MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Bivalvia | ... | Unionidae | Unioninae | Unionini | Nodularia

species Nodularia douglasiae (Griffith & Pidgeon, 1833): synonymy
     273 specimens + 225 Genbank vouchers

Unio douglasiae Griffith & Pidgeon, 1833
Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Unio (Nodularia) douglasiaeMartens (1905), Zhadin (1938)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Nodularia douglasiaeHaas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Wu et al. (2024), Hou et al. (2024), Hou et al. (2025), Hou et al. (2025), Dai et al. (2025)
Unio douglasiaeZhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Unio douglasiae douglasiaeHaas (1969)
Petit & Coan (2008)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia cf. douglasiaeHuang et al. (2019)
Nodularia douglasiae Clade CChoi et al. (2020)
Nodularia douglasiae Clade BChoi et al. (2020)
Nodularia douglasiae Clade AChoi et al. (2020)
Nodularia douglasiae douglasiaeBogatov (2022)
Nodularia sp. 2Dai et al. (2024)

Unio murchisonianus Lea, 1834
Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6.
Type(s): HOLOTYPE (MONOTYPY) USNM_84156, China [“Calcutta, India” penciled out].

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio osbeckii Philippi, 1844
Philippi, 1844. Z. Malak. 1: 164.
Philippi, 1847. Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio shanghaiensis Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121.
Type(s): HOLOTYPE BMNH_1965171, Shanghai, China.

Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae shanghaiensisHaas (1911)
Nodularia douglasiaeHaas (1920), Lopes-Lima et al. (2020)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio wrightii Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97.
Type(s): HOLOTYPE USNM_84129, China.

Nodularia (s.s.) douglasiaeSimpson (1900), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio wrigtii [sic]de Gregorio (1914)
Unio douglasiae douglasiaeHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio sculptus Deshayes, 1873
Deshayes, 1873. Nouvelles archives 9: 9, pl. 1, figs. 3-3a.
Type(s): SYNTYPES MNHN-IM-2000-1779, Petchily, Chine Centrale.

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Unio sculptusYu et al. (2001)
Nodularia douglasiaeProzorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio biwae Kobelt, 1879
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4.
Type(s): LECTOTYPE SMF_3650, Biwa-See, Japan.

Nodularia (s.s.) biwaeSimpson (1900), Haas (1911), Simpson (1914)
Unio biwaeHaas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae biwaeKondo (1997), Kondo (2008)
Nodularia biwaeBogan (2010)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae biwaeSano et al. (2017), Mabuchi & Nishida (2020)
Nodularia douglasiaeLopes-Lima et al. (2020), Mabuchi et al. (2021)

Unio douglasiae chloreus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Nodularia (s.s.) douglasiaeSimpson (1900)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae dactylinus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128e.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae deltae Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’].
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio favidens var. deltae Benson, 1862)

Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae firmus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)

Nodularia (s.s.) douglasiaeSimpson (1900)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae hainanensis Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128f.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio jourdyi Morlet, 1886
Morlet, 1886. J. de Conch. 34: 76, pl. 13, figs. 5-5a.
Type(s): SYNTYPES MNHN_MP_0150, Bac Hat etang de la riviere Claire.

Nodularia (s.s.) jourdyiSimpson (1900), Simpson (1914)
Unio jourdyiDautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Oxynaia jourdyiHaas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Nodularia jourdyiBolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021)
Nodularia douglasiaeWu et al. (2024)

Unio pictorum var. amurensis Mousson, 1887
Mousson, 1887. J. de Conch. 35: 26, pl. 1, fig. 8.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)

Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae amurensisHaas (1969)
Nodularia amurensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) amurensisZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae amurensisBogatov (2022)

Unio middendorffi Westerlund, 1890
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 113.
Type(s): NEOTYPE BIV229_5 (by Bolotov et al., 2020: Table S1), Gladkaya River, Primorye Region, Russia.

Unio pictorum middendorffiHaas (1940)
Unio douglasiae amurensisHaas (1969)
Nodularia middendorffiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae middendorffiBogatov (2022)

Unio pliculosus Martens, 1894
Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b.
Type(s): TYPE ZMB_55620, Singei, Korea.

Simpson (1900)
Unio (Nodularia) pliculosusMartens (1905)
Nodularia (s.s.) pliculosusHaas (1911)
Nodularia (s.s.) pliculosaSimpson (1914)
Unio douglasiae pliculosusHaas (1969)
Unio douglasiaeHabe (1977)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio verrucifer Martens, 1894
Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b.
Type(s): TYPE ZMB_55622, Hangang.

