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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Aculamprotula Wu, Liang, Wang & Ouyang, 1999: cladomics
     9 Recent valid species (9 total species)

Cladomics Summary

number of species = 9 (9 total)
analyzed in 103 cladograms in 57 publications
maximum number of terminals in an analysis = 5 (mean = 2.62, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 61
maximum number of included terminals = 5, (mean = 3.74 median = 4)
proportion of cladograms that support basic monophyly = 1


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Dai et al., 2025 [Publ 4440], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
55354017100
  +-------------------- Globunio                
  |                                             
  |           +-------- Aculamprotula_tortuosa **  
  |           |                                 
  |  +--------+     +-- Aculamprotula_fibrosa **   
--+  |        |  +--+                           
  |  |        |  |  +-- Aculamprotula_coreana **   
  |  |        +--+                              
  |  |           |  +-- Aculamprotula_scripta **   
  |  |           +--+                           
  +--+              +-- Aculamprotula_polysticta **
     |                                          
     |              +-- Middendorffinaia        
     |  +-----------+                           
     |  |           +-- Nodularia               
     |  |                                       
     +--+  +----------- Cuneopsis               
        |  |                                    
        |  |        +-- Diaurora                
        +--+--------+                           
           |        +-- Schistodesmus           
           |                                    
           |  +-------- Pseudocuneopsis         
           +--+                                 
              |  +----- Inversiunio             
              +--+                              
                 |  +-- Tchangsinaia            
                 +--+                           
                    +-- Unio                    

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
553544962
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_tientsinensis **
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Schistodesmus              

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
55254017101
  +----------------------- Globunio                
  |                                                
  |              +-------- Aculamprotula_tortuosa **  
  |              |                                 
--+  +-----------+  +----- Aculamprotula_coreana **   
  |  |           |  |                              
  |  |           +--+----- Aculamprotula_fibrosa **   
  |  |              |                              
  |  |              |  +-- Aculamprotula_polysticta **
  +--+              +--+                           
     |                 +-- Aculamprotula_scripta **   
     |                                             
     |  +----------------- Middendorffinaia        
     |  |                                          
     +--+  +-------------- Nodularia               
        |  |                                       
        +--+  +----------- Cuneopsis               
           |  |                                    
           |  |        +-- Diaurora                
           +--+--------+                           
              |        +-- Schistodesmus           
              |                                    
              |  +-------- Pseudocuneopsis         
              +--+                                 
                 |  +----- Unio                    
                 +--+                              
                    |  +-- Tchangsinaia            
                    +--+                           
                       +-- Inversiunio             

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
553548952
     +----------- Aculamprotula_tortuosa **     
  +--+                                       
  |  |  +-------- Aculamprotula_scripta **      
  |  +--+                                    
  |     |  +----- Aculamprotula_coreana **      
  |     +--+                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |     +-------- Unio                       
  |     |                                    
  +-----+     +-- Nodularia                  
        |  +--+                              
        |  |  +-- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5535482462
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_tientsinensis **
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
5535681562
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_tientsinensis **
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_coreana **      
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5535714472
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+     +-- Aculamprotula_coreana **      
  |           |  +--+                              
  |           |  |  +-- Aculamprotula_tientsinensis **
  |           +--+                                 
--+              |  +-- Aculamprotula_polysticta **   
  |              +--+                              
  |                 +-- Aculamprotula_scripta **      
  |                                                
  |  +----------------- Unio                       
  +--+                                             
     |  +-------------- Middendorffinaia           
     +--+                                          
        |  +----------- Pseudocuneopsis            
        +--+                                       
           |  +-------- Nodularia                  
           +--+                                    
              |  +----- Schistodesmus              
              +--+                                 
                 |  +-- Tchangsinaia               
                 +--+                              
                    +-- Cuneopsis                  

