The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Search the database pages:
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Pachynaias Crosse & Fischer, 1894: cladomics
1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total)
analyzed in 10 cladograms in 7 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 92 | 26 | -- | +-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
+--+ | +--+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias_spheniopsis **
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
+--+ | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Freeman et al., 2025 [Publ 4443], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | +-- Mesonaias
--+
+-- Pachynaias_spheniopsis **
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- | +----------------------- Cyrtonaias
|
--+ +-------------------- Ptychobranchus
| |
+--+ +----------------- Ligumia
| |
+--+ +-------------- Potamilus
| |
+--+ +----------- Pachynaias_spheniopsis **
| |
| | +-- Epioblasma
+--+ +-----+
| | +-- Truncilla
| |
+--+ +-- Mesonaias
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
MartÃnez et al., 2026 [Publ 4820], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 2 | 1 | -- | +----- Mesonaias_tampicoensis
--+
| +-- Mesonaias_explicata
+--+
+-- Pachynaias_spheniopsis **
Smith et al., 2020 [Publ 4129], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria
|
--+ +----------- Cyrtonaias
| |
| | +-- Pachynaias_spheniopsis **
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- | +-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias_spheniopsis **
|
+-- Mesonaias
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias_spheniopsis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +----------- Obliquaria
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio
+--+
| +-- Lampsilis
+--+
+-- Pachynaias_spheniopsis **
Smith et al., 2020 [Publ 4129], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 19 | 7 | -- | +-------------- Obliquaria
|
--+ +----------- Cyrtonaias
| |
| | +-- Pachynaias_spheniopsis **
+--+ +-----+
| | +-- Lampsilis
| |
+--+ +-- Ellipsaria
| +--+
| | +-- Truncilla
+--+
| +-- Atlanticoncha
+--+
+-- Potamilus
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- | +-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias_spheniopsis **
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
|
