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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Unionini | Unionina | Unio

species Unio caffer Krauss, 1848: synonymy
     186 specimens + 1 Genbank voucher

Unio caffer Krauss, 1848
Krauss, 1848. Südafrik. Moll.: 18, pl. 1, fig. 14.
Type(s): TYPE SMF_XXX, Rivers of Natal.

Nodularia (sec. Cafferia) cafferSimpson (1900)
Nodularia (Cafferia) caffraKobelt (1910)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio cafferOrtmann (1918), Heard & Vail (1976), Appleton (1979), Graf (2000), Graf & Ó Foighil (2000), ... Graf & Cummings (2021), Bieler (2021), Lopes-Lima et al. (2021), Lyubas et al. (2022), Campbell & Lydeard (2026)
Cafferia caffraConnolly (1925), Connolly (1931), Haas (1936), Modell (1942), Haas (1969), ... Hoeh et al. (2009), Bogan (2010), Graf & Cummings (2011), Reis et al. (2013), Marshall et al. (2014)
Unio (Cafferia) cafferPilsbry & Bequaert (1927), Thiele (1934), Connolly (1939), Graf (2002)
Cafferia cafferGraf & Cummings (2006), Jadhav & Jamkhedkar (2009)

Unio zeyheri Menke, 1849
Menke, 1849. Zeitschr. f. Malakozool. 5: 28.

Nodularia (sec. Cafferia) zeyheriSimpson (1900)
Nodularia (Cafferia) zeyheriKobelt (1910)
Unio (sec. Cafferia) cafferSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio cyamus Philippi, 1853
Philippi, 1853. Zeitschr. f. Malakozool. 8: 125.

Simpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio africanus Lea, 1856
Lea, 1856. Proc. Acad. Nat. Sci. 8: 94.
Lea, 1857. Jour. Acad. Nat. Sci. 3 [O. 6]: 300 [20], pl. 27, fig. 15.
Type(s): LECTOTYPE USNM_85233, C. Good Hope.

Nodularia (sec. Cafferia) caffer var. africanaSimpson (1900)
Nodularia (Cafferia) caffra var. africanaKobelt (1910)
Unio (sec. Cafferia) caffer var. africanusSimpson (1914)
Unio africanusPilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio verrauxi Küster, 1856
‘Charpentier mss.’ Küster, 1856. Conchyl. Cab. 9(2): 150, pl. 43, fig. 6.

Cafferia caffra verreauxi [sic]Connolly (1925)
Lampsilis (Cyrtonaias) umbrosaFrierson (1927)
Pilsbry & Bequaert (1927)
Unio (Cafferia) caffer verreauxi [sic]Connolly (1939)
Cafferia caffraHaas (1969), Daget (1998)
Unio cafferHeard & Vail (1976)

Unio verreauianus Lea, 1856
Lea, 1856. Proc. Acad. Nat. Sci. 8: 94.
Lea, 1857. Jour. Acad. Nat. Sci. 3 [O. 6]: 301 [21], pl. 27, fig. 16.
(renamed Unio verreauxianus Lea, 1867)
Type(s): LECTOTYPE USNM_84418 (by Boyko & Sage, 1996: 28), C. Good Hope.

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)
Cafferia caffra verreauxi [sic]Connolly (1925)
Unio verreauianusPilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)
Unio cafferBieler (2021)

Unio diminutis Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 151.
(incorrect original spelling of Unio diminutus Lea, 1859)
Type(s): LECTOTYPE BMNH_1965165, East Africa.

Nodularia (sec. Cafferia) diminutaSimpson (1900)
Unio (sec. Cafferia) diminutusSimpson (1914)
Unio diminutusPilsbry & Bequaert (1927)
Unio (Cafferia) caffer verreauxi [sic]Connolly (1939)
Cafferia caffraDaget (1998)
Unio cafferBieler (2021)

Unio diminutus Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 151 [as ‘diminutis’].
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 254, pl. 39, fig. 134.
(correct name for Unio diminutis Lea, 1859)
Type(s): BMNH_1965165, East Africa.

