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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Unionini | Unionina | Unio species Unio caffer Krauss, 1848: synonymy 186 specimens + 1 Genbank voucher Unio caffer Krauss, 1848
Krauss, 1848. Südafrik. Moll.: 18, pl. 1, fig. 14. Type(s): TYPE SMF_XXX, Rivers of Natal.
Nodularia (sec. Cafferia) caffer — Simpson (1900) Nodularia (Cafferia) caffra — Kobelt (1910) Unio (sec. Cafferia) caffer — Simpson (1914) Unio caffer — Ortmann (1918), Heard & Vail (1976), Appleton (1979), Graf (2000), Graf & Ó Foighil (2000), ... Graf & Cummings (2021), Bieler (2021), Lopes-Lima et al. (2021), Lyubas et al. (2022), Campbell & Lydeard (2026) Cafferia caffra — Connolly (1925), Connolly (1931), Haas (1936), Modell (1942), Haas (1969), ... Hoeh et al. (2009), Bogan (2010), Graf & Cummings (2011), Reis et al. (2013), Marshall et al. (2014) Unio (Cafferia) caffer — Pilsbry & Bequaert (1927), Thiele (1934), Connolly (1939), Graf (2002) Cafferia caffer — Graf & Cummings (2006), Jadhav & Jamkhedkar (2009) | |
Unio zeyheri Menke, 1849 Menke, 1849. Zeitschr. f. Malakozool. 5: 28.
Nodularia (sec. Cafferia) zeyheri — Simpson (1900) Nodularia (Cafferia) zeyheri — Kobelt (1910) Unio (sec. Cafferia) caffer — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio cyamus Philippi, 1853 Philippi, 1853. Zeitschr. f. Malakozool. 8: 125.
— Simpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927) Unio (sec. Cafferia) caffer — Simpson (1914) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio africanus Lea, 1856
Lea, 1856. Proc. Acad. Nat. Sci. 8: 94.
Lea, 1857. Jour. Acad. Nat. Sci. 3 [O. 6]: 300 [20], pl. 27, fig. 15. Type(s): LECTOTYPE USNM_85233, C. Good Hope.
Nodularia (sec. Cafferia) caffer var. africana — Simpson (1900) Nodularia (Cafferia) caffra var. africana — Kobelt (1910) Unio (sec. Cafferia) caffer var. africanus — Simpson (1914) Unio africanus — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio verrauxi Küster, 1856
‘Charpentier mss.’ Küster, 1856. Conchyl. Cab. 9(2): 150, pl. 43, fig. 6.
Cafferia caffra verreauxi [sic] — Connolly (1925) Lampsilis (Cyrtonaias) umbrosa — Frierson (1927) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer verreauxi [sic] — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) Unio caffer — Heard & Vail (1976) | |
Unio verreauianus Lea, 1856
Lea, 1856. Proc. Acad. Nat. Sci. 8: 94.
Lea, 1857. Jour. Acad. Nat. Sci. 3 [O. 6]: 301 [21], pl. 27, fig. 16. (renamed Unio verreauxianus Lea, 1867) Type(s): LECTOTYPE USNM_84418 (by Boyko & Sage, 1996: 28), C. Good Hope.
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) Cafferia caffra verreauxi [sic] — Connolly (1925) Unio verreauianus — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) Unio caffer — Bieler (2021) | |
Unio diminutis Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 151. (incorrect original spelling of Unio diminutus Lea, 1859) Type(s): LECTOTYPE BMNH_1965165, East Africa.
Nodularia (sec. Cafferia) diminuta — Simpson (1900) Unio (sec. Cafferia) diminutus — Simpson (1914) Unio diminutus — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer verreauxi [sic] — Connolly (1939) Cafferia caffra — Daget (1998) Unio caffer — Bieler (2021) | |
Unio diminutus Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 151 [as ‘diminutis’].
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 254, pl. 39, fig. 134. (correct name for Unio diminutis Lea, 1859) Type(s): BMNH_1965165, East Africa.