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
55351941454
  +-------- Aculamprotula_tortuosa **     
  |                                    
--+     +-- Aculamprotula_coreana **      
  |  +--+                              
  |  |  +-- Aculamprotula_scripta **      
  +--+                                 
     |  +-- Aculamprotula_fibrosa **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
5535351562
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_polysticta **   
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_scripta **      
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
55251941453
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+  +----- Aculamprotula_coreana **      
  |     |  |                                 
  |     +--+----- Aculamprotula_scripta **      
  |        |                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Schistodesmus              
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5535901762
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+        +-- Nodularia                  
     |  +-----+                              
     |  |     +-- Pseudocuneopsis            
     +--+                                    
        |  +----- Tchangsinaia               
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
5535631062
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
5535681062
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Schistodesmus              
              +--+                              
                 +-- Cuneopsis                  

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5535861762
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+  +-------- Nodularia                  
     |  |                                    
     +--+-------- Pseudocuneopsis            
        |                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
55351921354
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_scripta **      
  |     +--+                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
5535681062
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Tchangsinaia               

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
5535331872
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+     +-- Aculamprotula_tientsinensis **
  |           |  +--+                              
  |           |  |  +-- Aculamprotula_coreana **      
  |           +--+                                 
--+              |  +-- Aculamprotula_polysticta **   
  |              +--+                              
  |                 +-- Aculamprotula_scripta **      
  |                                                
  |  +----------------- Unio                       
  +--+                                             
     |  +-------------- Pseudocuneopsis            
     +--+                                          
        |  +----------- Nodularia                  
        +--+                                       
           |  +-------- Schistodesmus              
           +--+                                    
              |  +----- Cuneopsis                  
              +--+                                 
                 |  +-- Tchangsinaia               
                 +--+                              
                    +-- Inversiunio                

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5535381042
  +----------- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Aculamprotula_scripta **      
  +--+                                    
     |  +----- Aculamprotula_coreana **      
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55351162272
        +----------- Aculamprotula_tortuosa **     
  +-----+                                       
  |     |  +-------- Aculamprotula_scripta **      
  |     +--+                                    
  |        |  +----- Aculamprotula_coreana **      
  |        +--+                                 
--+           |  +-- Aculamprotula_fibrosa **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Middendorffinaia           
  |  |                                          
  +--+  +----------- Unio                       
     |  |                                       
     +--+  +-------- Nodularia                  
        |  |                                    
        +--+  +----- Schistodesmus              
           |  |                                 
           +--+----- Cuneopsis                  
              |                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Inversiunio                

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
55354214
  +-------------- Nodularia                  
--+                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_scripta **      
           +--+                              
              +-- Aculamprotula_coreana **      

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5535401044
        +-------- Nodularia                  
  +-----+                                    
  |     |  +----- Tchangsinaia               
  |     +--+                                 
  |        |  +-- Schistodesmus              
--+        +--+                              
  |           +-- Cuneopsis                  
  |                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_coreana **      
           +--+                              
              +-- Aculamprotula_scripta **      

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
5535832482
  +----------------- Globunio                   
  |                                             
  |        +-------- Aculamprotula_tortuosa **     
  |        |                                    
--+  +-----+     +-- Aculamprotula_polysticta **   
  |  |     |  +--+                              
  |  |     |  |  +-- Aculamprotula_scripta **      
  |  |     +--+                                 
  |  |        |  +-- Aculamprotula_coreana **      
  +--+        +--+                              
     |           +-- Aculamprotula_tientsinensis **
     |                                          
     |  +----------- Unio                       
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Pseudocuneopsis            
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5535861762
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+  +-------- Nodularia                  
     |  |                                    
     +--+-------- Pseudocuneopsis            
        |                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5535901762
              +-------- Aculamprotula_tortuosa **      
              |                                     
  +-----------+     +-- Aculamprotula_coreana **       
  |           |  +--+                               
  |           |  |  +-- Aculamprotula_tientsinensis ** 
  |           +--+                                  
  |              |  +-- Aculamprotula_polysticta **    
--+              +--+                               
  |                 +-- Aculamprotula_scripta **       
  |                                                 
  |  +----------------- Unio                        
  |  |                                              
  +--+  +-------------- Pseudocuneopsis             
     |  |                                           
     +--+  +----------- Nodularia                   
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus__1
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus__2
                 +--+                               
                    +-- Schistodesmus_spinosus      

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
442442963
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
--+           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Schistodesmus              

