The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini genus Pseudocuneopsis Huang, Dai, Chen & Wu et al., 2022: cladomics 5 Recent valid species (5 total species) Cladomics Summarynumber of species = 5 (5 total) analyzed in 37 cladograms in 22 publications maximum number of terminals in an analysis =
5 (mean = 2.24, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 23 maximum number of included terminals = 5, (mean =
3 median = 2) proportion of cladograms that support basic monophyly
= 1
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Hou et al., 2024 [Publ 4400], fig. 4 ** Top Monophyly Cladogram ** iT | tT | bi | riS | oT | roT | roS | iD | 5 | 5 | 3 | 5 | 71 | 44 | 7 | 0 |
+-------------------- Aculamprotula
|
--+ +----------------- Unio
| |
+--+ +-------------- Middendorffinaia
| |
| | +-------- Nodularia
+--+ +--+
| | | +----- Schistodesmus
| | +--+
| | | +-- Tchangsinaia
+--+ +--+
| +-- Cuneopsis
|
| +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Pseudocuneopsis_sichuanensis **
| +--+
| | +-- Pseudocuneopsis_yemaoi **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Dai et al., 2025 [Publ 4440], fig. 1 ** Top Sister-Group Cladogram ** iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 43 | 20 | 10 | -- |
+----------------------- Globunio
|
--+ +-------------------- Aculamprotula
| |
+--+ +----------------- Middendorffinaia
| |
+--+ +-------------- Nodularia
| |
| | +----------- Cuneopsis
+--+ |
| | +-- Diaurora
| +--------+
+--+ +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis **
| +-----+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----- Unio
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Hou et al., 2024 [Publ 4400], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 5 | 5 | 3 | 5 | 10 | 8 | 3 | 0 |
+----------- Tchangsinaia
|
+-----+ +-- Cuneopsis_kiangsiensis
| | +-----+
| | | +-- Cuneopsis_celtiformis
| +--+
| | +----- Cuneopsis_szechenyii
| +--+
--+ | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_demangei
|
| +-------------- Cuneopsis_rufescens
| |
+--+ +----------- Middendorffinaia
| |
+--+ +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Pseudocuneopsis_sichuanensis **
| +--+
| | +-- Pseudocuneopsis_yemaoi **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Hou et al., 2025 [Publ 4432], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 5 | 5 | 3 | 5 | 23 | 21 | 5 | 0 |
+-------------- Middendorffinaia
|
| +-------- Nodularia
--+ +--+
| | | +----- Schistodesmus
| | +--+
| | | +-- Tchangsinaia
+--+ +--+
| +-- Cuneopsis
|
| +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Pseudocuneopsis_sichuanensis **
| +--+
| | +-- Pseudocuneopsis_yemaoi **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Hou et al., 2025 [Publ 4401], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 5 | 5 | 3 | 5 | 23 | 21 | 5 | 0 |
+-------------- Middendorffinaia
|
| +-------- Nodularia
--+ +--+
| | | +----- Schistodesmus
| | +--+
| | | +-- Tchangsinaia
+--+ +--+
| +-- Cuneopsis
|
| +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Pseudocuneopsis_sichuanensis **
| +--+
| | +-- Pseudocuneopsis_yemaoi **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Liu et al., 2023 [Publ 4310], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 4 | 4 | 2 | 4 | 20 | 10 | 7 | 1 |
+-------------------------- Unio
|
--+ +----------------------- Aculamprotula
| |
+--+ +-------------------- Cuneopsis
| |
+--+ +-- Nodularia
| +--------------+
| | +-- Schistodesmus
+--+
| +-------------- Tchangsinaia
+--+
| +----------- Inversiunio
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Dai et al., 2025 [Publ 4440], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 4 | 4 | 2 | 4 | 84 | 25 | 8 | 1 |
+----------------------- Globunio
|
--+ +-------------------- Aculamprotula
| |
+--+ +----------------- Unio
| |
| | +----- Schistodesmus
+--+ +--------+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +----------- Nodularia
+--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_yangshuoensis **
+--+
| +-- Pseudocuneopsis_sichuanensis **
+--+
+-- Pseudocuneopsis_yemaoi **
Dai et al., 2024 [Publ 4358], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 4 | 4 | 2 | 4 | 48 | 13 | 6 | 1 |
+----------------- Aculamprotula
|
--+ +-------------- Unio
| |
| | +-------- Pseudocuneopsis_capitata **
+--+ +--+
| | | +----- Pseudocuneopsis_yangshuoensis **
| | +--+
| | | +-- Pseudocuneopsis_yemaoi **
+--+ +--+
| +-- Pseudocuneopsis_sichuanensis **
|
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Liu et al., 2023 [Publ 4310], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 4 | 4 | 2 | 4 | 20 | 10 | 7 | 1 |
+----------------- Unio
|
+----------------- Aculamprotula
|
--+ +-------------- Cuneopsis
| |
| | +-- Nodularia
| +-----------+
+--+ +-- Schistodesmus
|
| +----------- Tchangsinaia
| |
+--+----------- Inversiunio
|
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_sichuanensis **
+--+
| +-- Pseudocuneopsis_wuana **
+--+
+-- Pseudocuneopsis_yangshuoensis **
Dai et al., 2024 [Publ 4358], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 4 | 4 | 2 | 4 | 2 | 2 | 2 | 1 |
+-- Schistodesmus
+--------+
| +-- Nodularia
--+
| +-------- Pseudocuneopsis_capitata **
+--+
| +----- Pseudocuneopsis_yangshuoensis **
+--+
| +-- Pseudocuneopsis_sichuanensis **
+--+
+-- Pseudocuneopsis_yemaoi **
Wu et al., 2023 [Publ 4296], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 3 | 3 | 1 | 3 | 20 | 10 | 7 | 2 |
+----------------------- Unio
|
--+ +-------------------- Aculamprotula
| |
+--+ +----------------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----------+
| | +-- Schistodesmus
+--+
| +----------- Tchangsinaia
+--+
| +-------- Inversiunio
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Pseudocuneopsis_yangshuoensis **
+--+
+-- Pseudocuneopsis_sichuanensis **
Wu et al., 2022 [Publ 4238], fig. 2e iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 15 | 10 | 6 | -- |
