The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Cyprogenia Agassiz, 1852: cladomics
2 Recent valid species (2 total species) Cladomics Summarynumber of species = 2 (2 total)
analyzed in 15 cladograms in 9 publications
maximum number of terminals in an analysis =
2 (mean = 1.6, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 9
maximum number of included terminals = 2, (mean =
2 median = 2)
proportion of cladograms that support basic monophyly
= 0.33
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Serb, 2006 [Publ 2678], fig. 2 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 8 | 5 | 5 | 4 | 0 | +-------------- Obliquaria
|
| +-- Ptychobranchus
+-----------+
--+ +-- Lampsilis
|
| +-- Cyprogenia_aberti__6 **
| +--------+
| | +-- Dromus
+--+
| +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_aberti__4 **
| |
| | +-- Cyprogenia_stegaria__1 **
+--+ |
| +-- Cyprogenia_stegaria__2 **
| |
+--+-- Cyprogenia_aberti__1 **
|
+-- Cyprogenia_aberti__2 **
|
+-- Cyprogenia_aberti__3 **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 4 | 630 | 91 | 26 | 6 | +-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
+--+ | +--+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
+--+ | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2 **
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2 **
| +--+
| | +-- Cyprogenia_aberti__1 **
| +--+
| +-- Cyprogenia_stegaria__1 **
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Serb, 2006 [Publ 2678], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 7 | 5 | 5 | 4 | 3 | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 100 | 41 | 22 | -- | +----- Obliquaria
+-----------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Sagittunio
| |
| +-------------- Leaunio
| |
| +-------------- Potamilus
| |
| +-------------- Hamiota
| |
| | +-- Ellipsaria
--+ +-----------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +--------+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +--------+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+-----+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+-----+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +--------+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +-------- Medionidus_conradicus
| |
| | +-- Lemiox
+--+-----+
| +-- Medionidus_acutissimus
|
| +----- Dromus
+--+
| +-- Cyprogenia_aberti **
+--+
+-- Cyprogenia_stegaria **
Grobler et al., 2011 [Publ 3359], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 14 | 9 | -- | +----------- Lemiox
+--+
| | +-------- Medionidus
| +--+
| | +----- Dromus
| +--+
| | +-- Cyprogenia_stegaria **
--+ +--+
| +-- Cyprogenia_aberti **
|
| +----------- Potamilus
| |
| | +----- Lampsilis_siliquoidea
| | +--+
+--+ | | +-- Lampsilis_higginsii
| | +--+
| | +-- Ortmanniana
| |
| | +----- Lampsilis_fasciola
+--+--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-- Cambarunio
| |
+-----+-- Ligumia
|
+-- Epioblasma
Serb & Barnhart, 2008 [Publ 2674], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 8 | 5 | 5 | 4 | 2 | +-------------------------- Lampsilis
|
--+ +----------------------- Obliquaria
| |
+--+ +-------------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__4 **
| |
+--+ +----------- Cyprogenia_aberti__3 **
| |
+--+ +-------- Cyprogenia_aberti__5 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Serb, 2006 [Publ 2678], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 7 | 5 | 5 | 4 | 3 | +-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__5 **
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Serb & Barnhart, 2008 [Publ 2674], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 8 | 5 | 5 | 4 | 2 | +----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Ptychobranchus
| |
+--+ +-- Cyprogenia_aberti__6 **
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__5 **
| |
+--+ +----------- Cyprogenia_aberti__4 **
| |
+--+ +-------- Cyprogenia_aberti__3 **
| |
+--+ +----- Cyprogenia_stegaria__2 **
| |
+--+----- Cyprogenia_aberti__2 **
|
| +-- Cyprogenia_stegaria__1 **
+--+
+-- Cyprogenia_aberti__1 **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 100 | 41 | 22 | -- | +-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------------- Potamilus
| |
| | +-- Sagittunio
--+ | +--+
| +--------+ | +-- Medionidus_conradicus
| | | +--------+
| | | | | +-- Ptychobranchus
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis__2
| | |
| | | +----------- Medionidus_acutissimus
| | +--+
| | | +-------- Lemiox
| | +--+
+--+ | +----- Dromus
| +--+
| | +-- Cyprogenia_aberti **
| +--+
| +-- Cyprogenia_stegaria **
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 47 | 22 | -- | +-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia_stegaria **
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Roe, 2013 [Publ 3571], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 10 | 5 | -- | +----------- Obliquaria
|
--+----------- Lampsilis
|
| +-------- Medionidus
+--+
| +----- Ptychobranchus
+--+
| +-- Cyprogenia_aberti **
+--+
+-- Dromus
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia_stegaria **
| | +--+
| | +-- Ptychobranchus
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- | +----- Medionidus
--+
| +-- Dromus
+--+
+-- Cyprogenia_aberti **
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- | +----- Medionidus
--+
| +-- Dromus
+--+
+-- Cyprogenia_aberti **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 47 | 22 | -- | +-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia_stegaria **
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
|
