The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Unionina genus Unio Philipsson in Retzius, 1788: cladomics
29 Recent valid species (45 total species) Cladomics Summarynumber of species = 29 (45 total)
analyzed in 229 cladograms in 134 publications
maximum number of terminals in an analysis =
26 (mean = 3.47, median = 3) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 160
maximum number of included terminals = 26, (mean =
4.54 median = 3)
proportion of cladograms that support basic monophyly
= 0.94
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 26 | 26 | 24 | 27 | 606 | 0 | 0 | 0 | +-------------------------------- Unio_tumidus **
|
| +-- Unio_durieui **
| +--------------------------+
| | +-- Unio_gibbus **
--+ |
| | +----------------------- Unio_tumidiformis **
| | |
| | +--+ +-------------------- Unio_sesirmensis **
| | | | |
| | | +--+ +----------------- Unio_desectus **
+--+ | | |
| | +--+ +-------------- Unio_damascensis **
| | | |
| | +--+ +----------- Unio_ionicus **
| | | |
| | | | +-- Unio_carneus **
| | +--+ +-----+
| | | | +-- Unio_nanus **
| | | |
+--+ | | +-- Unio_crassus **
| +--+-----+
| | +-- Unio_gontierii **
| |
| | +----- Unio_vicarius **
| +--+
| | +-- Unio_bruguierianus **
| +--+
| +-- Unio_mardinensis **
|
| +-- Unio_delphinus **
| +--------------------+
| | +-- Unio_foucauldianus **
+--+
| +-------------------- Unio_ravoisieri **
| |
+--+ +----------------- Unio_mancus **
| |
+--+ +-------------- Unio_elongatulus **
| |
| | +-- Unio_hueti **
+--+ +--------+
| | +-- Unio_delicatus__2 **
| |
+--+ +-- Unio_eucirrus **
| +-----+
| | +-- Unio_pictorum **
+--+
| +----- Unio_caffer **
+--+
| +-- Unio_terminalis **
+--+
+-- Unio_delicatus__1 **
Lyubas et al., 2022 [Publ 4255], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 20 | 23 | 8 | 0 | 0 | 28 | +----------------------------- Unio_tumidus **
|
| +-- Unio_durieui **
| +-----------------------+
| | +-- Unio_gibbus **
--+ |
| | +-------------- Unio_damascensis **
| | |
| | +-------------- Unio_sesirmensis **
| | +--------+
+--+ | | +----------- Unio_ionicus **
| | | |
| | +--+ +-- Unio_bruguierianus **
| | | +-----+
| | | | +-- Unio_tumidiformis **
| | +--+
| | | +----- Unio_vicarius **
+--+ +--+
| | +-- Unio_carneus **
| +--+
| +-- Unio_crassus **
|
| +----- Unio_caffer **
| +--------------+
| | | +-- Unio_terminalis **
| | +--+
+--+ +-- Unio_delicatus **
|
| +----------------- Unio_pictorum **
| |
+--+ +-------------- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--------+
| | +-- Unio_mancus **
+--+
| +-------- Unio_ravoisieri **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
+--+
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Lyubas et al., 2022 [Publ 4255], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 14 | 23 | 8 | 0 | 0 | 22 | +----------------- Unio_tumidus **
|
| +-- Unio_durieui **
+--------------+
| +-- Unio_gibbus **
|
| +----------- Unio_sesirmensis **
| |
| +--+ +-------- Unio_damascensis **
--+ | | |
| | | | +----- Unio_ionicus **
| | +--+ |
| | | | +-- Unio_bruguierianus **
| | | +--+
| | +--+ +-- Unio_tumidiformis **
| | |
| | | +-- Unio_carneus **
| | | |
| | +--+-- Unio_crassus **
+--+ |
| +-- Unio_vicarius **
|
| +-------- Unio_elongatulus **
| |
| +-------- Unio_mancus **
| |
| | +-- Unio_pictorum **
| +-----+
| | +-- Unio_eucirrus **
| |
+-----+ +----- Unio_caffer **
+--+
| | +-- Unio_terminalis **
| +--+
| +-- Unio_delicatus **
|
| +----- Unio_ravoisieri **
| |
| | +-- Unio_delphinus **
+--+--+
| +-- Unio_foucauldianus **
|
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 19 | 57 | 0 | 0 | 23 | +-- Unio_gibbus **
+-----------------------------+
| +-- Unio_tumidus **
|
| +----------- Unio_terminalis **
--+ +-----------------+
| | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_pictorum **
| | +--+
+--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_mancus **
|
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_desectus **
| |
+--+ +-------------------- Unio_sesirmensis **
| |
+--+ +----------------- Unio_ionicus **
| |
+--+ +-------------- Unio_damascensis **
| |
| | +-- Unio_crassus **
+--+ +--------+
| | +-- Unio_gontierii **
| |
+--+ +-- Unio_bruguierianus **
| +-----+
| | +-- Unio_mardinensis **
+--+
| +----- Unio_vicarius **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 19 | 122 | 0 | 0 | 23 | +----------------------------- Unio_tumidus **
|
| +-------------------------- Unio_gibbus **
--+ |
| | +----------- Unio_terminalis **
| | +-----------+
+--+ | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_mancus **
| | +--+
+--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_pictorum **
|
| +-------------------- Unio_tumidiformis **
| |
+--+ +----------------- Unio_desectus **
| |
| | +----- Unio_damascensis **
+--+ +--------+
| | | +-- Unio_sesirmensis **
| | +--+
| | +-- Unio_ionicus **
+--+
| +-- Unio_crassus **
| +--------+
| | +-- Unio_gontierii **
+--+
| +-------- Unio_vicarius **
| |
+--+ +-- Unio_bruguierianus **
| +--+
