MUSSELpdb | valid gen. Pseudocuneopsis cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Pseudocuneopsis Huang, Dai, Chen & Wu et al., 2022: cladomics
7 Recent valid species (7 total species)

Cladomics Summary

number of species = 7 (7 total)
analyzed in 43 cladograms in 26 publications
maximum number of terminals in an analysis = 7 (mean = 2.51, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 7, (mean = 3.24 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Wang et al., 2025 [Publ 4488], fig. 3B

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	58	16	4	0

           +-------- Nodularia                    
  +--------+                                      
  |        |  +----- Schistodesmus                
  |        +--+                                   
  |           |  +-- Tchangsinaia                 
--+           +--+                                
  |              +-- Cuneopsis                    
  |                                               
  |  +-------------- Pseudocuneopsis_capitata **     
  |  |                                            
  +--+  +----------- Pseudocuneopsis_perflora **     
     |  |                                         
     +--+  +-------- Pseudocuneopsis_liuovatus **    
        |  |                                      
        +--+     +-- Pseudocuneopsis_sichuanensis ** 
           |  +--+                                
           |  |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                   
              |  +-- Pseudocuneopsis_wuana **        
              +--+                                
                 +-- Pseudocuneopsis_yangshuoensis **

Chen et al., 2023 [Publ 4302], fig. 1

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	12	6	--

  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Chen et al., 2025 [Publ 4448], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	3	2	1	3

  +-------------- Nodularia                    
  |                                            
--+  +----------- Pseudocuneopsis_perflora **     
  |  |                                         
  +--+  +-------- Pseudocuneopsis_capitata **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_yangshuoensis **
        |  +--+                                
        |  |  +-- Pseudocuneopsis_wuana **        
        +--+                                   
           |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                
              +-- Pseudocuneopsis_sichuanensis **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	80	13	5	1

        +-------- Nodularia                    
        |                                      
  +-----+     +-- Tongchuania                  
  |     |  +--+                                
  |     |  |  +-- Schistodesmus                
  |     +--+                                   
  |        |  +-- Tchangsinaia                 
--+        +--+                                
  |           +-- Cuneopsis                    
  |                                            
  |  +----------- Pseudocuneopsis_capitata **     
  |  |                                         
  +--+  +-------- Pseudocuneopsis_perflora **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_sichuanensis ** 
        |  +--+                                
        |  |  +-- Pseudocuneopsis_yemaoi **       
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4432], fig. 4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	23	20	4	2

     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4401], fig. 4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	23	20	4	2

     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2024 [Publ 4400], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	10	7	2	2

  +-------- Pseudocuneopsis_capitata **     
  |                                      
--+     +-- Pseudocuneopsis_sichuanensis ** 
  |  +--+                                
  |  |  +-- Pseudocuneopsis_yemaoi **       
  +--+                                   
     |  +-- Pseudocuneopsis_wuana **        
     +--+                                
        +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	71	19	4	2

     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	20	7	5	3

  +-------------- Cuneopsis                    
  |                                            
  |           +-- Nodularia                    
  +-----------+                                
--+           +-- Schistodesmus                
  |                                            
  |  +----------- Tchangsinaia                 
  |  |                                         
  +--+----------- Inversiunio                  
     |                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_sichuanensis ** 
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	84	12	4	3

           +----- Schistodesmus                
  +--------+                                   
  |        |  +-- Tchangsinaia                 
  |        +--+                                
--+           +-- Cuneopsis                    
  |                                            
  |  +----------- Nodularia                    
  +--+                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_yangshuoensis **
        +--+                                   
           |  +-- Pseudocuneopsis_sichuanensis ** 
           +--+                                
              +-- Pseudocuneopsis_yemaoi **

