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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

genus Yaukthwa Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan & Bolotov, 2019: cladomics
11 Recent valid species (11 total species)

Cladomics Summary

number of species = 11 (11 total)
analyzed in 33 cladograms in 21 publications
maximum number of terminals in an analysis = 10 (mean = 3.94, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 10, (mean = 4.46 median = 4)
proportion of cladograms that support basic monophyly = 0.89

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Bolotov et al., 2022 [Publ 4250], fig. S5

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
10	10	7	10	5	5	2	0

                 +-- Physunio               
  +--------------+                          
  |              +-- Trapezoideus           
  |                                         
  |              +-- Yaukthwa_elongatula **    
--+  +-----------+                          
  |  |           +-- Yaukthwa_baniensis **     
  |  |                                      
  |  |     +-------- Yaukthwa_panhai **        
  +--+  +--+                                
     |  |  |  +----- Yaukthwa_rectangularis ** 
     |  |  +--+                             
     |  |     |  +-- Yaukthwa_nesemanni **     
     +--+     +--+                          
        |        +-- Yaukthwa_zayleymanensis **
        |                                   
        |     +----- Yaukthwa_dalliana **      
        |     |                             
        +-----+  +-- Yaukthwa_paiensis **      
              |  |                          
              +--+-- Yaukthwa_inlenensis **    
                 |                          
                 +-- Yaukthwa_avaensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	14	4	--

        +-- Trapezoideus     
  +-----+                    
  |     +-- Physunio         
--+                          
  |  +----- Yaukthwa_paiensis **
  +--+                       
     |  +-- Solenaia         
     +--+                    
        +-- Lens

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	103	14	3	2

  +----------------------- Physunio               
  |                                               
--+  +-------------------- Trapezoideus           
  |  |                                            
  +--+  +----------------- Lens                   
     |  |                                         
     |  |              +-- Yaukthwa_elongatula **    
     +--+  +-----------+                          
        |  |           +-- Yaukthwa_baniensis **     
        |  |                                      
        +--+        +----- Yaukthwa_inlenensis **    
           |  +-----+                             
           |  |     |  +-- Yaukthwa_avaensis **      
           |  |     +--+                          
           +--+        +-- Yaukthwa_paiensis **      
              |                                   
              |  +-------- Yaukthwa_panhai **        
              +--+                                
                 |  +----- Yaukthwa_rectangularis ** 
                 +--+                             
                    |  +-- Yaukthwa_zayleymanensis **
                    +--+                          
                       +-- Yaukthwa_nesemanni **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	109	14	3	2

                    +-- Trapezoideus           
  +-----------------+                          
  |                 +-- Physunio               
  |                                            
--+  +----------------- Lens                   
  |  |                                         
  |  |              +-- Yaukthwa_elongatula **    
  +--+  +-----------+                          
     |  |           +-- Yaukthwa_baniensis **     
     |  |                                      
     +--+        +----- Yaukthwa_inlenensis **    
        |  +-----+                             
        |  |     |  +-- Yaukthwa_avaensis **      
        |  |     +--+                          
        +--+        +-- Yaukthwa_paiensis **      
           |                                   
           |  +-------- Yaukthwa_panhai **        
           +--+                                
              |  +----- Yaukthwa_rectangularis ** 
              +--+                             
                 |  +-- Yaukthwa_zayleymanensis **
                 +--+                          
                    +-- Yaukthwa_nesemanni **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	40	3	1	2

  +----------------- Trapezoideus           
  |                                         
  |              +-- Yaukthwa_elongatula **    
--+  +-----------+                          
  |  |           +-- Yaukthwa_baniensis **     
  |  |                                      
  +--+        +----- Yaukthwa_inlenensis **    
     |  +-----+                             
     |  |     |  +-- Yaukthwa_paiensis **      
     |  |     +--+                          
     +--+        +-- Yaukthwa_avaensis **      
        |                                   
        |  +-------- Yaukthwa_panhai **        
        +--+                                
           |  +----- Yaukthwa_rectangularis ** 
           +--+                             
              |  +-- Yaukthwa_nesemanni **     
              +--+                          
                 +-- Yaukthwa_zayleymanensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 25

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	1	1	1	3

  +-------------- Solenaia              
  |                                     
  |           +-- Yaukthwa_elongatula **   
--+  +--------+                         
  |  |        +-- Yaukthwa_baniensis **    
  |  |                                  
  +--+     +----- Yaukthwa_panhai **       
     |  +--+                            
     |  |  |  +-- Yaukthwa_rectangularis **
     |  |  +--+                         
     +--+     +-- Yaukthwa_nesemanni **    
        |                               
        |  +----- Yaukthwa_avaensis **     
        +--+                            
           |  +-- Yaukthwa_inlenensis **   
           +--+                         
              +-- Yaukthwa_paiensis **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	28	13	3	4