| | +-- Unio_mardinensis **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 17 | 5 | 0 | 0 | 31 | +-------------------------- Unio_gibbus **
|
--+ +----------------------- Unio_tumidus **
| |
| | +----- Unio_terminalis **
+--+ +--------------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +----------------- Unio_tumidiformis **
| |
| | +-- Unio_ionicus **
+--+ +-----------+
| | +-- Unio_carneus **
| |
| | +-- Unio_damascensis **
+--+ +--------+
| | +-- Unio_sesirmensis **
| |
| | +----- Unio_bruguierianus **
+--+ +--+
| | | +-- Unio_gontierii **
| | +--+
| | +-- Unio_mardinensis **
+--+
| +-- Unio_crassus **
| +--+
| | +-- Unio_nanus **
+--+
| +-- Unio_vicarius **
+--+
+-- Unio_desectus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 17 | 5 | 0 | 0 | 21 | +-- Unio_gibbus **
+-----------------------------+
| +-- Unio_tumidus **
|
--+ +----- Unio_terminalis **
| +-----------------------+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_sesirmensis **
| |
+--+ +-------------------- Unio_ionicus **
| |
+--+ +----------------- Unio_damascensis **
| |
+--+ +-------------- Unio_desectus **
| |
| | +-- Unio_crassus **
+--+ +--------+
| | +-- Unio_gontierii **
| |
+--+ +-- Unio_bruguierianus **
| +-----+
| | +-- Unio_mardinensis **
+--+
| +----- Unio_vicarius **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 16 | 9 | 0 | 0 | 25 | +----------------------- Unio_tumidus **
|
| +-------- Unio_damascensis **
| +-----------+
--+ | | +----- Unio_sesirmensis **
| | +--+
| | | +-- Unio_bruguierianus **
| | +--+
+--+ +-- Unio_nanus **
|
| +-- Unio_delicatus **
| +--------------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----------+
| | +-- Unio_hueti **
| |
+--+ +----------- Unio_ravoisieri **
| |
| | +-- Unio_pictorum **
+--+ +-----+
| | +-- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_foucauldianus **
+--+
+-- Unio_delphinus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 18 | 0 | 0 | 0 | 15 | +-- Unio_pictorum **
+--------------------------+
| +-- Unio_delphinus **
--+
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_sesirmensis **
| |
+--+ +-------------------- Unio_desectus **
| |
+--+ +----------------- Unio_damascensis **
| |
| | +-------------- Unio_ionicus **
+--+ |
| | +-- Unio_nanus **
| | +--------+
+--+ | +-- Unio_carneus **
| |
| | +-- Unio_gontierii **
| | |
| | +-- Unio_crassus__1 **
+--+--------+
| +-- Unio_crassus__2 **
| |
| +-- Unio_crassus__3 **
|
| +-------- Unio_vicarius **
| |
+--+ +----- Unio_bruguierianus__1 **
| |
+--+----- Unio_bruguierianus__2 **
|
| +-- Unio_bruguierianus__3 **
+--+
+-- Unio_mardinensis **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 9 | 14 | 5 | 0 | 0 | 20 | +-- Unio_durieui **
+--------------+
| +-- Unio_gibbus **
|
--+ +-------------- Unio_tumidus **
| |
| | +-------- Unio_tumidiformis **
| | +--+
+--+ | | +----- Unio_desectus **
| | +--+
| | | +-- Unio__1 **
| | +--+
+--+ +-- Unio_ionicus **
|
| +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
| |
+-----+ +-- Unio_pictorum **
| |
| +-- Unio_elongatulus **
| |
+--+-- Unio_mancus **
|
+-- Unio_ravoisieri **
|
+-- Unio__2 **
Froufe et al., 2016 [Publ 3738], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 12 | 2 | 0 | 0 | 21 | +-- Unio_gibbus **
+-----------------------+
| +-- Unio_durieui **
--+
| +----------------------- Unio_tumidus **
| |
+--+ +-- Unio **
| +-----------------+
| | +-- Unio_tumidiformis **
+--+
| +----------------- Unio_delphinus **
+--+
| +-------------- Unio_foucauldianus **
+--+
| +----------- Unio_ravoisieri **
+--+
| +-------- Unio_mancus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Prié & Puillandre, 2014 [Publ 3529], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 6 | 0 | 0 | 24 | +----- Unio_tumidus **
+-----------+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +----- Unio_tumidiformis **
| +--------+
| | | +-- Unio **
| | +--+
+--+ +-- Unio_crassus **
|
| +----------- Unio_delphinus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 0 | 0 | 0 | 25 | +----------------- Unio_tumidus **
|
--+ +-------------- Unio_bruguierianus **
| |
| | +-- Unio_delicatus **
+--+ +--------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----+
| | +-- Unio_hueti **
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_eucirrus **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 5 | 9 | 8 | 0 | 0 | 26 | +-- Unio_tumidus **
+--------+
| +-- Unio_gibbus **
|
--+ +-- Unio **
| +-----+
| | +-- Unio_tumidiformis **
+--+
| +----- Unio_ravoisieri **
| |
+--+ +-- Unio_mancus **
| |
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
|
+-- Unio_elongatulus **
Khalloufi et al., 2011 [Publ 2748], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 9 | 2 | 0 | 0 | 24 | +----- Unio_tumidus **
+-----+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio **
| |
+--+----- Unio_pictorum **
|
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Marrone et al., 2019 [Publ 3907], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 8 | 0 | 0 | 0 | 23 | +-------------- Unio_tumidus **
|
--+ +----------- Unio_delphinus **
| |
+--+ +-------- Unio_foucauldianus **
