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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Coelaturini

genus Coelatura Conrad, 1853: cladomics
     24 Recent valid species (25 total species)

Cladomics Summary

number of species = 24 (25 total)
analyzed in 50 cladograms in 32 publications
maximum number of terminals in an analysis = 18 (mean = 2.4, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 18, (mean = 4.68 median = 3)
proportion of cladograms that support basic monophyly = 0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
1011122018110
  +----------------- Coelatura_horei **          
  |                                           
  |              +-- Coelatura_gabonensis **     
  |     +--------+                            
  |     |        +-- Coelatura_aegyptiaca__4 **  
  |     |                                     
  |  +--+  +-------- Coelatura_kunenensis **     
--+  |  |  |                                  
  |  |  |  |  +----- Coelatura_luapulaensis__1 **
  |  |  +--+  |                               
  |  |     |  +----- Coelatura_luapulaensis__3 **
  |  |     |  |                               
  |  |     +--+----- Coelatura_luapulaensis__4 **
  |  |        |                               
  |  |        +----- Coelatura_choziensis__2 **  
  +--+        |                               
     |        |  +-- Coelatura_luapulaensis__2 **
     |        +--+                            
     |           +-- Coelatura_choziensis__1 **  
     |                                        
     |  +----------- Coelatura_alluaudi__2 **    
     |  |                                     
     |  +----------- Nitia                    
     |  |                                     
     |  |  +-------- Coelatura_aegyptiaca__1 **  
     +--+  |                                  
        |  +-------- Coelatura_bakeri__2 **      
        |  |                                  
        |  |     +-- Coelatura_aegyptiaca__2 **  
        |  +-----+                            
        +--+     +-- Coelatura_stuhlmanni__1 **  
           |                                  
           |     +-- Coelatura_stuhlmanni__2 **  
           +-----+                            
           |     +-- Coelatura_bakeri__1 **      
           |                                  
           |  +----- Coelatura_aegyptiaca__3 **  
           +--+                               
              |  +-- Coelatura_alluaudi__1 **    
              +--+                            
                 +-- Coelatura_hauttecoeuri **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
18211118189622
  +-------------------------------- Grandidieria                 
  |                                                              
  |                             +-- Pseudospatha                 
--+  +--------------------------+                                
  |  |                          +-- Moncetia                     
  |  |                                                           
  |  |  +-------------------------- Nitia                        
  +--+  |                                                        
     |  |                       +-- Coelatura_aegyptiaca **         
     |  |                       |                                
     |  |                       +-- Coelatura_alluaudi **           
     +--+                       |                                
        |  +--------------------+-- Coelatura_bakeri **             
        |  |                    |                                
        |  |                    +-- Coelatura_hauttecoeuri **       
        |  |                    |                                
        |  |                    +-- Coelatura_stuhlmanni **         
        +--+                                                     
           |                    +-- Coelatura_leopoldvillensis **   
           |  +-----------------+                                
           |  |                 +-- Coelatura_mesafricana **        
           |  |                                                  
           |  |                 +-- Coelatura_horei **              
           +--+                 |                                
              |  +--------------+-- Coelatura_ratidota **           
              |  |              |                                
              |  |              +-- Coelatura **                    
              +--+                                               
                 |  +-------------- Coelatura_gabonensis **         
                 |  |                                            
                 |  +-------------- Coelatura_stanleyvillensis **   
                 +--+                                            
                    |  +----------- Coelatura_kunenensis **         
                    |  |                                         
                    |  |     +----- Prisodontopsis_aviculaeformis
                    +--+  +--+                                   
                       |  |  |  +-- Coelatura_briarti **            
                       |  |  +--+                                
                       |  |     +-- Prisodontopsis_mweruensis    
                       +--+                                      
                          |     +-- Coelatura_choziensis **         
                          |     |                                
                          |     +-- Coelatura_hypsiprymna **        
                          |     |                                
                          +-----+-- Coelatura_luapulaensis **       
                                |                                
                                +-- Coelatura_mossambicensis **     
                                |                                
                                +-- Nyassunio

