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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Leoparreysiini

genus Leoparreysia Vikhrev, Bolotov & Aksenova in Bolotov et al., 2017: cladomics
     9 Recent valid species (9 total species)

Cladomics Summary

number of species = 9 (9 total)
analyzed in 41 cladograms in 27 publications
maximum number of terminals in an analysis = 5 (mean = 2.88, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 5, (mean = 3.57 median = 3)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Bolotov et al., 2019 [Publ 3974], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
553513000
        +-- Leoparreysia_whitteni **  
  +-----+                          
  |     +-- Leoparreysia_burmana **   
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_tavoyensis **
     +--+                          
        +-- Leoparreysia_canefrii **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
553544000
        +-- Leoparreysia_whitteni **  
  +-----+                          
  |     +-- Leoparreysia_burmana **   
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_canefrii **  
     +--+                          
        +-- Leoparreysia_tavoyensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
5535627002
  +----------- Leoparreysia_burmana **   
--+                                   
  |  +-------- Leoparreysia_whitteni **  
  +--+                                
     |  +----- Leoparreysia_olivacea **  
     +--+                             
        |  +-- Leoparreysia_canefrii **  
        +--+                          
           +-- Leoparreysia_tavoyensis **

Bolotov et al., 2022 [Publ 4250], fig. S3

iTtTbiriSoTroTroSiD
55350002
  +----------- Leoparreysia_burmana **   
--+                                   
  |  +-------- Leoparreysia_whitteni **  
  +--+                                
     |  +----- Leoparreysia_olivacea **  
     +--+                             
        |  +-- Leoparreysia_tavoyensis **
        +--+                          
           +-- Leoparreysia_canefrii **

Bolotov et al., 2024 [Publ 4369], fig. 2

iTtTbiriSoTroTroSiD
553557000
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_canefrii **  
     +--+                          
        +-- Leoparreysia_tavoyensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5535112002
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_tavoyensis **
  +--+                             
     |  +-- Leoparreysia_olivacea **  
     +--+                          
        +-- Leoparreysia_canefrii **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
553556000
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_canefrii **  
     +--+                          
        +-- Leoparreysia_tavoyensis **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
553558000
        +-- Leoparreysia_whitteni **  
  +-----+                          
  |     +-- Leoparreysia_burmana **   
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_tavoyensis **
     +--+                          
        +-- Leoparreysia_canefrii **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
553553002
  +----------- Leoparreysia_burmana **   
--+                                   
  |  +-------- Leoparreysia_whitteni **  
  +--+                                
     |  +----- Leoparreysia_olivacea **  
     +--+                             
        |  +-- Leoparreysia_canefrii **  
        +--+                          
           +-- Leoparreysia_tavoyensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
553553000
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_canefrii **  
     +--+                          
        +-- Leoparreysia_tavoyensis **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
550556003
  +-- Leoparreysia_burmana **   
  |                          
  +-- Leoparreysia_canefrii **  
  |                          
--+-- Leoparreysia_olivacea **  
  |                          
  +-- Leoparreysia_tavoyensis **
  |                          
  +-- Leoparreysia_whitteni **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
553553000
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_canefrii **  
     +--+                          
        +-- Leoparreysia_tavoyensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
5535106000
        +-- Leoparreysia_burmana **   
  +-----+                          
  |     +-- Leoparreysia_whitteni **  
--+                                
  |  +----- Leoparreysia_olivacea **  
  +--+                             
     |  +-- Leoparreysia_tavoyensis **
     +--+                          
        +-- Leoparreysia_canefrii **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
331379002
  +----- Leoparreysia_olivacea **  
--+                             
  |  +-- Leoparreysia_canefrii **  
  +--+                          
     +-- Leoparreysia_tavoyensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3313196004
  +----- Leoparreysia_canefrii **  
--+                             
  |  +-- Leoparreysia_tavoyensis **
  +--+                          
     +-- Leoparreysia_olivacea **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3303198003
  +-- Leoparreysia_tavoyensis **
  |                          
--+-- Leoparreysia_canefrii **  
  |                          
  +-- Leoparreysia_olivacea **

Konopleva et al., 2020 [Publ 4072], fig. 2

iTtTbiriSoTroTroSiD
33039003
  +-- Leoparreysia_olivacea **  
  |                          
--+-- Leoparreysia_canefrii **  
  |                          
  +-- Leoparreysia_tavoyensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
3324122003
  +-------- Leoparreysia_olivacea **     
--+                                   
  |  +----- Leoparreysia_tavoyensis__2 **
  +--+                                
     |  +-- Leoparreysia_canefrii **     
     +--+                             
        +-- Leoparreysia_tavoyensis__1 **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
331388002
  +----- Leoparreysia_olivacea **  
--+                             
  |  +-- Leoparreysia_canefrii **  
  +--+                          
     +-- Leoparreysia_tavoyensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
331388002
  +----- Leoparreysia_olivacea **  
--+                             
  |  +-- Leoparreysia_canefrii **  
  +--+                          
     +-- Leoparreysia_tavoyensis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3313198004
  +----- Leoparreysia_tavoyensis **
--+                             
  |  +-- Leoparreysia_canefrii **  
  +--+                          
     +-- Leoparreysia_olivacea **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
220213700--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
22025500--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Chen et al., 2026 [Publ 4833], fig. 1

iTtTbiriSoTroTroSiD
22022500--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Bolotov et al., 2022 [Publ 4250], fig. S2

iTtTbiriSoTroTroSiD
22021300--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
22022900--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
22025500--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
22022600--
  +-- Leoparreysia_tavoyensis **
--+                          
  +-- Leoparreysia_olivacea **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
220213700--
  +-- Leoparreysia_olivacea **  
--+                          
  +-- Leoparreysia_tavoyensis **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
22023800--
  +-- Leoparreysia_olivacea **  
--+                          
  +-- Leoparreysia_tavoyensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112100--
-- Leoparreysia_tavoyensis **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iTtTbiriSoTroTroSiD
11013500--
-- Leoparreysia_tavoyensis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014400--
-- Leoparreysia_burmana **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Leoparreysia_tavoyensis **

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
1101900--
-- Leoparreysia_canefrii **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Leoparreysia_tavoyensis **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
11013500--
-- Leoparreysia_tavoyensis **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
11012200--
-- Leoparreysia_canefrii **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Leoparreysia_tavoyensis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11018000--
-- Leoparreysia_tavoyensis **

Graf et al., 2014 [Publ 3591], fig. 3

iTtTbiriSoTroTroSiD
11011000--
-- Leoparreysia_tavoyensis **
 
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