| |
+--+ +----- Unio_mancus **
| |
| +----- Unio_elongatulus **
+--+
+----- Unio_ravoisieri **
|
| +-- Unio_pictorum **
+--+
+-- Unio_terminalis **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 8 | 0 | 0 | 24 | +-------------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--------+
| | +-- Unio__2 **
+--+
| +-------- Unio__1 **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 0 | 0 | 0 | 22 | +----------------- Unio_tumidus **
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Unio_sesirmensis **
+--+
| +-------- Unio_damascensis **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio **
+--+
+-- Unio_nanus **
Araujo et al., 2009 [Publ 2589], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 4 | 0 | 0 | 25 | +-------------- Unio_gibbus **
|
--+ +----------- Unio_tumidus **
| |
| | +-- Unio_tumidiformis **
+--+ +-----+
| | +-- Unio_crassus **
| |
+--+ +-- Unio_pictorum__1 **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum__2 **
+--+
+-- Unio_caffer **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 36 | 0 | 0 | 23 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 86 | 0 | 0 | 25 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 3 | 7 | 5 | 0 | 0 | 26 | +-- Unio_gibbus **
|
+-----+-- Unio_tumidus **
| |
| +-- Anodonta
--+
| +----- Unio_pictorum__1 **
| |
| +----- Unio_pictorum__2 **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 6 | 90 | 0 | 0 | 24 | +----------- Unio_tumidus **
|
--+ +-------- Unio_nanus **
| |
+--+ +----- Unio_elongatulus **
| |
+--+ +-- Unio_delphinus **
| |
+--+-- Unio_mancus **
|
+-- Unio_pictorum **
Araujo et al., 2009 [Publ 2589], fig. 7C iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 7 | 4 | 0 | 0 | 24 | +-- Unio_gibbus **
+-----+
| +-- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--+
| | +-- Unio_crassus **
+--+
| +-- Unio_pictorum__1 **
| |
+--+-- Unio_pictorum__2 **
|
+-- Unio_mancus **
Reis et al., 2013 [Publ 3580], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 8 | 0 | 0 | 21 | +----------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio_delphinus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 86 | 0 | 0 | 25 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 82 | 0 | 0 | 23 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 90 | 0 | 0 | 25 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 48 | 0 | 0 | 23 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 47 | 0 | 0 | 23 | +-------------- Unio_tumidus **
--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 7 | 0 | 0 | 24 | +----------- Unio_tumidus **
--+
| +-------- Unio_nanus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Bashê, 2022 [Publ 4265], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 1 | 0 | 0 | 27 | +-------------- Unio_pictorum__2 **
--+
| +----------- Unio_foucauldianus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_tigridis **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum__1 **
Prie et al., 2012 [Publ 3487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 1 | 0 | 0 | 24 | +----------- Unio_tumidus **
--+
| +-------- Unio_crassus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 42 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 38 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 70 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Soroka, 2005 [Publ 2726], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 1 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_caffer **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 25 | +-------- Unio_tumidus **
--+
| +----- Unio **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 196 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 52 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 76 | 0 | 0 | 24 | +----- Unio_gibbus **
|
--+----- Unio_tumidus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 194 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Klymus et al., 2025 [Publ 4814], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 32 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 196 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Klymus et al., 2025 [Publ 4814], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 32 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 25 | +-------- Unio_tumidus **
--+
| +----- Unio **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Nagel et al., 1998 [Publ 1946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 0 | 0 | 0 | 24 | +----- Unio_tumidus **
|
--+ +-- Unio_pictorum **
| |
+--+-- Unio_crassus **
|
+-- Unio_mancus **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 65 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 19 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Nagel et al., 1998 [Publ 1946], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 25 | +-------- Unio_crassus **
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 50 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 92 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 70 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 6 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 118 | 0 | 0 | 24 | +----- Unio_gibbus **
