The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Villosa Frierson, 1927: cladomics
4 Recent valid species (4 total species) Cladomics Summarynumber of species = 4 (4 total)
analyzed in 15 cladograms in 11 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 0
maximum number of included terminals = , (mean =
median = )
proportion of cladograms that support basic monophyly
= 0
Campbell et al., 2005 [Publ 2660], fig. 1 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa_villosa **
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Buhay et al., 2002 [Publ 2661], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 3 | -- | +----- Lampsilis
|
--+----- Epioblasma
|
| +-- Cambarunio
+--+
+-- Villosa_villosa **
Roe et al., 2002 [Publ 2689], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 9 | 4 | -- | +----- Ligumia
|
+----- Obliquaria
--+
+----- Lampsilis
|
| +-- Villosa_villosa **
+--+
+-- Hamiota
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 8 | 6 | -- | +-------------- Potamilus_fragilis
|
--+ +----------- Ptychobranchus
| |
+--+ +-------- Medionidus
| |
+--+ +----- Potamilus_purpuratus
| |
+--+----- Lampsilis
|
| +-- Ellipsaria
+--+
+-- Villosa_delumbis **
Inoue et al., 2013 [Publ 3528], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 9 | 5 | -- | +-------- Lampsilis
|
--+ +----- Leaunio
| |
+--+----- Obovaria
|
| +-- Villosa_villosa **
+--+
+-- Cambarunio
Turner et al., 2000 [Publ 2722], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 8 | 2 | -- | +-------------------- Medionidus
|
--+ +----------------- Lampsilis_teres
| |
| +----------------- Villosa_delumbis **
+--+
| +-- Lampsilis_satura
| +-----------+
| | +-- Lampsilis_cardium
+--+
| +----------- Lampsilis_reeveiana
| |
+--+ +-------- Lampsilis_hydiana__1
| |
+--+-------- Lampsilis_hydiana__2
|
| +----- Lampsilis_siliquoidea__2
+--+
| +-- Lampsilis_siliquoidea__1
+--+
+-- Lampsilis_powellii
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +-- Villosa_delumbis **
|
+-- Ellipsaria
|
--+-- Lampsilis
|
+-- Potamilus
|
+-- Ptychobranchus
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 8 | 6 | -- | +-------------------- Obliquaria
--+
| +----------------- Medionidus
+--+
| +-------------- Ptychobranchus
+--+
| +----------- Potamilus_fragilis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ellipsaria
+--+
| +-- Villosa_delumbis **
+--+
+-- Lampsilis
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 10 | 7 | -- | +----- Ptychobranchus
+--------+
| | +-- Potamilus_fragilis
| +--+
| +-- Medionidus
--+
| +-- Villosa_delumbis **
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- | +----------------- Toxolasma
|
| +-------------- Potamilus
| |
| | +-- Ellipsaria
| +-----------+
--+ | +-- Truncilla
| |
| | +----- Medionidus
| +--------+
| | | +-- Lemiox
| | +--+
+--+ +-- Dromus
|
| +----------- Cambarunio
| |
| +----------- Leaunio
| |
| +----------- Villosa_villosa **
| |
| | +-- Sagittunio
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha
+-----+
| | +-- Epioblasma
| +--+
| +-- Obovaria
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Patterson et al., 2021 [Publ 4222], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa_villosa **
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 7 | 6 | -- | +----------------- Medionidus
--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis
+--+
| +-- Villosa_delumbis **
+--+
+-- Ellipsaria
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 42 | 22 | -- | +----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa_villosa **
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Buhay et al., 2002 [Publ 2661], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 6 | 3 | -- | +----- Lampsilis
|
--+----- Epioblasma
|
| +-- Cambarunio
+--+
+-- Villosa_villosa **
|
