The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Cyrtonaias Crosse & Fischer, 1894: cladomics
4 Recent valid species (4 total species) Cladomics Summarynumber of species = 4 (4 total)
analyzed in 42 cladograms in 21 publications
maximum number of terminals in an analysis =
2 (mean = 1.07, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 3
maximum number of included terminals = 2, (mean =
2 median = 2)
proportion of cladograms that support basic monophyly
= 0
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Pfeiffer et al., 2019 [Publ 3866], fig. 1 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 13 | 0 | 2 | 33 | 10 | 10 | 0 | +-- Cyrtonaias_tampicoensis **
+--------------------+
| +-- Glebula
|
--+ +-------------------- Potamilus
| |
| | +-- Truncilla
+--+ +--------------+
| | +-- Ptychobranchus
| |
+--+ +-------------- Pachynaias
| |
| | +-- Cyrtonaias_explicata **
+--+ +--------+
| | +-- Sphenonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- | +-- Cyrtonaias_tampicoensis **
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 13 | 0 | 2 | 24 | 10 | 10 | 5 | +-- Glebula
+-----------+
| +-- Cyrtonaias_tampicoensis **
|
--+ +----------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +-----+
| | +-- Truncilla
| |
| | +----- Ortmanniana
+--+ |
| +----- Epioblasma
| |
| +----- Lampsilis
| |
+--+----- Ligumia
|
+----- Cambarunio
|
+----- Pachynaias
|
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 12 | 0 | 2 | 23 | 9 | 9 | 16 | +----------------------------- Cyrtonaias_tampicoensis **
|
--+ +-------------------------- Ptychobranchus
| |
+--+ +----------------------- Ligumia
| |
+--+ +-------------------- Potamilus
| |
+--+ +----------------- Pachynaias
| |
| | +-- Epioblasma
+--+ +-----------+
| | +-- Truncilla
| |
+--+ +-- Ortmanniana
| +--------+
| | +-- Lampsilis
+--+
| +-------- Cambarunio
+--+
| +----- Sphenonaias_microdon__2
+--+
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon__1
Hoeh et al., 2002 [Publ 2673], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- | +----------- Potamilus
--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- | +----------- Cyrtonaias_tampicoensis **
--+
| +-------- Potamilus
+--+
| +----- Ligumia
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria
--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- | +----------- Potamilus
--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Smith et al., 2020 [Publ 4129], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria
|
--+ +----------- Cyrtonaias_tampicoensis **
| |
| | +-- Pachynaias
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria
--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +-- Glebula
--+
+-- Cyrtonaias_tampicoensis **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- | -- Cyrtonaias_tampicoensis **
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 5 | 5 | -- | +-- Cyrtonaias_tampicoensis **
+-----+
| +-- Glebula
|
--+ +-- Lampsilis
| +--+
| | +-- Leaunio
+--+
| +-- Truncilla
+--+
+-- Potamilus
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- | +-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias_tampicoensis **
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Campbell et al., 2008 [Publ 2675], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- | -- Cyrtonaias_tampicoensis **
Walker et al., 2006 [Publ 2666], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 4 | 4 | -- | +----------- Potamilus
--+
| +-------- Cyrtonaias_tampicoensis **
+--+
| +----- Ligumia
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Smith et al., 2020 [Publ 4129], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria
|
--+ +----------- Cyrtonaias_tampicoensis **
| |
| | +-- Pachynaias
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 5 | 5 | -- | +-- Glebula
--+
+-- Cyrtonaias_tampicoensis **
Campbell et al., 2008 [Publ 2675], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- | -- Cyrtonaias_tampicoensis **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- | -- Cyrtonaias_tampicoensis **
Freeman et al., 2025 [Publ 4443], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 2 | 2 | -- | +-- Glebula
--+
+-- Cyrtonaias_tampicoensis **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 42 | 4 | 3 | -- | -- Cyrtonaias_tampicoensis **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 4 | 3 | -- | -- Cyrtonaias_tampicoensis **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias_tampicoensis **
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Hoeh et al., 2002 [Publ 2673], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- | +----------- Cyrtonaias_tampicoensis **
--+
| +-------- Ortmanniana
+--+
| +----- Lampsilis
+--+
| +-- Potamilus
+--+
+-- Ligumia
Walker et al., 2006 [Publ 2666], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 4 | -- | -- Cyrtonaias_tampicoensis **
Chapman et al., 2008 [Publ 2744], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Campbell et al., 2008 [Publ 2675], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | -- Cyrtonaias_tampicoensis **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- | -- Cyrtonaias_tampicoensis **
Burlakova et al., 2012 [Publ 3486], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- | -- Cyrtonaias_tampicoensis **
Chapman et al., 2008 [Publ 2667], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 12 | 10 | -- | +-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias_tampicoensis **
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- | -- Cyrtonaias_tampicoensis **
Freeman et al., 2025 [Publ 4443], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 2 | 2 | -- | +----- Obliquaria
--+
| +-- Glebula
+--+
+-- Cyrtonaias_tampicoensis **
Freeman et al., 2025 [Publ 4443], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | -- Cyrtonaias_explicata **
Freeman et al., 2025 [Publ 4443], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 3 | 3 | -- | +-------- Toxolasma
--+
| +----- Obliquaria
+--+
| +-- Glebula
+--+
+-- Cyrtonaias_tampicoensis **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- | +-- Cyrtonaias_tampicoensis **
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Ortmanniana
+--+
| +-- Ligumia
+--+
+-- Potamilus
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +----- Obliquaria
+--------+
| | +-- Cyrtonaias_tampicoensis **
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- | +-- Cyrtonaias_tampicoensis **
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Ortmanniana
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- | +----------- Cyrtonaias_tampicoensis **
--+
| +-------- Potamilus
+--+
| +----- Ligumia
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Campbell et al., 2008 [Publ 2675], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | -- Cyrtonaias_tampicoensis **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- | +-- Glebula
--+
+-- Cyrtonaias_tampicoensis **
Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- | +-- Cyrtonaias_tampicoensis **
--+
+-- Glebula
|