           +----- Lens                  
  +--------+                            
  |        |  +-- Trapezoideus          
  |        +--+                         
--+           +-- Physunio              
  |                                     
  |  +----------- Yaukthwa_elongatula **   
  |  |                                  
  +--+        +-- Yaukthwa_inlenensis **   
     |  +-----+                         
     |  |     +-- Yaukthwa_paiensis **     
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_nesemanni **    
           +--+                         
              +-- Yaukthwa_rectangularis **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	26	9	3	5

  +-------------- Physunio              
  |                                     
  |           +-- Trapezoideus          
--+  +--------+                         
  |  |        +-- Lens                  
  |  |                                  
  +--+        +-- Yaukthwa_paiensis **     
     |  +-----+                         
     |  |     +-- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_rectangularis **
           +--+                         
              +-- Yaukthwa_nesemanni **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	47	11	3	5

        +----- Physunio              
  +-----+                            
  |     |  +-- Trapezoideus          
  |     +--+                         
  |        +-- Lens                  
--+                                  
  |        +-- Yaukthwa_paiensis **     
  |  +-----+                         
  |  |     +-- Yaukthwa_inlenensis **   
  +--+                               
     |  +----- Yaukthwa_panhai **       
     +--+                            
        |  +-- Yaukthwa_nesemanni **    
        +--+                         
           +-- Yaukthwa_rectangularis **

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	13	0	0	7

  +----------- Yaukthwa_inlenensis **   
--+                                  
  |  +-------- Yaukthwa_elongatula **   
  +--+                               
     |  +----- Yaukthwa_panhai **       
     +--+                            
        |  +-- Yaukthwa_nesemanni **    
        +--+                         
           +-- Yaukthwa_rectangularis **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	195	8	2	6

  +----------- Physunio_modelli      
  |                                  
  +----------- Lens                  
  |                                  
  +----------- Physunio_superbus     
  |                                  
  |  +-------- Yaukthwa_inlenensis **   
  +--+                               
--+  |  +----- Yaukthwa_panhai **       
  |  +--+                            
  |     |  +-- Yaukthwa_nesemanni **    
  |     +--+                         
  |        +-- Yaukthwa_rectangularis **
  |                                  
  |        +-- Lens_comptus          
  |  +-----+                         
  |  |     +-- Lens_novoselovi       
  +--+                               
     |  +----- Lens_eximius          
     +--+                            
        |  +-- Lens_contradens       
        +--+                         
           +-- Lens_micropterus

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	135	8	2	6

  +-------------- Physunio              
--+                                     
  |  +----------- Lens                  
  +--+                                  
     |  +-------- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_rectangularis **
           +--+                         
              +-- Yaukthwa_nesemanni **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	195	8	2	6

  +-------------- Physunio              
--+                                     
  |  +----------- Lens                  
  +--+                                  
     |  +-------- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_nesemanni **    
           +--+                         
              +-- Yaukthwa_rectangularis **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	121	8	2	6

  +-------------- Physunio              
--+                                     
  |  +----------- Lens                  
  +--+                                  
     |  +-------- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_nesemanni **    
           +--+                         
              +-- Yaukthwa_rectangularis **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	135	7	2	6

  +-------------- Lens                  
--+                                     
  |  +----------- Physunio              
  +--+                                  
     |  +-------- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_rectangularis **
           +--+                         
              +-- Yaukthwa_nesemanni **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	193	8	2	6

  +-------------- Physunio              
--+                                     
  |  +----------- Lens                  
  +--+                                  
     |  +-------- Yaukthwa_inlenensis **   
     +--+                               
        |  +----- Yaukthwa_panhai **       
        +--+                            
           |  +-- Yaukthwa_nesemanni **    
           +--+                         
              +-- Yaukthwa_rectangularis **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	29	4	3	7

  +-------------- Physunio           
--+                                  
  |  +----------- Trapezoideus       
  +--+                               
     |  +-------- Lens               
     +--+                            
        |  +----- Yaukthwa_elongatula **
        +--+                         
           |  +-- Yaukthwa_inlenensis **
           +--+                      
              +-- Yaukthwa_paiensis **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	34	4	3	7