Nodularia (sec. Cafferia) diminutaSimpson (1900)
Nodularia (Cafferia) diminutaKobelt (1910)
Unio (sec. Cafferia) diminutusSimpson (1914)
Unio diminutusPilsbry & Bequaert (1927)
Haas (1936)
Unio (Cafferia) caffer verreauxi [sic]Connolly (1939)
Cafferia caffraDaget (1998)
Unio cafferBieler (2021)

Unio gratiosus Küster, 1861
‘Parreiss in sched.’ Küster, 1861. Conchyl. Cab. 9(2): 239, pl. 80, fig. 3.
(homonym of Unio gratiosus Philippi, 1843)

Unio diminutusPilsbry & Bequaert (1927)

Unio natalensis Lea, 1864
Lea, 1864. Proc. Acad. Nat. Sci. 16: 113.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 59 [63], pl. 20, fig. 57.
Type(s): HOLOTYPE (MONOTYPY) ANSP_41686, Umpingave River, Natal, S. Afr.

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio natalensisPilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Graf & Cummings (2011)

Unio zelebori Dunker, 1866
Dunker, 1866. Verh. Zool.-Bot. Ges. Wien 16: 915.
(homonym of Unio zelebori Zelebor, 1851)

Diplodon (Hyridella s.s.) zeleboriSimpson (1900), Simpson (1914)
Hyridella (Echyridella) menziesiMcMichael & Hiscock (1958), Haas (1969)
Cafferia caffraMarshall & Fenwick (2006), Marshall et al. (2014)

Unio (Hyridella) fissidens O. Boettger, 1866
O. Boettger, 1866. Ber. Senckenb. Naturf. Ges. 1885/86: 27, pl. 2, figs. 6-7.
Type(s): TYPE SMF_3442, Hygap östl. v. Keheum, S. Kalahari.

Nodularia (sec. Cafferia) fissidensSimpson (1900)
Nodularia (Cafferia) fissidensKobelt (1910)
Unio (sec. Cafferia) fissidensSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio (Hyridella) hygapanus O. Boettger, 1866
O. Boettger, 1866. Ber. Senckenb. Naturf. Ges. 1885/86: 26, pl. 2, fig. 5.
Type(s): HOLOTYPE SMF_3441, Hygap östl. v. Keheum, S. Kalahari.

Nodularia (sec. Cafferia) hygapanusSimpson (1900)
Nodularia (Cafferia) hygapanusKobelt (1910)
Unio (sec. Cafferia) hygapanusSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Marshall & Fenwick (2006)

Unio verreauxianus Lea, 1867
Lea, 1867. Obs., index I-XI: errata.
Lea, 1870. Synopsis Fam. Naiades: 36.
(new name for Unio verreauianus Lea, 1856)

Nodularia (sec. Cafferia) cafferSimpson (1900)
Nodularia (Cafferia) caffra var. verreauxianaKobelt (1910)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio verreauianusPilsbry & Bequaert (1927)
Cafferia caffraDaget (1998)
Unio cafferBieler (2021)

Unio rectilinearis Sowerby, 1868
Sowerby, 1868. Conch. Icon. 16: pl. 65, sp. 332.
Type(s): HOLOTYPE BMNH_74-12-11-4, Riv. Columbia.

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio vaalensis Chaper, 1885
Chaper, 1885. Bull. Soc. Zool. France 10: 480, pl. 11, figs. 1-3.
Type(s): SYNTYPES MNHN_MP_0180, la Vaal, Afrique Australe.

Nodularia (sec. Cafferia) caffer var. vaalensisSimpson (1900)
Nodularia (Cafferia) caffra var. vaalensisKobelt (1910)
Unio (sec. Cafferia) caffer var. vaalensisSimpson (1914)
Unio natalensisPilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Marshall & Fenwick (2006), Graf & Cummings (2011)

Unio gratiosus Paetel, 1890
Paetel, 1890. Cat. Conch.-Samml. 3: 154.
(reference to Unio gratiosus Küster, 1861)

Nodularia (sec. Cafferia) diminutaSimpson (1900)
Unio (sec. Cafferia) diminutusSimpson (1914)

Unio verreauxi E.A. Smith, 1891
‘Kuster’ of Authors.
E.A. Smith, 1891. Ann. Mag. Nat. Hist. 8: 319.
Sturany, 1899. Denk. Ak. Wien 67: 627.
Connolly, 1925. Rec. Albany Mus. 3: [1].
Connolly, 1939. Ann. S. Afr. Mus. 33: 608.
Heard & Vail, 1976. Zool. Africana 11(1): 45.
(error for Unio verrauxi Küster, 1856)

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio (Cafferia) caffer verreauxi [sic]Connolly (1939)
Cafferia caffraDaget (1998)

Unio caffer var. pentheri Sturany, 1898
Sturany, 1898. Anz. Kais. Akad. Wiss. , Math.-Nat. Classe 35(16): 161.
Sturany, 1899. Denk. Ak. Wien 67: 627, pl. 3, figs. 64-65.
(intro'd as intra-sp. of Unio caffer Krauss, 1848)
Type(s): SYNTYPES NMW_XXX, Panda ma tinka (Zambezi drainage).