|
--+----- Unio_tumidus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 34 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Klishko et al., 2017 [Publ 3946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 2 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_crassus **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 36 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 16 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 14 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 35 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 82 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Wang et al., 2025 [Publ 4488], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 61 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 13 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 40 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 7 | 0 | 0 | 24 | +----- Unio_tumidus **
|
--+----- Unio_gibbus **
|
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 45 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 43 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 40 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 70 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 44 | 0 | 0 | 23 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Klishko et al., 2018 [Publ 3945], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 4 | 0 | 0 | 24 | +----- Unio_tumidus **
|
--+----- Unio_gibbus **
|
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 11 | 0 | 0 | 24 | +----- Unio_tumidus **
|
--+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 3 | 4 | 0 | 0 | 25 | +-- Unio_tumidus **
|
--+-- Unio_pictorum **
|
+-- Unio_nanus **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 79 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 0 | 0 | 24 | +----- Unio_nanus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 2 | 0 | 0 | 26 | +----- Unio_nanus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Zieritz et al., 2012 [Publ 3673], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 26 | +----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 8b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 135 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Kilikowska et al., 2020 [Publ 4335], fig. 4a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 0 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_nanus **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 53 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 2 | 0 | 0 | 26 | +----- Unio_pictorum **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_crassus **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 15 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 26 | +----- Unio_nanus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Källersjö et al., 2005 [Publ 2684], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 3 | 4 | 0 | 0 | 25 | +-- Unio_crassus **
|
--+-- Unio_pictorum **
|
+-- Unio_tumidus **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 19 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 33 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 10 | 0 | 0 | 26 | +----- Unio_crassus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 15 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 8a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Soroka, 2010 [Publ 3404], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 135 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 3a-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 26 | +----- Unio_nanus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 16 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 19 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 10 | 0 | 3 | 121 | 0 | 0 | 32 | +-- Unio_crassus **
|
+-- Unio_pictorum **
|
+-- Unio_tumidus **
|
+-- Scriptolamprotula
|
+-- Cuneopsis_szechenyii
--+
+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus
Soroka, 2010 [Publ 3404], fig. 3b-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Froufe et al., 2017 [Publ 3802], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 7 | 0 | 0 | 24 | +----- Unio_mancus **
--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 20 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_nanus **
Bogatov et al., 2018 [Publ 3859], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 3 | 3 | 0 | 0 | 26 | +-------- Unio_tumidus **
|
--+ +-- Anodonta
| +--+
| | +-- Pseudanodonta
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Bogatov et al., 2018 [Publ 3859], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 3 | 3 | 0 | 0 | 28 | +----------- Unio_crassus **
--+
| +-------- Unio_pictorum **
+--+
| +----- Unio_tumidus **
+--+
| +-- Anodonta
+--+
+-- Pseudanodonta