Simpson (1900)
Unio (Nodularia) verruciferMartens (1905)
Nodularia (s.s.) verruciferaHaas (1911), Simpson (1914)
Unio douglasiae verruciferHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio abbreviatus Westerlund, 1897
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 132.
(homonym of Unio requienii var. abbreviatus de Joannis, 1859)
Type(s): PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio (Nodularia) douglasiae var. schrenki [sic]Zhadin (1938)
Unio douglasiae var. abbreviatusZhadin (1965)
Unio douglasiae amurensisHaas (1969)
Nodularia abbreviataMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) abbreviataZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrenckiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Unio schrencki Westerlund, 1897
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 131.
Type(s): PARATYPE SMF_3663, Amur - Fluss.

Nodularia (s.s.) douglasiaeSimpson (1900), Simpson (1914)
Unio (Nodularia) douglasiae var. schrenki [sic]Zhadin (1938)
Unio douglasiae var. schrenckiZhadin (1965)
Unio douglasiae amurensisHaas (1969)
Nodularia schrenckiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia (s.s.) schrenckiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Unio boettgeri Neumayr, 1899
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’].
(homonym of Unio firmus var. boettgeri Ihering, 1893)

Simpson (1900), Haas (1969)
Nodularia (s.s.) boettgeriSimpson (1914)
Nodularia douglasiaeHaas (1920)
Nodularia (s.s.) douglasiaeHaas (1924)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio kobelti Neumayr, 1899
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1.
(homonym of Unio kobelti Kobelt & Rolle, 1895)

Unio kobeltiSimpson (1914)
Nodularia douglasiaeHaas (1920)
Nodularia (s.s.) douglasiaeHaas (1924)
Unio douglasiaeLiu et al. (1979)
Nodularidia douglasiaeHe & Zhuang (2013)

Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905
Pilsbry & Hirase, 1905. Proc. Acad. Nat. Sci. 57: 750.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): HOLOTYPE ANSP_58170, Formosa.

Nodularia (s.s.) douglasiae var. taiwanicaSimpson (1914)
Nodularia (s.s.) douglasiae taiwanicaHaas (1924)
Unio douglasiae taiwanicusKuroda (1941)
Unio douglasiae taiwanicaHaas (1969)
Unio douglasiaeHabe (1977), Liu et al. (1979), Wu (1982)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio jourdyi var. corrugata Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905. J. de Conch. 53: 205.
(homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0149, Bas-Luc Nam.

Unio jourdyi var. corrugataDautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. corrugataSimpson (1914)

Unio jourdyi var. ponderosa Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905. J. de Conch. 53: 453.
(homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.

Unio jourdyi var. ponderosaDautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. ponderosaSimpson (1914)

Nodularia denserugata Haas, 1910
Haas, 1910. Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.

Nodularia (s.s.) denserugataHaas (1911), Simpson (1914)
Unio denserugatusHaas (1969)
Middendorffinaia (Sujfununio) denserugataMoskvicheva & Starobogatov (1973)
Nodularia denserugataGraf & Cummings (2007), Bogan (2010)
Ptychorhynchus denserugataHe & Zhuang (2013), Zieritz et al. (2018)
Ptychorhynchus denserugatumGraf & Cummings (2021)
Nodularia douglasiaeWu et al. (2024)

Nodularia verrucosa Haas, 1910
Haas, 1910. Nachr. Bl. Dtsch. Malak. Ges. 42: 99.
Type(s): TYPE SMF_3660, Hunan, M. China.

Nodularia verrucosaHaas (1910), Simpson (1914)
Pilsbry & Bequaert (1927), Haas (1936), Haas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)

Nodularia douglasiae crassidens Haas, 1910
Haas, 1910. Ann. Mag. Nat. Hist. 6: 499.
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 75.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3659, Hunan, M. China.

Nodularia (s.s.) douglasiae crassidensHaas (1911)
Nodularia (s.s.) douglasiae var. crassidensSimpson (1914)
Unio douglasiae douglasiaeHaas (1969)
Nodularia douglasiae crassidensDang et al. (1980), Do et al. (2018)
Nodularia crassidensProzorova et al. (2005)
Nodularia dorriIUCN (2012)
Nodularidia douglasiaeHe & Zhuang (2013)

Nodularia douglasiae flavoviridis Haas, 1910
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3654, W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.