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
442430113
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442452743
     +-------- Aculamprotula_tortuosa **     
  +--+                                    
  |  |  +----- Aculamprotula_scripta **      
  |  +--+                                 
  |     |  +-- Aculamprotula_tientsinensis **
--+     +--+                              
  |        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus              

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
442425663
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
--+           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Tchangsinaia               

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
442430113
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **      

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442410113
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **      

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442452853
  +-------------- Middendorffinaia           
  |                                          
  |     +-------- Aculamprotula_tortuosa **     
--+  +--+                                    
  |  |  |  +----- Aculamprotula_scripta **      
  |  |  +--+                                 
  |  |     |  +-- Aculamprotula_tientsinensis **
  +--+     +--+                              
     |        +-- Aculamprotula_coreana **      
     |                                       
     |  +-------- Unio                       
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis                  

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
442418223
           +-- Unio                       
  +--------+                              
  |        +-- Nodularia                  
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_coreana **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
331341744
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus              

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
331388854
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Cuneopsis                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus              

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
331335744
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_coreana **      
  |     +--+                              
--+        +-- Aculamprotula_tientsinensis **
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Nodularia                  
        +--+                              
           +-- Tchangsinaia               

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3313651054
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Inversiunio                
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Tchangsinaia               

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
331337854
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_coreana **      
  |        +--+                              
--+           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Tchangsinaia               
        +--+                                 
           |  +-- Cuneopsis                  
           +--+                              
              +-- Schistodesmus              

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
331339634
     +----- Aculamprotula_tortuosa **     
  +--+                                 
  |  |  +-- Aculamprotula_coreana **      
  |  +--+                              
--+     +-- Aculamprotula_tientsinensis **
  |                                    
  |  +----- Unio                       
  +--+                                 
     |  +-- Tchangsinaia               
     +--+                              
        +-- Nodularia                  

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3313651054
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Inversiunio                
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Tchangsinaia               

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3313651054
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Inversiunio                
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus              

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331315224
  +----- Aculamprotula_tortuosa **     
--+                                 
  |  +-- Aculamprotula_tientsinensis **
  +--+                              
     +-- Aculamprotula_scripta **      

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
331318434
        +-- Tchangsinaia               
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_tientsinensis **
     +--+                              
        +-- Aculamprotula_scripta **      

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
331318224
        +-- Unio                       
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_coreana **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
331388854
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_scripta **      
  |        +--+                              
--+           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Cuneopsis                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus              

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3313191124
  +----- Aculamprotula_tortuosa **     
--+                                 
  |  +-- Aculamprotula_scripta **      
  +--+                              
     +-- Aculamprotula_tientsinensis **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
331328224
        +-- Unio                       
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_tientsinensis **
     +--+                              
        +-- Aculamprotula_scripta **      

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
331320744
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Nodularia                  

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
331345744
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Schistodesmus              
        +--+                              
           +-- Tchangsinaia               

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
331341744
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus              

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
331350964
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+        +-- Schistodesmus              
     |  +-----+                              
     |  |     +-- Cuneopsis                  
     +--+                                    
        |  +----- Pseudocuneopsis            
        +--+                                 
           |  +-- Nodularia                  
           +--+                              
              +-- Tchangsinaia               

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22023444--
           +-- Aculamprotula_tientsinensis **
  +--------+                              
  |        +-- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Nodularia                  
        +--+                              
           +-- Tchangsinaia               

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
22023211--
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
220222127--
  +----------------- Unio                       
  |                                             
--+              +-- Aculamprotula_scripta **      
  |  +-----------+                              
  |  |           +-- Aculamprotula_tientsinensis **
  +--+                                          
     |  +----------- Cuneopsis                  
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Schistodesmus              
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Inversiunio                
              +--+                              
                 +-- Pseudocuneopsis            

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
220221117--
  +----------------- Unio                       
  |                                             
--+              +-- Aculamprotula_scripta **      
  |  +-----------+                              
  |  |           +-- Aculamprotula_tientsinensis **
  +--+                                          
     |  +----------- Cuneopsis                  
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Schistodesmus              
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Inversiunio                
              +--+                              
                 +-- Pseudocuneopsis            