Unio (sec. Cafferia) caffer var. pentheriSimpson (1914)
Unio (Cafferia) caffer var. pentheriPilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Marshall & Fenwick (2006)
Cafferia cafferGraf & Cummings (2006)

Unio charon Preston, 1910
Preston, 1910. Ann. Mag. Nat. Hist. 6: 60, pl. 4, fig. 6.
Type(s): FIGURED SYNTYPE MRAC_56592, Brit. C. Afr.: Silongwe.

Unio charonSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Graf & Cummings (2011)

Unio mashonae Preston, 1910
Preston, 1910. Ann. Mag. Nat. Hist. 6: 61, pl. 4, fig. 10 [as ‘mashonæ’].
Type(s): HOLOTYPE (ORIG. DESIG.) BMNH_1910-1-15-6, Banks of a strait, or on the veldt about 16 miles from Enkeldoorn, Mashonaland.

Unio (sec. Cafferia) mashonaeSimpson (1914)
Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Graf & Cummings (2011)
Cafferia cafferGraf & Cummings (2006)

Unio vicinus Preston, 1910
Preston, 1910. Ann. Mag. Nat. Hist. 6: 60, pl. 4, fig. 7.
(renamed Unio silongweensis Preston, 1912; homonym of Unio vicinus Lea, 1857)
Type(s): FIGURED SYNTYPE MRAC_56589, Brit. C. Afr.: Silongwe.

Unio silongweensisSimpson (1914), Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998), Marshall & Fenwick (2006)

Unio silongweensis Preston, 1912
Preston, 1912. Nautilus 26: 35.
(new name for Unio vicinus Preston, 1910)

Unio silongweensisSimpson (1914), Pilsbry & Bequaert (1927)
Unio (Cafferia) cafferConnolly (1939)
Cafferia caffraHaas (1969), Mandahl-Barth (1988), Daget (1998)

Unio (Cafferia) connollyi Pilsbry, 1923
Pilsbry, 1923. Proc. Acad. Nat. Sci. 75: 276, text fig. 1; pl. 19, figs. 3-4.
Pilsbry & Bequaert, 1927. Bull. AMNH 53: 380.
Type(s): HOLOTYPE (ORIG. DESIG.) ANSP_125501A, Port Natal, Africa.

Unio (Cafferia) connollyiPilsbry & Bequaert (1927)
Unio (Cafferia) caffer connollyiConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)
Unio cafferHeard & Vail (1976)

Cafferia caffra obesa Connolly, 1925
Connolly, 1925. Rec. Albany Mus. 3: 261, pl. 1, figs. 1-4.
(intro'd as intra-sp. of Unio caffer Krauss, 1848)
Type(s): POSSIBLE SYNTYPES MCZ_52457, Bushmans River, Alicedale, South Africa.

Cafferia caffra obesaConnolly (1925)
Unio (Cafferia) connollyiPilsbry & Bequaert (1927)
Unio (Cafferia) caffer connollyiConnolly (1939)
Cafferia caffraHaas (1969), Daget (1998)

Unio caffer Küster, 1856
‘Krauss’ Küster, 1856. Conchyl. Cab. 9(2): 143, pl. 42, fig. 3. [in part]
(reference to Unio caffer Küster, 1856)

Nodularia (sec. Cafferia) caffer var. africanaSimpson (1900)
Unio (sec. Cafferia) caffer var. africanusSimpson (1914)

Unio caffer Küster, 1856
‘Krauss’ Küster, 1856. Conchyl. Cab. 9(2): 143, pl. 42, fig. 2. [in part]
(reference to Unio caffer Krauss, 1848)

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)

Unio niloticus Küster, 1856
‘Caillaud’ Küster, 1856. Conchyl. Cab. 9(2): 158, pl. 45, fig. 5.
(misidentified reference to Unio niloticus Cailliaud, 1827)

Nodularia (sec. Cafferia) caffer var. africanaSimpson (1900)
Nodularia (Cælatura [sic]) niloticaPallary (1909)
Unio (sec. Cafferia) caffer var. africanusSimpson (1914)