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	20	7	5	3

  +-------------------- Cuneopsis                    
  |                                                  
--+                 +-- Nodularia                    
  |  +--------------+                                
  |  |              +-- Schistodesmus                
  +--+                                               
     |  +-------------- Tchangsinaia                 
     +--+                                            
        |  +----------- Inversiunio                  
        +--+                                         
           |  +-------- Pseudocuneopsis_capitata **     
           +--+                                      
              |  +----- Pseudocuneopsis_sichuanensis ** 
              +--+                                   
                 |  +-- Pseudocuneopsis_wuana **        
                 +--+                                
                    +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2024 [Publ 4358], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	2	2	2	3

           +-- Schistodesmus                
  +--------+                                
  |        +-- Nodularia                    
--+                                         
  |  +-------- Pseudocuneopsis_capitata **     
  +--+                                      
     |  +----- Pseudocuneopsis_yangshuoensis **
     +--+                                   
        |  +-- Pseudocuneopsis_sichuanensis ** 
        +--+                                
           +-- Pseudocuneopsis_yemaoi **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	49	10	4	3

     +-------- Pseudocuneopsis_capitata **     
  +--+                                      
  |  |  +----- Pseudocuneopsis_yangshuoensis **
  |  +--+                                   
  |     |  +-- Pseudocuneopsis_yemaoi **       
--+     +--+                                
  |        +-- Pseudocuneopsis_sichuanensis ** 
  |                                         
  |  +-------- Nodularia                    
  +--+                                      
     |  +----- Schistodesmus                
     +--+                                   
        |  +-- Tchangsinaia                 
        +--+                                
           +-- Cuneopsis

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	20	7	5	4

  +----------------- Cuneopsis                    
  |                                               
--+              +-- Nodularia                    
  |  +-----------+                                
  |  |           +-- Schistodesmus                
  +--+                                            
     |  +----------- Tchangsinaia                 
     +--+                                         
        |  +-------- Inversiunio                  
        +--+                                      
           |  +----- Pseudocuneopsis_capitata **     
           +--+                                   
              |  +-- Pseudocuneopsis_yangshuoensis **
              +--+                                
                 +-- Pseudocuneopsis_sichuanensis **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	72	10	4	--

           +-- Pseudocuneopsis_capitata **    
  +--------+                               
  |        +-- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2e

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	16	7	4	--

           +-- Pseudocuneopsis_sichuanensis **
  +--------+                               
  |        +-- Pseudocuneopsis_capitata **    
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Tchangsinaia                
     +--+                                  
        |  +-- Cuneopsis                   
        +--+                               
           +-- Schistodesmus

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	10	4	--

              +-- Pseudocuneopsis_capitata **    
  +-----------+                               
  |           +-- Pseudocuneopsis_sichuanensis **
--+                                           
  |  +----------- Nodularia                   
  |  |                                        
  +--+        +-- Schistodesmus_lampreyanus   
     |  +-----+                               
     |  |     +-- Cuneopsis                   
     +--+                                     
        |  +----- Tchangsinaia                
        +--+                                  
           |  +-- Schistodesmus_xinyuensis    
           +--+                               
              +-- Schistodesmus_spinosus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	10	4	--

  +-------- Nodularia                   
  |                                     
  |     +-- Pseudocuneopsis_capitata **    
--+-----+                               
  |     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Schistodesmus               
  +--+                                  
     |  +-- Tchangsinaia                
     +--+                               
        +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	11	6	--

  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	10	9	5	--

  +-------------- Nodularia                   
  |                                           
--+           +-- Pseudocuneopsis_capitata **    
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_sichuanensis **
  +--+                                        
     |  +-------- Schistodesmus               
     +--+                                     
        |  +----- Cuneopsis                   
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Inversiunio

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	11	6	--

           +-- Tchangsinaia                
  +--------+                               
  |        +-- Inversiunio                 
  |                                        
--+     +----- Cuneopsis                   
  |  +--+                                  
  |  |  |  +-- Pseudocuneopsis_sichuanensis **
  |  |  +--+                               
  +--+     +-- Pseudocuneopsis_capitata **    
     |                                     
     |  +----- Nodularia                   
     +--+                                  
        |  +-- Diaurora                    
        +--+                               
           +-- Schistodesmus