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
22023054--
           +-- Aculamprotula_tortuosa **
  +--------+                         
  |        +-- Aculamprotula_coreana ** 
--+                                  
  |  +-------- Unio                  
  +--+                               
     |  +----- Schistodesmus         
     +--+                            
        |  +-- Tchangsinaia          
        +--+                         
           +-- Nodularia             

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
220250156--
                 +-- Aculamprotula_scripta **      
  +--------------+                              
  |              +-- Aculamprotula_tientsinensis **
--+                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis                  

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
22021411--
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
220222127--
  +-------- Unio                       
  |                                    
  |     +-- Aculamprotula_tientsinensis **
  +-----+                              
  |     +-- Aculamprotula_scripta **      
--+                                    
  |  +----- Cuneopsis                  
  |  |                                 
  |  |  +-- Nodularia                  
  |  +--+                              
  +--+  +-- Schistodesmus              
     |                                 
     |  +-- Tchangsinaia               
     |  |                              
     +--+-- Inversiunio                
        |                              
        +-- Pseudocuneopsis            

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22021422--
     +-- Nodularia                  
  +--+                              
  |  +-- Tchangsinaia               
--+                                 
  |  +-- Aculamprotula_tientsinensis **
  +--+                              
     +-- Aculamprotula_tortuosa **     

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
22022954--
           +-- Aculamprotula_tortuosa **
  +--------+                         
  |        +-- Aculamprotula_coreana ** 
--+                                  
  |  +-------- Unio                  
  +--+                               
     |  +----- Schistodesmus         
     +--+                            
        |  +-- Nodularia             
        +--+                         
           +-- Tchangsinaia          

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
22023644--
  +-- Aculamprotula_tientsinensis **
--+                              
  +-- Aculamprotula_tortuosa **     

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012242--
  +----- Schistodesmus         
--+                            
  |  +-- Aculamprotula_tortuosa **
  +--+                         
     +-- Unio                  

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013074--
  +----------- Aculamprotula_tientsinensis **
--+                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Cuneopsis                  

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113895--
  +-------------- Unio                  
--+                                     
  |  +----------- Aculamprotula_tortuosa **
  +--+                                  
     |  +-------- Nodularia             
     +--+                               
        |  +----- Cuneopsis             
        +--+                            
           |  +-- Tchangsinaia          
           +--+                         
              +-- Schistodesmus         

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012411--
-- Aculamprotula_tortuosa **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011941--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015266--
  +-------------- Aculamprotula_tientsinensis **
  |                                          
--+  +----------- Unio                       
  |  |                                       
  +--+        +-- Pseudocuneopsis            
     |  +-----+                              
     |  |     +-- Nodularia                  
     +--+                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Cuneopsis                  
           +--+                              
              +-- Tchangsinaia               

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012111--
-- Aculamprotula_tortuosa **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
110116104--
-- Aculamprotula_tientsinensis **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
110139169--
  +----------------- Aculamprotula_fibrosa **
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  +--+  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113895--
  +----------- Unio                  
  |                                  
--+  +-------- Aculamprotula_tortuosa **
  |  |                               
  |  +-------- Schistodesmus         
  +--+                               
     +-------- Nodularia             
     |                               
     |  +----- Cuneopsis_szechenyii  
     +--+                            
        |  +-- Tchangsinaia          
        +--+                         
           +-- Cuneopsis_rufescens   

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
110139169--
  +----------------- Aculamprotula_fibrosa **
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  +--+  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
110111116--
  +----------------- Aculamprotula_tientsinensis **
--+                                             
  |  +-------------- Nodularia                  
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Schistodesmus              
        +--+                                    
           |  +----- Cuneopsis                  
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Inversiunio                

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017595--
  +----------------- Unio                  
--+                                        
  |  +-------------- Aculamprotula_tortuosa **
  +--+                                     
     |  +----------- Nodularia             
     +--+                                  
        |  +-------- Schistodesmus         
        +--+                               
           |  +----- Cuneopsis_szechenyii  
           +--+                            
              |  +-- Tchangsinaia          
              +--+                         
                 +-- Cuneopsis_rufescens   

