The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Obliquaria Rafinesque, 1820: cladomics
1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total)
analyzed in 55 cladograms in 34 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 92 | 26 | -- | Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- | Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- | +----------------- Obliquaria_reflexa **
|
| +----- Potamilus
--+ +--------+
| | | +-- Dromus
| | +--+
+--+ +-- Lemiox
|
| +----------- Lampsilis_ovata
| |
+--+ +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
Roe & Lydeard, 1998 [Publ 2693], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 8 | 2 | -- | Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- | +-------- Obliquaria_reflexa **
--+
| +----- Potamilus
+--+
| +-- Lampsilis
+--+
+-- Truncilla
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- | Chapman et al., 2008 [Publ 2744], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- | +-------- Obliquaria_reflexa **
--+
| +----- Potamilus
+--+
| +-- Lampsilis
+--+
+-- Truncilla
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 15 | 9 | -- | Freeman et al., 2025 [Publ 4443], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 2 | 2 | -- | +----- Obliquaria_reflexa **
--+
| +-- Glebula
+--+
+-- Cyrtonaias
Serb, 2006 [Publ 2678], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------- Obliquaria_reflexa **
|
| +-- Ptychobranchus
+-----------+
--+ +-- Lampsilis
|
| +-- Cyprogenia_aberti__6
| +--------+
| | +-- Dromus
+--+
| +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_aberti__4
| |
| | +-- Cyprogenia_stegaria__1
+--+ |
| +-- Cyprogenia_stegaria__2
| |
+--+-- Cyprogenia_aberti__1
|
+-- Cyprogenia_aberti__2
|
+-- Cyprogenia_aberti__3
Serb, 2006 [Publ 2678], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria_reflexa **
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- | Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 8 | 6 | -- | +-------------------- Obliquaria_reflexa **
--+
| +----------------- Medionidus
+--+
| +-------------- Ptychobranchus
+--+
| +----------- Potamilus_fragilis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ellipsaria
+--+
| +-- Villosa
+--+
+-- Lampsilis
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- | +-------------- Potamilus_alatus
|
+-------------- Potamilus_capax
|
+-------------- Glebula
|
| +-- Toxolasma
+-----------+
--+ +-- Paetulunio
|
| +-- Dromus
+-----------+
| +-- Lemiox
|
| +-- Lampsilis_ovata
| +--------+
| | +-- Obliquaria_reflexa **
+--+
| +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
Porto-Hannes et al., 2019 [Publ 3969], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 16 | 5 | -- | Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 2 | 1 | -- | Freeman et al., 2025 [Publ 4443], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 3 | 3 | -- | Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +----- Obliquaria_reflexa **
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 3 | 3 | -- | +-------- Obliquaria_reflexa **
--+
| +----- Truncilla
+--+
| +-- Lampsilis
+--+
+-- Potamilus
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- | +-- Lampsilis
--+
+-- Obliquaria_reflexa **
Szumowski et al., 2012 [Publ 3569], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- | +----------------------- Ellipsaria
|
--+ +-------------------- Truncilla
| |
+--+ +----------------- Obliquaria_reflexa **
| |
| | +-------------- Potamilus
+--+ |
| | +----- Lampsilis_ornata
| | +-----+
| | | | +-- Lampsilis_cardium
+--+ | +--+
| | +-- Lampsilis_ovata
| |
| | +-------- Lampsilis_teres
| +--+
+--+ | +----- Lampsilis_siliquoidea
| +--+
| | +-- Ortmanniana
| +--+
| +-- Lampsilis_higginsii
|
| +----- Hamiota
| |
+-----+ +-- Epioblasma
| |
+--+-- Cambarunio
|
+-- Ligumia
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 10 | 7 | -- | Roe et al., 2002 [Publ 2689], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 9 | 4 | -- | +----- Ligumia
|
+----- Obliquaria_reflexa **
--+
+----- Lampsilis
|
| +-- Villosa
+--+
+-- Hamiota
Smith et al., 2020 [Publ 4129], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria_reflexa **
|
--+ +----------- Cyrtonaias
| |
| | +-- Pachynaias
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- | Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- | +----------- Truncilla
|