Nodularia (sec. Cafferia) diminuta — Simpson (1900) Nodularia (Cafferia) diminuta — Kobelt (1910) Unio (sec. Cafferia) diminutus — Simpson (1914) Unio diminutus — Pilsbry & Bequaert (1927) — Haas (1936) Unio (Cafferia) caffer verreauxi [sic] — Connolly (1939) Cafferia caffra — Daget (1998) Unio caffer — Bieler (2021) | |
Unio gratiosus Küster, 1861
‘Parreiss in sched.’ Küster, 1861. Conchyl. Cab. 9(2): 239, pl. 80, fig. 3. (homonym of Unio gratiosus Philippi, 1843)
Unio diminutus — Pilsbry & Bequaert (1927) | |
Unio natalensis Lea, 1864
Lea, 1864. Proc. Acad. Nat. Sci. 16: 113.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 59 [63], pl. 20, fig. 57. Type(s): HOLOTYPE (MONOTYPY) ANSP_41686, Umpingave River, Natal, S. Afr.
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) Unio natalensis — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Graf & Cummings (2011) | |
Unio zelebori Dunker, 1866 Dunker, 1866. Verh. Zool.-Bot. Ges. Wien 16: 915. (homonym of Unio zelebori Zelebor, 1851)
Diplodon (Hyridella s.s.) zelebori — Simpson (1900), Simpson (1914) Hyridella (Echyridella) menziesi — McMichael & Hiscock (1958), Haas (1969) Cafferia caffra — Marshall & Fenwick (2006), Marshall et al. (2014) | |
Unio (Hyridella) fissidens O. Boettger, 1866
O. Boettger, 1866. Ber. Senckenb. Naturf. Ges. 1885/86: 27, pl. 2, figs. 6-7. Type(s): TYPE SMF_3442, Hygap östl. v. Keheum, S. Kalahari.
Nodularia (sec. Cafferia) fissidens — Simpson (1900) Nodularia (Cafferia) fissidens — Kobelt (1910) Unio (sec. Cafferia) fissidens — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio (Hyridella) hygapanus O. Boettger, 1866
O. Boettger, 1866. Ber. Senckenb. Naturf. Ges. 1885/86: 26, pl. 2, fig. 5. Type(s): HOLOTYPE SMF_3441, Hygap östl. v. Keheum, S. Kalahari.
Nodularia (sec. Cafferia) hygapanus — Simpson (1900) Nodularia (Cafferia) hygapanus — Kobelt (1910) Unio (sec. Cafferia) hygapanus — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Marshall & Fenwick (2006) | |
Unio verreauxianus Lea, 1867 Lea, 1867. Obs., index I-XI: errata. Lea, 1870. Synopsis Fam. Naiades: 36. (new name for Unio verreauianus Lea, 1856)
Nodularia (sec. Cafferia) caffer — Simpson (1900) Nodularia (Cafferia) caffra var. verreauxiana — Kobelt (1910) Unio (sec. Cafferia) caffer — Simpson (1914) Unio verreauianus — Pilsbry & Bequaert (1927) Cafferia caffra — Daget (1998) Unio caffer — Bieler (2021) | |
Unio rectilinearis Sowerby, 1868
 Sowerby, 1868. Conch. Icon. 16: pl. 65, sp. 332. Type(s): HOLOTYPE BMNH_74-12-11-4, Riv. Columbia.
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio vaalensis Chaper, 1885
 Chaper, 1885. Bull. Soc. Zool. France 10: 480, pl. 11, figs. 1-3. Type(s): SYNTYPES MNHN_MP_0180, la Vaal, Afrique Australe.
Nodularia (sec. Cafferia) caffer var. vaalensis — Simpson (1900) Nodularia (Cafferia) caffra var. vaalensis — Kobelt (1910) Unio (sec. Cafferia) caffer var. vaalensis — Simpson (1914) Unio natalensis — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Marshall & Fenwick (2006), Graf & Cummings (2011) | |
Unio gratiosus Paetel, 1890 Paetel, 1890. Cat. Conch.-Samml. 3: 154. (reference to Unio gratiosus Küster, 1861)
Nodularia (sec. Cafferia) diminuta — Simpson (1900) Unio (sec. Cafferia) diminutus — Simpson (1914) | |
Unio verreauxi E.A. Smith, 1891 ‘Kuster’ of Authors. E.A. Smith, 1891. Ann. Mag. Nat. Hist. 8: 319. Sturany, 1899. Denk. Ak. Wien 67: 627. Connolly, 1925. Rec. Albany Mus. 3: [1]. Connolly, 1939. Ann. S. Afr. Mus. 33: 608. Heard & Vail, 1976. Zool. Africana 11(1): 45. (error for Unio verrauxi Küster, 1856)
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) Unio (Cafferia) caffer verreauxi [sic] — Connolly (1939) Cafferia caffra — Daget (1998) | |
Unio caffer var. pentheri Sturany, 1898 Sturany, 1898. Anz. Kais. Akad. Wiss. , Math.-Nat. Classe 35(16): 161.