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012142--
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Schistodesmus         
  +--+                         
     +-- Unio                  

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
110115116--
-- Aculamprotula_tientsinensis **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011421--
-- Aculamprotula_tientsinensis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1101611--
  +-- Unio                  
--+                         
  +-- Aculamprotula_tortuosa **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011222--
-- Aculamprotula_tortuosa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015386--
  +----------------- Aculamprotula_tortuosa **
--+                                        
  |  +-------------- Unio                  
  +--+                                     
     |  +----------- Pseudocuneopsis       
     +--+                                  
        |  +-------- Nodularia             
        +--+                               
           |  +----- Tchangsinaia          
           +--+                            
              |  +-- Cuneopsis             
              +--+                         
                 +-- Schistodesmus         

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011521--
-- Aculamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013364--
-- Aculamprotula_tortuosa **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011295--
-- Aculamprotula_scripta **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112495--
  +-- Unio_crassus          
  |                         
  +-- Unio_pictorum         
  |                         
  +-- Unio_tumidus          
  |                         
  +-- Aculamprotula_tortuosa **
  |                         
  +-- Cuneopsis_szechenyii  
--+                         
  +-- Tchangsinaia          
  |                         
  +-- Cuneopsis_rufescens   
  |                         
  +-- Nodularia_douglasiae  
  |                         
  +-- Nodularia_nuxpersicae 
  |                         
  +-- Schistodesmus         

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014042--
-- Aculamprotula_tortuosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017595--
  +----------------- Unio                  
--+                                        
  |  +-------------- Aculamprotula_tortuosa **
  +--+                                     
     |  +----------- Nodularia             
     +--+                                  
        |  +-------- Schistodesmus         
        +--+                               
           |  +----- Cuneopsis_szechenyii  
           +--+                            
              |  +-- Tchangsinaia          
              +--+                         
                 +-- Cuneopsis_rufescens   

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014042--
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Schistodesmus         
  +--+                         
     +-- Unio                  

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011221--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
110116116--
  +----------------- Aculamprotula_tortuosa **
--+                                        
  |  +-------------- Unio                  
  +--+                                     
     |  +----------- Pseudocuneopsis       
     +--+                                  
        |  +-------- Nodularia             
        +--+                               
           |  +----- Tchangsinaia          
           +--+                            
              |  +-- Cuneopsis             
              +--+                         
                 +-- Schistodesmus         

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
110179105--
  +-------------- Unio                  
--+                                     
  |  +----------- Aculamprotula_tortuosa **
  +--+                                  
     |  +-------- Nodularia             
     +--+                               
        |  +----- Schistodesmus         
        +--+                            
           |  +-- Tchangsinaia          
           +--+                         
              +-- Cuneopsis             

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011322--
  +-- Schistodesmus         
--+                         
  +-- Aculamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011631--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
  +-- Unio                       
--+                              
  +-- Aculamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
110120106--
  +----------------- Aculamprotula_scripta **
--+                                       
  |  +-------------- Unio                 
  +--+                                    
     |  +----------- Middendorffinaia     
     +--+                                 
        |  +-------- Nodularia            
        +--+                              
           |  +----- Cuneopsis            
           +--+                           
              |  +-- Inversiunio          
              +--+                        
                 +-- Schistodesmus        

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011221--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011421--
-- Aculamprotula_tientsinensis **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013063--
  +-------- Aculamprotula_tientsinensis **
--+                                    
  |  +----- Unio                       
  +--+                                 
     |  +-- Tchangsinaia               
     +--+                              
        +-- Nodularia                  

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
110120106--
-- Aculamprotula_scripta **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101941--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012411--
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio                  

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012142--
  +----- Schistodesmus         
--+                            
  |  +-- Aculamprotula_tortuosa **
  +--+                         
     +-- Unio                  

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Unio                  
--+                         
  +-- Aculamprotula_tortuosa **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011433--
  +-------- Aculamprotula_tortuosa **
--+                               
  |  +----- Schistodesmus         
  +--+                            
     |  +-- Nodularia             
     +--+                         
        +-- Unio                  
 
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