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
18211118189622
                       +----- Grandidieria                 
  +--------------------+                                   
  |                    |  +-- Pseudospatha                 
  |                    +--+                                
  |                       +-- Moncetia                     
  |                                                        
--+  +----------------------- Nitia                        
  |  |                                                     
  |  |                    +-- Coelatura_aegyptiaca **         
  |  |                    |                                
  |  |                    +-- Coelatura_alluaudi **           
  +--+                    |                                
     |  +-----------------+-- Coelatura_bakeri **             
     |  |                 |                                
     |  |                 +-- Coelatura_hauttecoeuri **       
     |  |                 |                                
     |  |                 +-- Coelatura_stuhlmanni **         
     +--+                                                  
        |                 +-- Coelatura_leopoldvillensis **   
        |  +--------------+                                
        |  |              +-- Coelatura_mesafricana **        
        |  |                                               
        |  |              +-- Coelatura_horei **              
        +--+              |                                
           |  +-----------+-- Coelatura_ratidota **           
           |  |           |                                
           |  |           +-- Coelatura **                    
           |  |                                            
           +--+           +-- Coelatura_gabonensis **         
              |  +--------+                                
              |  |        +-- Coelatura_stanleyvillensis **   
              |  |                                         
              +--+  +-------- Coelatura_kunenensis **         
                 |  |                                      
                 |  |     +-- Coelatura_briarti **            
                 |  |     |                                
                 +--+  +--+-- Prisodontopsis_mweruensis    
                    |  |  |                                
                    |  |  +-- Prisodontopsis_aviculaeformis
                    |  |                                   
                    +--+  +-- Coelatura_choziensis **         
                       |  |                                
                       |  +-- Coelatura_hypsiprymna **        
                       |  |                                
                       +--+-- Coelatura_luapulaensis **       
                          |                                
                          +-- Coelatura_mossambicensis **     
                          |                                
                          +-- Nyassunio

Graf et al., 2014 [Publ 3591], fig. 3

iTtTbiriSoTroTroSiD
57101163213
  +-------------------- Nitia                    
  |                                              
--+  +----------------- Coelatura_aegyptiaca__2 **  
  |  |                                           
  +--+  +-------------- Coelatura_aegyptiaca__1 **  
     |  |                                        
     +--+  +----------- Coelatura_gabonensis **     
        |  |                                     
        +--+  +-------- Coelatura_kunenensis **     
           |  |                                  
           |  |  +----- Prisodontopsis           
           +--+  |                               
              |  |  +-- Coelatura_choziensis__1 **  
              |  |  |                            
              +--+  +-- Coelatura_choziensis__2 **  
                 |  |                            
                 |  +-- Coelatura_luapulaensis__1 **
                 |  |                            
                 +--+-- Coelatura_luapulaensis__2 **
                    |                            
                    +-- Coelatura_luapulaensis__3 **
                    |                            
                    +-- Coelatura_luapulaensis__4 **
                    |                            
                    +-- Coelatura_luapulaensis__5 **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
58061203222
  +-- Nitia                        
  |                                
  +-- Coelatura_gabonensis **         
  |                                
  +-- Prisodontopsis_mweruensis    
  |                                
  +-- Coelatura_kunenensis **         
  |                                
--+-- Coelatura_aegyptiaca__1 **      
  |                                
  +-- Coelatura_aegyptiaca__2 **      
  |                                
  +-- Prisodontopsis_aviculaeformis
  |                                
  +-- Coelatura_luapulaensis **       
  |                                
  +-- Coelatura_choziensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
35441962219
  +----------- Coelatura_aegyptiaca__2 **
  |                                   
--+        +-- Coelatura_aegyptiaca__1 **
  |  +-----+                          
  |  |     +-- Nitia                  
  +--+                                
     |  +----- Coelatura_kunenensis **   
     +--+                             
        |  +-- Coelatura_gabonensis **   
        +--+                          
           +-- Prisodontopsis

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
34341362219
  +-------------- Nitia                  
--+                                      
  |  +----------- Coelatura_aegyptiaca__2 **
  +--+                                   
     |  +-------- Coelatura_aegyptiaca__1 **
     +--+                                
        |  +----- Prisodontopsis         
        +--+                             
           |  +-- Coelatura_gabonensis **   
           +--+                          
              +-- Coelatura_kunenensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
34341362219
  +-------------- Nitia                  
--+                                      
  |  +----------- Coelatura_aegyptiaca__1 **
  +--+                                   
     |  +-------- Coelatura_aegyptiaca__2 **
     +--+                                
        |  +----- Coelatura_gabonensis **   
        +--+                             
           |  +-- Coelatura_kunenensis **   
           +--+                          
              +-- Prisodontopsis