--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma
+--+
| +----- Ligumia
+--+
| +-- Obliquaria_reflexa **
+--+
+-- Ortmanniana
Porto-Hannes et al., 2019 [Publ 3969], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 16 | 5 | -- | +----------------------- Obliquaria_reflexa **
|
| +-- Cambarunio
| +-----------------+
| | +-- Leaunio
--+ |
| | +----------- Lampsilis_teres
| | |
| +--------+ +-------- Ortmanniana
| | | |
| | +--+ +-- Lampsilis_hydiana
+--+ | +--+
| | | +-- Lampsilis_virescens
| +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Lampsilis_radiata
|
| +----------------- Obovaria
| |
+--+ +-------------- Lampsilis_bracteata
| |
+--+ +----------- Lampsilis_ornata
| |
+--+ +-------- Lampsilis_ovata
| |
+--+ +----- Lampsilis_cardium__3
| |
+--+ +-- Lampsilis_satura
| |
+--+-- Lampsilis_cardium__1
|
+-- Lampsilis_cardium__2
Serb & Barnhart, 2008 [Publ 2674], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------------- Lampsilis
|
--+ +----------------------- Obliquaria_reflexa **
| |
+--+ +-------------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__4
| |
+--+ +----------- Cyprogenia_aberti__3
| |
+--+ +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- | +-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria_reflexa **
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Roe, 2013 [Publ 3571], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 10 | 5 | -- | +----------- Obliquaria_reflexa **
|
--+----------- Lampsilis
|
| +-------- Medionidus
+--+
| +----- Ptychobranchus
+--+
| +-- Cyprogenia
+--+
+-- Dromus
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 13 | 11 | -- | Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 13 | 11 | -- | Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +-------------- Obliquaria_reflexa **
|
+-------------- Truncilla
--+
+-------------- Ptychobranchus
|
| +----------- Ligumia
+--+
| +-------- Cambarunio
+--+
| +----- Epioblasma
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Smith et al., 2020 [Publ 4129], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria_reflexa **
|
--+ +----------- Cyrtonaias
| |
| | +-- Pachynaias
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria_reflexa **
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- | Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria_reflexa **
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 15 | 9 | -- | +----------- Toxolasma
|
| +-------- Obliquaria_reflexa **
--+ |
| | +----- Obovaria
| | |
| | +----- Lampsilis_cardium
+--+ |
| +----- Potamilus
| |
| | +-- Epioblasma
| +--+
| | +-- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana
+--+
| +-- Lampsilis_higginsii
|
| +-- Ligumia
+--+
| +-- Lampsilis_teres
|
| +-- Ellipsaria
+--+
+-- Truncilla
Chapman et al., 2008 [Publ 2667], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria_reflexa **
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- | Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 10 | 7 | -- | Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 8 | 6 | -- | Serb, 2006 [Publ 2678], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +-------------------- Obliquaria_reflexa **
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- | +-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria_reflexa **
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- | Freeman et al., 2025 [Publ 4443], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 3 | 3 | -- | +-------- Toxolasma
--+
| +----- Obliquaria_reflexa **
+--+
| +-- Glebula
+--+
+-- Cyrtonaias
Serb & Barnhart, 2008 [Publ 2674], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 4 | -- | +----------------------- Obliquaria_reflexa **
|
+----------------------- Lampsilis
--+
| +-------------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__5
| |
+--+ +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Porto-Hannes et al., 2021 [Publ 4227], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 8 | 3 | -- | +-- Obovaria
+--------+
| +-- Ortmanniana
|
| +-- Obliquaria_reflexa **
+--------+
--+ +-- Lampsilis_ornata
|
| +----- Lampsilis_radiata__2
| +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_radiata__1
|
| +----- Lampsilis_virescens
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria_reflexa **
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria_reflexa **
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
|