+----------------- Aculamprotula
|
--+ +-------------- Unio
| |
+--+ +-- Pseudocuneopsis_sichuanensis **
| +--------+
| | +-- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Dai et al., 2025 [Publ 4440], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 43 | 20 | 10 | -- |
+-------------------- Globunio
|
--+ +----------------- Aculamprotula
| |
| | +-- Middendorffinaia
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
| +--------+
+--+ +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis **
| +-----+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Wu et al., 2022 [Publ 4238], fig. 2d iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 36 | 21 | 7 | -- |
+-------------------- Aculamprotula
|
--+ +----------------- Unio
| |
+--+ +-- Pseudocuneopsis_capitata **
| +-----------+
| | +-- Pseudocuneopsis_sichuanensis **
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 93 | 20 | 6 | -- |
+-------------- Aculamprotula
|
--+ +----------- Unio
| |
| | +----- Nodularia
+--+ +--+
| | | +-- Pseudocuneopsis_capitata **
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis **
|
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 89 | 20 | 6 | -- |
+-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
| | +-- Pseudocuneopsis_capitata **
+--+-----+
| +-- Pseudocuneopsis_sichuanensis **
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 89 | 20 | 6 | -- |
+-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
| | +-- Pseudocuneopsis_capitata **
+--+-----+
| +-- Pseudocuneopsis_sichuanensis **
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Chen et al., 2023 [Publ 4302], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 38 | 15 | 9 | -- |
+----------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
| +--------+
+--+ +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis **
| +-----+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 93 | 20 | 6 | -- |
+-------------------- Aculamprotula
|
--+ +----------------- Unio
| |
+--+ +-- Pseudocuneopsis_capitata **
| +-----------+
| | +-- Pseudocuneopsis_sichuanensis **
+--+
| +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus__1
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_lampreyanus__2
+--+
+-- Schistodesmus_spinosus
Chen et al., 2023 [Publ 4302], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 38 | 15 | 9 | -- |
+----------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
| +--------+
+--+ +-- Schistodesmus
|
| +-- Pseudocuneopsis_sichuanensis **
| +-----+
| | +-- Pseudocuneopsis_capitata **
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 10 | 10 | 6 | -- |
+----------------- Aculamprotula
|
--+ +-------------- Nodularia
| |
+--+ +-- Pseudocuneopsis_capitata **
| +--------+
| | +-- Pseudocuneopsis_sichuanensis **
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Wu et al., 2024 [Publ 4379], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 51 | 27 | 6 | -- |
+----------------- Aculamprotula
|
--+ +-------------- Unio
| |
+--+ +-- Pseudocuneopsis_capitata **
| +--------+
| | +-- Pseudocuneopsis_sichuanensis **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2022 [Publ 4238], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 2 | 2 | 0 | 2 | 71 | 18 | 6 | -- |
+----------------- Aculamprotula
|
--+ +-------------- Unio
| |
+--+ +-- Pseudocuneopsis_capitata **
| +--------+
| | +-- Pseudocuneopsis_sichuanensis **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 3-BI iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 72 | 14 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2020 [Publ 4121], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 48 | 13 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Dai et al., 2024 [Publ 4386], fig. 6 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 39 | 19 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4388], fig. 6 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 53 | 8 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_sichuanensis **
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4147], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 28 | 9 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2021 [Publ 4366], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 67 | 14 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 3-ML iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 72 | 14 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Dai et al., 2023 [Publ 4285], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 52 | 6 | 6 | -- |
+-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-- Pseudocuneopsis_capitata **
| +-----+
| | +-- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Hou et al., 2025 [Publ 4432], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 10 | 9 | 3 | -- |
+-------- Tchangsinaia
--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Middendorffinaia
+--+
+-- Cuneopsis
Wu et al., 2019 [Publ 3864], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 45 | 12 | 6 | -- |
+----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis_capitata **
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2024 [Publ 4379], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 11 | 11 | 3 | -- |
+----- Nodularia
|
--+ +-- Cuneopsis
| |
+--+-- Tchangsinaia
|
+-- Pseudocuneopsis_capitata **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 52 | 11 | 6 | -- |
+-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Zhang et al., 2004 [Publ 2735], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 13 | 6 | 4 | -- |
+-------------- Tchangsinaia
--+
| +----------- Cuneopsis_heudei
+--+
| +-------- Nodularia
+--+
| +----- Pseudocuneopsis_capitata **
+--+
| +-- Cuneopsis_rufescens
+--+
+-- Schistodesmus
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 15 | 11 | 6 | -- |
+----------- Unio
|
--+ +-------- Pseudocuneopsis_capitata **
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
|