Sturany, 1899. Denk. Ak. Wien 67: 627, pl. 3, figs. 64-65. (intro'd as intra-sp. of Unio caffer Krauss, 1848) Type(s): SYNTYPES NMW_XXX, Panda ma tinka (Zambezi drainage).
Unio (sec. Cafferia) caffer var. pentheri — Simpson (1914) Unio (Cafferia) caffer var. pentheri — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Marshall & Fenwick (2006) Cafferia caffer — Graf & Cummings (2006) | |
Unio charon Preston, 1910
 Preston, 1910. Ann. Mag. Nat. Hist. 6: 60, pl. 4, fig. 6. Type(s): FIGURED SYNTYPE MRAC_56592, Brit. C. Afr.: Silongwe.
Unio charon — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Graf & Cummings (2011) | |
Unio mashonae Preston, 1910
 Preston, 1910. Ann. Mag. Nat. Hist. 6: 61, pl. 4, fig. 10 [as ‘mashonæ’]. Type(s): HOLOTYPE (ORIG. DESIG.) BMNH_1910-1-15-6, Banks of a strait, or on the veldt about 16 miles from Enkeldoorn, Mashonaland.
Unio (sec. Cafferia) mashonae — Simpson (1914) — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Graf & Cummings (2011) Cafferia caffer — Graf & Cummings (2006) | |
Unio vicinus Preston, 1910
 Preston, 1910. Ann. Mag. Nat. Hist. 6: 60, pl. 4, fig. 7. (renamed Unio silongweensis Preston, 1912; homonym of Unio vicinus Lea, 1857) Type(s): FIGURED SYNTYPE MRAC_56589, Brit. C. Afr.: Silongwe.
Unio silongweensis — Simpson (1914), Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998), Marshall & Fenwick (2006) | |
Unio silongweensis Preston, 1912 Preston, 1912. Nautilus 26: 35. (new name for Unio vicinus Preston, 1910)
Unio silongweensis — Simpson (1914), Pilsbry & Bequaert (1927) Unio (Cafferia) caffer — Connolly (1939) Cafferia caffra — Haas (1969), Mandahl-Barth (1988), Daget (1998) | |
Unio (Cafferia) connollyi Pilsbry, 1923
Pilsbry, 1923. Proc. Acad. Nat. Sci. 75: 276, text fig. 1; pl. 19, figs. 3-4. Pilsbry & Bequaert, 1927. Bull. AMNH 53: 380. Type(s): HOLOTYPE (ORIG. DESIG.) ANSP_125501A, Port Natal, Africa.
Unio (Cafferia) connollyi — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer connollyi — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) Unio caffer — Heard & Vail (1976) | |
Cafferia caffra obesa Connolly, 1925
Connolly, 1925. Rec. Albany Mus. 3: 261, pl. 1, figs. 1-4. (intro'd as intra-sp. of Unio caffer Krauss, 1848) Type(s): POSSIBLE SYNTYPES MCZ_52457, Bushmans River, Alicedale, South Africa.