Unio douglasiae amurensisHaas (1969)
Nodularia flavoviridisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) flavoviridisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrenckiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)

Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1.
‘Moskv. et Star.’ Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 41.
(incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973)

Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973
Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п.
(correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)

Middendorffinaia (s.s.) arsenieviMoskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (s.s.) arsenjevi [sic]Zatravkin (1983)
Middendorffinaia arsenieviKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicusGraf (2007)
Middendorffinaia (s.s.) ussuriensisBogatov (2012), Vinarski & Kantor (2016)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ussuriensisBogatov (2022)

Nodularia sujfunica Moskvicheva, 1973
Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 с-т.

Nodularia sujfunicaMoskvicheva (1973), Zatravkin (1983)
Nodularia (s.s.) sujfunicaZatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensisStarobogatov et al. (2004)
Nodularia vladivostokensisKantor & Sysoev (2005)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia suifunica [sic]Klishko et al. (2018)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae schrenckiBogatov (2022)

Nodularia vladivostokensis Moskvicheva, 1973
Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 п-р.

Nodularia vladivostokensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) vladivostokensisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae douglasiaeBogatov (2022)

Nodularia lebedevi Zatravkin & Starobogatov, 1984
Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1787, figs. 5-8.

Nodularia sp.n.Zatravkin (1983)
Nodularia (Amurunio) lebedeviZatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia lebedeviKantor & Sysoev (2005), Kantor et al. (2010)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) middendorffiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia middendorffiVinarski & Kantor (2016)
Nodularia douglasiae middendorffiBogatov (2022)

Scabies longata Liu et al., 1991
Liu et al., 1991. Acta Zootax. Sinica 16(3): 279 [281], fig’d.

Scabies longataHu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Scabies crispataHe & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiaeDai et al. (2024)

Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992
Martynov & Chernyshev, 1992. Zool. Zhur. 71: 19, figs. 1-3.

Nodularia (Magadaninaia) extremalisMartynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia extremalisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992
Bogatov & Starobogatov, 1992. Zool. Zhur. 71: 134, fig’d.

Nodularia (s.s.) moskvichevaeStarobogatov et al. (2004)
Nodularia moskvichevaeKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae middendorffiBogatov (2022)

Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000
Bogatov, 2000. Zool. Zhur. 79: 861, text fig. a-r.

Middendorffinaia (s.s.) ochoticaStarobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Middendorffinaia ochoticaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicusGraf (2007)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ochoticaBogatov (2022)

Nodularia sakhalinensis Bogatov, 2001
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 75.

Nodularia (s.s.) sakhalinensisStarobogatov et al. (2004)
Nodularia sakhalinensisKantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Acuticosta jianghanensis He & Zhuang, 2013
He & Zhuang, 2013. Freshwater Bivalves China: 23, fig. 47-48.

Acuticosta jianghanensisHe & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiaeBogatov & Prozorova (2017)

Nodularia douglasiae nipponensis Sano et al., 2017
‘Martens’ of Authors.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part]
(misidentified reference to Unio nipponensis Martens, 1877)

Nodularia douglasiaeSano et al. (2020), Choi et al. (2020)

Nodularia suifunica Klishko et al., 2018
‘Moskvicheva’ of Authors.
Klishko et al., 2018. Systemat. Biodiv. 16(3): 288.
Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 59.
(error for Nodularia sujfunica Moskvicheva, 1973)

Unio ezonis of Authors, unknown year
‘Kuroda, MS’ in ANSP coll. of Authors.

Unio föttgeri of Authors, unknown year
of Authors.

Unio douglasiaeLiu et al. (1979)

Margarita (Unio) merchisonianus Lea, 1836
Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata].
(error for Unio murchisonianus Lea, 1834)

Unio pictorum var. longirostris Middendorff, 1851
Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3.
Schrenck, 1867. Reise Forsch. Amurlande 2: 696.
(misidentified reference to Unio longirostris Rossmässler, 1836)

Nodularia middendorffiMoskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Bogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio murchinsonianus Küster, 1862
‘Lea’ Küster, 1862. Conchyl. Cab. 9(2): 309.
(error for Unio murchisonianus Lea, 1834)

