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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Bivalvia | ... | Unionidae | Unioninae | Unionini | Nodularia species Nodularia douglasiae (Griffith & Pidgeon, 1833): synonymy 273 specimens + 225 Genbank vouchers Unio douglasiae Griffith & Pidgeon, 1833
Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’]. ‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998) Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Wu et al. (2024), Hou et al. (2024), Hou et al. (2025), Hou et al. (2025), Dai et al. (2025) Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018) Unio douglasiae douglasiae — Haas (1969) — Petit & Coan (2008) Nodularidia douglasiae — He & Zhuang (2013) Nodularia cf. douglasiae — Huang et al. (2019) Nodularia douglasiae Clade C — Choi et al. (2020) Nodularia douglasiae Clade B — Choi et al. (2020) Nodularia douglasiae Clade A — Choi et al. (2020) Nodularia douglasiae douglasiae — Bogatov (2022) Nodularia sp. 2 — Dai et al. (2024) | |
Unio murchisonianus Lea, 1834
 Lea, 1834. Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6. Type(s): HOLOTYPE (MONOTYPY) USNM_84156, China [“Calcutta, India” penciled out].
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Unio (Nodularia) douglasiae — Martens (1905) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae douglasiae — Haas (1969) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae — Lopes-Lima et al. (2020) | |
Unio osbeckii Philippi, 1844 Philippi, 1844. Z. Malak. 1: 164.
Philippi, 1847. Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Unio (Nodularia) douglasiae — Martens (1905) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae douglasiae — Haas (1969) Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio shanghaiensis Lea, 1859
Lea, 1859. Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121. Type(s): HOLOTYPE BMNH_1965171, Shanghai, China.
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Preston (1912), Simpson (1914) Unio (Nodularia) douglasiae — Martens (1905) Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911) Nodularia douglasiae — Haas (1920), Lopes-Lima et al. (2020) Unio douglasiae douglasiae — Haas (1969) Unio douglasiae — Liu et al. (1979) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio wrightii Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97. Type(s): HOLOTYPE USNM_84129, China.
Nodularia (s.s.) douglasiae — Simpson (1900), Simpson (1914) Unio (Nodularia) douglasiae — Martens (1905) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio wrigtii [sic] — de Gregorio (1914) Unio douglasiae douglasiae — Haas (1969) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae — Lopes-Lima et al. (2020) | |
Unio sculptus Deshayes, 1873
Deshayes, 1873. Nouvelles archives 9: 9, pl. 1, figs. 3-3a. Type(s): SYNTYPES MNHN-IM-2000-1779, Petchily, Chine Centrale.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae douglasiae — Haas (1969) Unio douglasiae — Liu et al. (1979) Unio sculptus — Yu et al. (2001) Nodularia douglasiae — Prozorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio biwae Kobelt, 1879
Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4. Type(s): LECTOTYPE SMF_3650, Biwa-See, Japan.
Nodularia (s.s.) biwae — Simpson (1900), Haas (1911), Simpson (1914) Unio biwae — Haas (1969), Habe (1977), Habe (1991), Slugina (2006) Unio douglasiae biwae — Kondo (1997), Kondo (2008) Nodularia biwae — Bogan (2010) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae biwae — Sano et al. (2017), Mabuchi & Nishida (2020) Nodularia douglasiae — Lopes-Lima et al. (2020), Mabuchi et al. (2021) | |
Unio douglasiae chloreus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Nodularia (s.s.) douglasiae — Simpson (1900) Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio douglasiae dactylinus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128e. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae douglasiae — Haas (1969) Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio douglasiae deltae Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’]. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio favidens var. deltae Benson, 1862)
Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio douglasiae firmus Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)
Nodularia (s.s.) douglasiae — Simpson (1900) Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio douglasiae hainanensis Heude, 1885
Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128f. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Unio douglasiae — Liu et al. (1979) Nodularia douglasiae — Prozorova et al. (2005), Lopes-Lima et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio jourdyi Morlet, 1886
Morlet, 1886. J. de Conch. 34: 76, pl. 13, figs. 5-5a. Type(s): SYNTYPES MNHN_MP_0150, Bac Hat etang de la riviere Claire.
Nodularia (s.s.) jourdyi — Simpson (1900), Simpson (1914) Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905) Oxynaia jourdyi — Haas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018) Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021) Nodularia douglasiae — Wu et al. (2024) | |
Unio pictorum var. amurensis Mousson, 1887 Mousson, 1887. J. de Conch. 35: 26, pl. 1, fig. 8. (intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae amurensis — Haas (1969) Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae amurensis — Bogatov (2022) | |
Unio middendorffi Westerlund, 1890 Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris]. Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 113. Type(s): NEOTYPE BIV229_5 (by Bolotov et al., 2020: Table S1), Gladkaya River, Primorye Region, Russia.
Unio pictorum middendorffi — Haas (1940) Unio douglasiae amurensis — Haas (1969) Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae middendorffi — Bogatov (2022) | |
Unio pliculosus Martens, 1894
Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b. Type(s): TYPE ZMB_55620, Singei, Korea.