           +-- Physunio           
  +--------+                      
  |        +-- Trapezoideus       
--+                               
  |  +-------- Lens               
  +--+                            
     |  +----- Yaukthwa_elongatula **
     +--+                         
        |  +-- Yaukthwa_inlenensis **
        +--+                      
           +-- Yaukthwa_paiensis **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
3	7	1	3	34	4	3	12

  +-- Physunio           
  |                      
  +-- Yaukthwa_paiensis **  
  |                      
  +-- Yaukthwa_inlenensis **
--+                      
  +-- Trapezoideus       
  |                      
  +-- Yaukthwa_elongatula **
  |                      
  +-- Lens

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	34	4	3	7

           +-- Physunio           
  +--------+                      
  |        +-- Trapezoideus       
--+                               
  |  +-------- Lens               
  +--+                            
     |  +----- Yaukthwa_elongatula **
     +--+                         
        |  +-- Yaukthwa_inlenensis **
        +--+                      
           +-- Yaukthwa_paiensis **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	34	4	3	7

           +-- Physunio           
  +--------+                      
  |        +-- Trapezoideus       
--+                               
  |  +-------- Lens               
  +--+                            
     |  +----- Yaukthwa_elongatula **
     +--+                         
        |  +-- Yaukthwa_inlenensis **
        +--+                      
           +-- Yaukthwa_paiensis **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	35	4	3	7

           +-- Physunio           
  +--------+                      
  |        +-- Trapezoideus       
--+                               
  |  +-------- Lens               
  +--+                            
     |  +----- Yaukthwa_elongatula **
     +--+                         
        |  +-- Yaukthwa_inlenensis **
        +--+                      
           +-- Yaukthwa_paiensis **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
3	7	5	3	34	4	3	11

  +----------- Yaukthwa_elongatula **
  |                               
--+  +-------- Trapezoideus       
  |  |                            
  +--+     +-- Physunio           
     |  +--+                      
     |  |  +-- Yaukthwa_inlenensis **
     +--+                         
        |  +-- Yaukthwa_paiensis **  
        +--+                      
           +-- Lens

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	10	0	3	80	7	2	15

  +-- Lens_eximius       
  |                      
  +-- Physunio_modelli__1
  |                      
  +-- Yaukthwa_nesemanni ** 
  |                      
  +-- Yaukthwa_panhai **    
  |                      
  +-- Lens_comptus       
  |                      
--+-- Lens_novoselovi    
  |                      
  +-- Physunio_modelli__2
  |                      
  +-- Physunio           
  |                      
  +-- Yaukthwa_inlenensis **
  |                      
  +-- Lens_contradens    
  |                      
  +-- Lens_micropterus

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	43	5	3	7

           +-- Physunio           
  +--------+                      
  |        +-- Trapezoideus       
--+                               
  |  +-------- Lens               
  +--+                            
     |  +----- Yaukthwa_elongatula **
     +--+                         
        |  +-- Yaukthwa_inlenensis **
        +--+                      
           +-- Yaukthwa_paiensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	2	2	--

  +-------- Physunio          
--+                           
  |  +----- Lens              
  +--+                        
     |  +-- Yaukthwa_nesemanni **
     +--+                     
        +-- Yaukthwa_panhai **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	2	2	--

  +-------- Physunio          
--+                           
  |  +----- Lens              
  +--+                        
     |  +-- Yaukthwa_nesemanni **
     +--+                     
        +-- Yaukthwa_panhai **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	80	17	3	--

        +-- Physunio           
  +-----+                      
  |     +-- Trapezoideus       
--+                            
  |  +----- Lens               
  +--+                         
     |  +-- Yaukthwa_nesemanni ** 
     +--+                      
        +-- Yaukthwa_elongatula **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	19	8	3	--

        +-- Trapezoideus      
  +-----+                     
  |     +-- Physunio          
--+                           
  |  +----- Lens              
  +--+                        
     |  +-- Yaukthwa_nesemanni **
     +--+                     
        +-- Yaukthwa_panhai **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

        +-- Trapezoideus     
  +-----+                    
  |     +-- Physunio         
--+                          
  |  +----- Yaukthwa_paiensis **
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Solenaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

        +-- Trapezoideus     
  +-----+                    
  |     +-- Physunio         
--+                          
  |  +----- Yaukthwa_paiensis **
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Solenaia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	120	2	2	--

-- Yaukthwa_nesemanni **

Pfeiffer et al., 2021 [Publ 4126], fig. 18

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	7	2	--

  +----- Yaukthwa_paiensis **
--+                       
  |  +-- Trapezoideus     
  +--+                    
     +-- Physunio

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