Unio pictorum Schrenck, 1867
‘L.’ Schrenck, 1867. Reise Forsch. Amurlande 2: 696-699.
(misidentified reference to Mya pictorum Linnaeus, 1758)

Nodularia middendorffiMoskvicheva (1973)

Unio pictorum var. longirostris Westerlund, 1877
Westerlund, 1877. Kongl. Svenska Vet. Akad. Handl. 14(12): 74.
(misidentified reference to Unio longirostris Rossmässler, 1836)

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae amurensisHaas (1969)
Nodularia amurensisVinarski & Kantor (2016)

Unio nipponensis Kobelt, 1879
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3.
(misidentified reference to Unio nipponensis Martens, 1877)

Nodularia (s.s.) douglasiae var. nipponensisSimpson (1900)
Unio (Nodularia) nipponensisMartens (1905)

Unio douglasiae var. osbeckii Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128i.
(reference to Unio osbeckii Philippi, 1844)

Unio (Nodularia) douglasiae var. sinuolatusMartens (1905)

Unio schanghaiensis Neumayr, 1899
‘Lea’ Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645.
(error for Unio shanghaiensis Lea, 1859)

Unio wrigtii de Gregorio, 1914
‘Lea’ de Gregorio, 1914. Il Nat. Siciliano 2: 40.
(error for Unio wrightii Lea, 1865)

Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938
‘Westerlund’ Zhadin, 1938. Faune USSR Mollusques 4(1): 99.
(error for Unio schrencki Westerlund, 1897)

Unio taiwanensis Habe, 1977
‘Pilsbry’ Habe, 1977. Mollusc. Japan: 114.
(error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905)

Unio firnus Liu et al., 1979
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae firmus Heude, 1885)

Unio douglasiae chlorens Liu et al., 1979
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae chloreus Heude, 1885)

Unio douglasiae cleltae Liu et al., 1979
Liu et al., 1979. Econ. Fauna China: 74.
(error for Unio douglasiae deltae Heude, 1885)

Nodularia doctylina Liu et al., 1980
Liu et al., 1980. Acta Zool. Sinica 26(4): 366.
(error for Unio douglasiae dactylinus Heude, 1885)

Nodularia sachalinensis Bogatov, 2001
Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 59.
(error for Nodularia sakhalinensis Bogatov, 2001)

Unio douglassiae Kantor & Sysoev, 2005
‘Griffith et Pidgeon’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 337.
(error for Unio douglasiae Griffith & Pidgeon, 1833)

Nodularia jourdyi Bolotov et al., 2018
Bolotov et al., 2018. Sci. Repts. 8: 4.
Pfeiffer et al., 2019. MPE 137: 117.
(reference to Unio jourdyi Morlet, 1886)

Nodularia douglasiaeLopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024)

Unio osbreckii Lopes-Lima et al., 2020
‘Philippi’ Lopes-Lima et al., 2020. MPE 146: Appendix 1.
(error for Unio osbeckii Philippi, 1844)

Taxonomic History (since 1900)