Unio verrauianus Sowerby, 1868
‘Lea’ Sowerby, 1868. Conch. Icon. 16: pl. 69, sp. 352.
(error for Unio verreauianus Lea, 1856)

Nodularia (sec. Cafferia) cafferSimpson (1900)
Unio (sec. Cafferia) cafferSimpson (1914)

Unio teyheri Lea, 1870
‘Menke’ Lea, 1870. Synopsis Fam. Naiades: 64.
(error for Unio zeyheri Menke, 1849)

Unio gracilis Jickeli, 1874
‘Parreyss’ Jickeli, 1874. Nova Acta Dtsch. Akad. Naturf. 37(1): 280.
(intro'd as syn of Unio diminutus Lea, 1859)

Unio diminutusPilsbry & Bequaert (1927)

Taxonomic History (since 1900)

Nodularia (sec. Cafferia) cafferSimpson (1900)
Nodularia (sec. Cafferia) zeyheriSimpson (1900)
Simpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927), Haas (1936)
Nodularia (sec. Cafferia) caffer var. africanaSimpson (1900)
Nodularia (sec. Cafferia) diminutaSimpson (1900)
Diplodon (Hyridella s.s.) zeleboriSimpson (1900), Simpson (1914)
Nodularia (sec. Cafferia) fissidensSimpson (1900)
Nodularia (sec. Cafferia) hygapanusSimpson (1900)
Nodularia (sec. Cafferia) caffer var. vaalensisSimpson (1900)
Nodularia (Cælatura [sic]) niloticaPallary (1909)
Nodularia (Cafferia) caffraKobelt (1910)
Nodularia (Cafferia) zeyheriKobelt (1910)
Nodularia (Cafferia) caffra var. africanaKobelt (1910)
Nodularia (Cafferia) diminutaKobelt (1910)
Nodularia (Cafferia) fissidensKobelt (1910)
Nodularia (Cafferia) hygapanusKobelt (1910)
Nodularia (Cafferia) caffra var. verreauxianaKobelt (1910)
Nodularia (Cafferia) caffra var. vaalensisKobelt (1910)
Unio (sec. Cafferia) cafferSimpson (1914)
Unio (sec. Cafferia) caffer var. africanusSimpson (1914)
Unio (sec. Cafferia) diminutusSimpson (1914)
Unio (sec. Cafferia) fissidensSimpson (1914)
Unio (sec. Cafferia) hygapanusSimpson (1914)
Unio (sec. Cafferia) caffer var. vaalensisSimpson (1914)
Unio (sec. Cafferia) caffer var. pentheriSimpson (1914)
Unio charonSimpson (1914)
Unio (sec. Cafferia) mashonaeSimpson (1914)
Unio silongweensisSimpson (1914), Pilsbry & Bequaert (1927)
Unio cafferOrtmann (1918), Heard & Vail (1976), Appleton (1979), Graf (2000), Graf & Ó Foighil (2000), ... Graf & Cummings (2021), Bieler (2021), Lopes-Lima et al. (2021), Lyubas et al. (2022), Campbell & Lydeard (2026)
Cafferia caffraConnolly (1925), Connolly (1931), Haas (1936), Modell (1942), Haas (1969), ... Hoeh et al. (2009), Bogan (2010), Graf & Cummings (2011), Reis et al. (2013), Marshall et al. (2014)
Cafferia caffra verreauxi [sic]Connolly (1925)
Cafferia caffra obesaConnolly (1925)
Unio (Cafferia) cafferPilsbry & Bequaert (1927), Thiele (1934), Connolly (1939), Graf (2002)
Unio africanusPilsbry & Bequaert (1927)
Lampsilis (Cyrtonaias) umbrosaFrierson (1927)
Unio verreauianusPilsbry & Bequaert (1927)
Unio diminutusPilsbry & Bequaert (1927)
Unio natalensisPilsbry & Bequaert (1927)
Unio (Cafferia) caffer var. pentheriPilsbry & Bequaert (1927)
Unio (Cafferia) connollyiPilsbry & Bequaert (1927)
Unio (Cafferia) caffer verreauxi [sic]Connolly (1939)
Unio (Cafferia) caffer connollyiConnolly (1939)
Hyridella (Echyridella) menziesiMcMichael & Hiscock (1958), Haas (1969)
Cafferia cafferGraf & Cummings (2006), Jadhav & Jamkhedkar (2009)

 
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