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
35241982220
  +-------- Coelatura_aegyptiaca__2 **
  |                                
--+  +----- Coelatura_aegyptiaca__1 **
  |  |                             
  |  +----- Nitia                  
  +--+                             
     |  +-- Coelatura_gabonensis **   
     |  |                          
     +--+-- Coelatura_kunenensis **   
        |                          
        +-- Prisodontopsis

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
35241982221
  +-------- Coelatura_aegyptiaca__2 **
  |                                
  +-------- Nitia                  
--+                                
  |  +----- Coelatura_aegyptiaca__1 **
  |  |                             
  +--+  +-- Coelatura_gabonensis **   
     |  |                          
     +--+-- Coelatura_kunenensis **   
        |                          
        +-- Prisodontopsis

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
2302612220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Nyassunio           
     +--+                       
        +-- Coelatura_kunenensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
2302562220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Coelatura_kunenensis **
     +--+                       
        +-- Nyassunio

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
2402592221
  +-- Coelatura_aegyptiaca **
  |                       
  +-- Coelatura_kunenensis **
--+                       
  +-- Nitia               
  |                       
  +-- Nyassunio

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
2302562220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Coelatura_kunenensis **
     +--+                       
        +-- Nyassunio

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
2302562220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Coelatura_kunenensis **
     +--+                       
        +-- Nyassunio

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
2303342220
  +----------- Nitia                  
--+                                   
  |  +-------- Coelatura_aegyptiaca__2 **
  +--+                                
     |  +----- Coelatura_aegyptiaca__1 **
     +--+                             
        |  +-- Coelatura_gabonensis **   
        +--+                          
           +-- Prisodontopsis

Bolotov et al., 2024 [Publ 4369], fig. 2

iTtTbiriSoTroTroSiD
2302602220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Coelatura_kunenensis **
     +--+                       
        +-- Nyassunio

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
2303342220
  +-------- Nitia                  
  |                                
--+  +----- Coelatura_aegyptiaca__1 **
  |  |                             
  +--+----- Coelatura_aegyptiaca__2 **
     |                             
     |  +-- Coelatura_gabonensis **   
     +--+                          
        +-- Prisodontopsis

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
2302592220
  +-------- Nitia               
--+                             
  |  +----- Coelatura_aegyptiaca **
  +--+                          
     |  +-- Nyassunio           
     +--+                       
        +-- Coelatura_kunenensis **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112100--
-- Coelatura_aegyptiaca **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Coelatura_aegyptiaca **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
11012722--
  +----- Nitia               
--+                          
  |  +-- Coelatura_aegyptiaca **
  +--+                       
     +-- Prisodontopsis

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Coelatura_aegyptiaca **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019000--
-- Coelatura_aegyptiaca **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
110111111--
  +-- Coelatura_aegyptiaca **
--+                       
  +-- Nitia

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Coelatura_aegyptiaca **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
11012200--
-- Coelatura_aegyptiaca **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111711--
  +-- Coelatura_aegyptiaca **
--+                       
  +-- Nitia

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014500--
-- Coelatura_aegyptiaca **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018211--
  +-- Coelatura_aegyptiaca **
--+                       
  +-- Nitia

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014511--
  +-- Coelatura_aegyptiaca **
--+                       
  +-- Nitia

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Coelatura_aegyptiaca **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015611--
  +-- Coelatura_choziensis **
--+                       
  +-- Prisodontopsis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015611--
  +-- Coelatura_choziensis **
--+                       
  +-- Prisodontopsis

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014500--
-- Coelatura_aegyptiaca **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014400--
-- Coelatura_aegyptiaca **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014600--
-- Coelatura_aegyptiaca **

Bolotov et al., 2019 [Publ 3974], fig. 1

iTtTbiriSoTroTroSiD
11011711--
  +-- Nitia               
--+                       
  +-- Coelatura_aegyptiaca **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013000--
-- Coelatura_aegyptiaca **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014400--
-- Coelatura_aegyptiaca **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Coelatura_aegyptiaca **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Coelatura_aegyptiaca **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014600--
-- Coelatura_aegyptiaca **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019000--
-- Coelatura_aegyptiaca **
 
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