Cafferia caffra obesa — Connolly (1925) Unio (Cafferia) connollyi — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer connollyi — Connolly (1939) Cafferia caffra — Haas (1969), Daget (1998) | |
Unio caffer Küster, 1856
‘Krauss’ Küster, 1856. Conchyl. Cab. 9(2): 143, pl. 42, fig. 3. [in part] (reference to Unio caffer Küster, 1856)
Nodularia (sec. Cafferia) caffer var. africana — Simpson (1900) Unio (sec. Cafferia) caffer var. africanus — Simpson (1914) | |
Unio caffer Küster, 1856
‘Krauss’ Küster, 1856. Conchyl. Cab. 9(2): 143, pl. 42, fig. 2. [in part] (reference to Unio caffer Krauss, 1848)
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) | |
Unio niloticus Küster, 1856
‘Caillaud’ Küster, 1856. Conchyl. Cab. 9(2): 158, pl. 45, fig. 5. (misidentified reference to Unio niloticus Cailliaud, 1827)
Nodularia (sec. Cafferia) caffer var. africana — Simpson (1900) Nodularia (Cælatura [sic]) nilotica — Pallary (1909) Unio (sec. Cafferia) caffer var. africanus — Simpson (1914) | |
Unio verrauianus Sowerby, 1868
‘Lea’ Sowerby, 1868. Conch. Icon. 16: pl. 69, sp. 352. (error for Unio verreauianus Lea, 1856)
Nodularia (sec. Cafferia) caffer — Simpson (1900) Unio (sec. Cafferia) caffer — Simpson (1914) | |
Unio teyheri Lea, 1870 ‘Menke’ Lea, 1870. Synopsis Fam. Naiades: 64. (error for Unio zeyheri Menke, 1849) | |
Unio gracilis Jickeli, 1874 ‘Parreyss’ Jickeli, 1874. Nova Acta Dtsch. Akad. Naturf. 37(1): 280. (intro'd as syn of Unio diminutus Lea, 1859)
Unio diminutus — Pilsbry & Bequaert (1927) | |
Taxonomic History (since 1900)Nodularia (sec. Cafferia) caffer — Simpson (1900) Nodularia (sec. Cafferia) zeyheri — Simpson (1900) — Simpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927), Haas (1936) Nodularia (sec. Cafferia) caffer var. africana — Simpson (1900) Nodularia (sec. Cafferia) diminuta — Simpson (1900) Diplodon (Hyridella s.s.) zelebori — Simpson (1900), Simpson (1914) Nodularia (sec. Cafferia) fissidens — Simpson (1900) Nodularia (sec. Cafferia) hygapanus — Simpson (1900) Nodularia (sec. Cafferia) caffer var. vaalensis — Simpson (1900) Nodularia (Cælatura [sic]) nilotica — Pallary (1909) Nodularia (Cafferia) caffra — Kobelt (1910) Nodularia (Cafferia) zeyheri — Kobelt (1910) Nodularia (Cafferia) caffra var. africana — Kobelt (1910) Nodularia (Cafferia) diminuta — Kobelt (1910) Nodularia (Cafferia) fissidens — Kobelt (1910) Nodularia (Cafferia) hygapanus — Kobelt (1910) Nodularia (Cafferia) caffra var. verreauxiana — Kobelt (1910) Nodularia (Cafferia) caffra var. vaalensis — Kobelt (1910) Unio (sec. Cafferia) caffer — Simpson (1914) Unio (sec. Cafferia) caffer var. africanus — Simpson (1914) Unio (sec. Cafferia) diminutus — Simpson (1914) Unio (sec. Cafferia) fissidens — Simpson (1914) Unio (sec. Cafferia) hygapanus — Simpson (1914) Unio (sec. Cafferia) caffer var. vaalensis — Simpson (1914) Unio (sec. Cafferia) caffer var. pentheri — Simpson (1914) Unio charon — Simpson (1914) Unio (sec. Cafferia) mashonae — Simpson (1914) Unio silongweensis — Simpson (1914), Pilsbry & Bequaert (1927) Unio caffer — Ortmann (1918), Heard & Vail (1976), Appleton (1979), Graf (2000), Graf & Ó Foighil (2000), ... Graf & Cummings (2021), Bieler (2021), Lopes-Lima et al. (2021), Lyubas et al. (2022), Campbell & Lydeard (2026) Cafferia caffra — Connolly (1925), Connolly (1931), Haas (1936), Modell (1942), Haas (1969), ... Hoeh et al. (2009), Bogan (2010), Graf & Cummings (2011), Reis et al. (2013), Marshall et al. (2014) Cafferia caffra verreauxi [sic] — Connolly (1925) Cafferia caffra obesa — Connolly (1925) Unio (Cafferia) caffer — Pilsbry & Bequaert (1927), Thiele (1934), Connolly (1939), Graf (2002) Unio africanus — Pilsbry & Bequaert (1927) Lampsilis (Cyrtonaias) umbrosa — Frierson (1927) Unio verreauianus — Pilsbry & Bequaert (1927) Unio diminutus — Pilsbry & Bequaert (1927) Unio natalensis — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer var. pentheri — Pilsbry & Bequaert (1927) Unio (Cafferia) connollyi — Pilsbry & Bequaert (1927) Unio (Cafferia) caffer verreauxi [sic] — Connolly (1939) Unio (Cafferia) caffer connollyi — Connolly (1939) Hyridella (Echyridella) menziesi — McMichael & Hiscock (1958), Haas (1969) Cafferia caffer — Graf & Cummings (2006), Jadhav & Jamkhedkar (2009)
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