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	12	6	--

  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	10	4	--

     +----- Nodularia                   
  +--+                                  
  |  |  +-- Pseudocuneopsis_capitata **    
  |  +--+                               
--+     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Tchangsinaia                
  +--+                                  
     |  +-- Schistodesmus               
     +--+                               
        +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	37	13	5	--

              +-- Pseudocuneopsis_capitata **    
  +-----------+                               
  |           +-- Pseudocuneopsis_sichuanensis **
--+                                           
  |  +----------- Nodularia                   
  +--+                                        
     |  +-------- Schistodesmus               
     +--+                                     
        |  +----- Cuneopsis                   
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Inversiunio

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	52	17	4	--

           +-- Pseudocuneopsis_capitata **    
  +--------+                               
  |        +-- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Hou et al., 2025 [Publ 4445], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	20	4	--

           +-- Pseudocuneopsis_sichuanensis **
  +--------+                               
  |        +-- Pseudocuneopsis_capitata **    
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	10	4	--

  +-------- Nodularia                   
  |                                     
  |     +-- Pseudocuneopsis_capitata **    
--+-----+                               
  |     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Schistodesmus               
  +--+                                  
     |  +-- Tchangsinaia                
     +--+                               
        +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	11	6	--

  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	11	6	--

              +-- Tchangsinaia                
  +-----------+                               
  |           +-- Inversiunio                 
  |                                           
--+           +-- Pseudocuneopsis_sichuanensis **
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_capitata **    
  +--+                                        
     |  +-------- Cuneopsis                   
     +--+                                     
        |  +----- Nodularia                   
        +--+                                  
           |  +-- Diaurora                    
           +--+                               
              +-- Schistodesmus

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	4	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Hou et al., 2025 [Publ 4432], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	8	2	--

-- Pseudocuneopsis_capitata **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	6	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Tchangsinaia            
        +--+                           
           +-- Cuneopsis

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	5	4	--

        +-- Cuneopsis               
  +-----+                           
  |     +-- Schistodesmus           
--+                                 
  |  +----- Pseudocuneopsis_capitata **
  +--+                              
     |  +-- Nodularia               
     +--+                           
        +-- Tchangsinaia

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	6	4	--

  +-------------- Tchangsinaia            
--+                                       
  |  +----------- Cuneopsis_heudei        
  +--+                                    
     |  +-------- Nodularia               
     +--+                                 
        |  +----- Pseudocuneopsis_capitata **
        +--+                              
           |  +-- Cuneopsis_rufescens     
           +--+                           
              +-- Schistodesmus

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	7	4	--

  +-------- Pseudocuneopsis_capitata **
  |                                 
--+     +-- Cuneopsis               
  |  +--+                           
  |  |  +-- Schistodesmus           
  +--+                              
     |  +-- Tchangsinaia            
     +--+                           
        +-- Nodularia

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	6	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	4	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	4	4	--

        +-- Pseudocuneopsis_capitata **
  +-----+                           
  |     +-- Nodularia               
--+                                 
  |  +----- Schistodesmus           
  +--+                              
     |  +-- Cuneopsis               
     +--+                           
        +-- Tchangsinaia

Wu et al., 2020 [Publ 4147], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	5	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Tchangsinaia

Wu et al., 2024 [Publ 4379], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	12	3	--

  +----- Nodularia               
  |                              
--+  +-- Cuneopsis               
  |  |                           
  +--+-- Tchangsinaia            
     |                           
     +-- Pseudocuneopsis_capitata **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	10	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Tchangsinaia            
        +--+                           
           +-- Cuneopsis

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	6	4	--

  +----------- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Tchangsinaia                
     +--+                                  
        |  +-- Cuneopsis                   
        +--+                               
           +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	6	4	--

  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Tchangsinaia

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