| +--------+
| | +-- Truncilla
+--+
| +-------- Leaunio
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- | +----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Serb & Barnhart, 2008 [Publ 2674], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------------- Lampsilis
|
--+ +----------------------- Obliquaria
| |
+--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__4
| |
+--+ +----------- Cyprogenia_aberti__3
| |
+--+ +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Serb & Barnhart, 2008 [Publ 2674], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__5
| |
+--+ +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Chapman et al., 2008 [Publ 2667], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 11 | 8 | -- | +----------- Ligumia
|
+----------- Truncilla
|
| +-- Ptychobranchus_fasciolaris **
+--------+
--+ +-- Sagittunio
|
| +-- Lampsilis_siliquoidea
| +-----+
| | +-- Epioblasma
| |
+--+ +----- Ortmanniana
| |
| +----- Cambarunio
+--+
+----- Leaunio
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
|