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 0 | 0 | 24 | +----- Unio_nanus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 16 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 20 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 0 | 0 | 24 | +----- Unio_nanus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 0 | 0 | 24 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 14 | 0 | 0 | 27 | +-------- Unio_pictorum **
--+
| +----- Anodonta
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 13 | 0 | 0 | 27 | +-- Anodonta
|
+-- Unio_ravoisieri **
--+
+-- Unio_pictorum **
|
+-- Scriptolamprotula
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 32 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 9 | 0 | 2 | 13 | 0 | 0 | 32 | +-- Scriptolamprotula
|
+-- Lamprotula
|
+-- Lampsilis
|
+-- Potomida
|
--+-- Unio_pictorum **
|
+-- Anodonta
|
+-- Unio_ravoisieri **
|
+-- Lamellidens
|
+-- Elliptio
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 30 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_pictorum **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 0 | 0 | -- | +-- Unio_nanus **
--+
+-- Unio_tumidus **
Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_delphinus **
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 2 | 29 | 0 | 0 | 28 | +-------- Unio_pictorum **
|
--+-------- Unio_tumidus **
|
| +----- Anodonta
+--+
| +-- Strophitus
+--+
+-- Pyganodon
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 0 | 0 | -- | +-- Unio_nanus **
--+
+-- Unio_tumidus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 0 | 0 | -- | +-- Unio_nanus **
--+
+-- Unio_delphinus **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 2 | 29 | 0 | 0 | 28 | +----------- Unio_pictorum **
--+
| +-------- Unio_tumidus **
+--+
| +----- Anodonta
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 7 | 0 | 3 | 23 | 0 | 0 | 31 | +-------------- Unio_pictorum__2 **
|
--+ +-- Unio_pictorum__1 **
| +--------+
| | +-- Unio_caffer **
+--+
| +-------- Anodonta
| |
+--+ +----- Lasmigona
| |
+--+ +-- Strophitus
| |
+--+-- Alasmidonta
|
+-- Pyganodon
Soroka, 2010 [Publ 3404], fig. 8b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 1 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 0 | 0 | -- | +-- Unio_mancus **
--+
+-- Unio_elongatulus **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 0 | 0 | -- | +-- Unio_crassus **
--+
+-- Unio_pictorum **
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 8a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 1 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 0 | 0 | -- | +-- Unio_crassus **
--+
+-- Unio_delphinus **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_nanus **
Tomilova et al., 2020 [Publ 4097], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- | +-- Unio_tumidus **
--+
+-- Unio_pictorum **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 37 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- | Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 0 | 0 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Bolotov et al., 2015 [Publ 3646], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 0 | 0 | -- | Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- | Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 0 | 0 | -- | Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 0 | 0 | -- | Klunzinger et al., 2021 [Publ 4086], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Wang et al., 2016 [Publ 3714], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- | Araujo et al., 2009 [Publ 2395], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Choi et al., 2020 [Publ 4169], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Wu et al., 2020 [Publ 4147], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 0 | 0 | -- | Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Taylor et al., 2007 [Publ 2696], fig. 1-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Wu et al., 2026 [Publ 4811], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Combosch et al., 2017 [Publ 3806], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Huang et al., 2018 [Publ 3828], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 0 | 0 | -- | Combosch et al., 2017 [Publ 3806], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Wu et al., 2020 [Publ 4121], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Vikhrev et al., 2018 [Publ 3812], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Bolotov et al., 2015 [Publ 3646], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 0 | 0 | -- | Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- | Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Graf et al., 2015 [Publ 3619], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Graf et al., 2014 [Publ 3591], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Wu et al., 2020 [Publ 4121], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 0 | 0 | -- | Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 0 | 0 | -- | Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 0 | 0 | -- | Nagel et al., 1998 [Publ 1946], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 28 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
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