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) biwaeSimpson (1900), Haas (1911), Simpson (1914)
Nodularia (s.s.) douglasiae var. nipponensisSimpson (1900)
Nodularia (s.s.) jourdyiSimpson (1900), Simpson (1914)
Simpson (1900), Pilsbry & Bequaert (1927), Haas (1936), Haas (1969), Petit & Coan (2008)
Unio (Nodularia) douglasiaeMartens (1905), Zhadin (1938)
Unio (Nodularia) nipponensisMartens (1905)
Unio (Nodularia) douglasiae var. sinuolatusMartens (1905)
Unio jourdyiDautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Unio (Nodularia) pliculosusMartens (1905)
Unio (Nodularia) verruciferMartens (1905)
Unio jourdyi var. corrugataDautzenberg & Fischer (1905)
Unio jourdyi var. ponderosaDautzenberg & Fischer (1905)
Nodularia verrucosaHaas (1910), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Nodularia (s.s.) douglasiae shanghaiensisHaas (1911)
Nodularia (s.s.) pliculosusHaas (1911)
Nodularia (s.s.) verruciferaHaas (1911), Simpson (1914)
Nodularia (s.s.) denserugataHaas (1911), Simpson (1914)
Nodularia (s.s.) douglasiae crassidensHaas (1911)
Oxynaia jourdyiHaas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018)
Unio wrigtii [sic]de Gregorio (1914)
Nodularia (s.s.) pliculosaSimpson (1914)
Nodularia (s.s.) boettgeriSimpson (1914)
Unio kobeltiSimpson (1914)
Nodularia (s.s.) douglasiae var. taiwanicaSimpson (1914)
Nodularia (s.s.) jourdyi var. corrugataSimpson (1914)
Nodularia (s.s.) jourdyi var. ponderosaSimpson (1914)
Nodularia (s.s.) douglasiae var. crassidensSimpson (1914)
Nodularia douglasiaeHaas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Wu et al. (2024), Hou et al. (2024), Hou et al. (2025), Hou et al. (2025), Dai et al. (2025)
Nodularia (s.s.) douglasiae taiwanicaHaas (1924)
Unio (Nodularia) douglasiae var. schrenki [sic]Zhadin (1938)
Unio pictorum middendorffiHaas (1940)
Unio douglasiae taiwanicusKuroda (1941)
Unio douglasiaeZhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018)
Unio douglasiae var. abbreviatusZhadin (1965)
Unio douglasiae var. schrenckiZhadin (1965)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiae amurensisHaas (1969)
Unio biwaeHaas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae pliculosusHaas (1969)
Unio douglasiae verruciferHaas (1969)
Unio douglasiae taiwanicaHaas (1969)
Unio denserugatusHaas (1969)
Nodularia middendorffiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia amurensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia abbreviataMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia schrenckiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Middendorffinaia (Sujfununio) denserugataMoskvicheva & Starobogatov (1973)
Nodularia flavoviridisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Middendorffinaia (s.s.) arsenieviMoskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia sujfunicaMoskvicheva (1973), Zatravkin (1983)
Nodularia vladivostokensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia douglasiae crassidensDang et al. (1980), Do et al. (2018)
Middendorffinaia (s.s.) arsenjevi [sic]Zatravkin (1983)
Nodularia sp.n.Zatravkin (1983)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) amurensisZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) abbreviataZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) schrenckiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) flavoviridisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) sujfunicaZatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (Amurunio) lebedeviZatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia (Magadaninaia) extremalisMartynov & Chernyshev (1992), Starobogatov et al. (2004)
Unio douglasiae biwaeKondo (1997), Kondo (2008)
Unio sculptusYu et al. (2001)
Nodularia (s.s.) moskvichevaeStarobogatov et al. (2004)
Middendorffinaia (s.s.) ochoticaStarobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Nodularia (s.s.) sakhalinensisStarobogatov et al. (2004)
Nodularia crassidensProzorova et al. (2005)
Middendorffinaia arsenieviKantor & Sysoev (2005), Kantor et al. (2010)
Nodularia lebedeviKantor & Sysoev (2005), Kantor et al. (2010)
Scabies longataHu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Nodularia extremalisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia moskvichevaeKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Middendorffinaia ochoticaKantor & Sysoev (2005), Kantor et al. (2010)
Nodularia sakhalinensisKantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia denserugataGraf & Cummings (2007), Bogan (2010)
Unio crassus mongolicusGraf (2007)
Nodularia biwaeBogan (2010)
Nodularia dorriIUCN (2012)
Middendorffinaia (s.s.) ussuriensisBogatov (2012), Vinarski & Kantor (2016)
Nodularidia douglasiaeHe & Zhuang (2013)
Ptychorhynchus denserugataHe & Zhuang (2013), Zieritz et al. (2018)
Scabies crispataHe & Zhuang (2013), Graf & Cummings (2021)
Acuticosta jianghanensisHe & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiae biwaeSano et al. (2017), Mabuchi & Nishida (2020)
Nodularia jourdyiBolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021)
Nodularia suifunica [sic]Klishko et al. (2018)
Nodularia cf. douglasiaeHuang et al. (2019)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiae Clade CChoi et al. (2020)
Nodularia douglasiae Clade BChoi et al. (2020)
Nodularia douglasiae Clade AChoi et al. (2020)
Ptychorhynchus denserugatumGraf & Cummings (2021)
Nodularia douglasiae douglasiaeBogatov (2022)
Nodularia douglasiae amurensisBogatov (2022)
Nodularia douglasiae middendorffiBogatov (2022)
Nodularia douglasiae schrenckiBogatov (2022)
Middendorffinaia mongolica ussuriensisBogatov (2022)
Middendorffinaia mongolica ochoticaBogatov (2022)
Nodularia sp. 2Dai et al. (2024)

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."