— Simpson (1900) Unio (Nodularia) pliculosus — Martens (1905) Nodularia (s.s.) pliculosus — Haas (1911) Nodularia (s.s.) pliculosa — Simpson (1914) Unio douglasiae pliculosus — Haas (1969) Unio douglasiae — Habe (1977) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae — Lopes-Lima et al. (2020) | |
Unio verrucifer Martens, 1894
Martens, 1894. S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905. Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b. Type(s): TYPE ZMB_55622, Hangang.
— Simpson (1900) Unio (Nodularia) verrucifer — Martens (1905) Nodularia (s.s.) verrucifera — Haas (1911), Simpson (1914) Unio douglasiae verrucifer — Haas (1969) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae — Lopes-Lima et al. (2020) | |
Unio abbreviatus Westerlund, 1897
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 132. (homonym of Unio requienii var. abbreviatus de Joannis, 1859) Type(s): PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938) Unio douglasiae var. abbreviatus — Zhadin (1965) Unio douglasiae amurensis — Haas (1969) Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularia (s.s.) schrencki — Bogatov (2012) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae schrencki — Bogatov (2022) | |
Unio schrencki Westerlund, 1897
Westerlund, 1897. Ann. Mus. St. Pétersbourg 1897: 131. Type(s): PARATYPE SMF_3663, Amur - Fluss.
Nodularia (s.s.) douglasiae — Simpson (1900), Simpson (1914) Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938) Unio douglasiae var. schrencki — Zhadin (1965) Unio douglasiae amurensis — Haas (1969) Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017) Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae schrencki — Bogatov (2022) | |
Unio boettgeri Neumayr, 1899
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’]. (homonym of Unio firmus var. boettgeri Ihering, 1893)
— Simpson (1900), Haas (1969) Nodularia (s.s.) boettgeri — Simpson (1914) Nodularia douglasiae — Haas (1920) Nodularia (s.s.) douglasiae — Haas (1924) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio kobelti Neumayr, 1899
Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1. (homonym of Unio kobelti Kobelt & Rolle, 1895)
Unio kobelti — Simpson (1914) Nodularia douglasiae — Haas (1920) Nodularia (s.s.) douglasiae — Haas (1924) Unio douglasiae — Liu et al. (1979) Nodularidia douglasiae — He & Zhuang (2013) | |
Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905
Pilsbry & Hirase, 1905. Proc. Acad. Nat. Sci. 57: 750. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833) Type(s): HOLOTYPE ANSP_58170, Formosa.
Nodularia (s.s.) douglasiae var. taiwanica — Simpson (1914) Nodularia (s.s.) douglasiae taiwanica — Haas (1924) Unio douglasiae taiwanicus — Kuroda (1941) Unio douglasiae taiwanica — Haas (1969) Unio douglasiae — Habe (1977), Liu et al. (1979), Wu (1982) Nodularidia douglasiae — He & Zhuang (2013) | |
Unio jourdyi var. corrugata Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905. J. de Conch. 53: 205. (homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886) Type(s): SYNTYPE MNHN_MP_0149, Bas-Luc Nam.
Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905) Nodularia (s.s.) jourdyi var. corrugata — Simpson (1914) | |
Unio jourdyi var. ponderosa Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905. J. de Conch. 53: 453. (homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886) Type(s): SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.
Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905) Nodularia (s.s.) jourdyi var. ponderosa — Simpson (1914) | |
Nodularia denserugata Haas, 1910 Haas, 1910. Ann. Mag. Nat. Hist. 6: 496. Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.
Nodularia (s.s.) denserugata — Haas (1911), Simpson (1914) Unio denserugatus — Haas (1969) Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973) Nodularia denserugata — Graf & Cummings (2007), Bogan (2010) Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018) Ptychorhynchus denserugatum — Graf & Cummings (2021) Nodularia douglasiae — Wu et al. (2024) | |
Nodularia verrucosa Haas, 1910
Haas, 1910. Nachr. Bl. Dtsch. Malak. Ges. 42: 99. Type(s): TYPE SMF_3660, Hunan, M. China.
Nodularia verrucosa — Haas (1910), Simpson (1914) — Pilsbry & Bequaert (1927), Haas (1936), Haas (1969) Nodularidia douglasiae — He & Zhuang (2013) | |
Nodularia douglasiae crassidens Haas, 1910
Haas, 1910. Ann. Mag. Nat. Hist. 6: 499. Haas, 1910. Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911. Conch. Cab. 9 (pt. 2, h. 44): 75. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833) Type(s): TYPE SMF_3659, Hunan, M. China.
Nodularia (s.s.) douglasiae crassidens — Haas (1911) Nodularia (s.s.) douglasiae var. crassidens — Simpson (1914) Unio douglasiae douglasiae — Haas (1969) Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018) Nodularia crassidens — Prozorova et al. (2005) Nodularia dorri — IUCN (2012) Nodularidia douglasiae — He & Zhuang (2013) | |
Nodularia douglasiae flavoviridis Haas, 1910
Haas, 1910. Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2. (intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833) Type(s): TYPE SMF_3654, W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.
Unio douglasiae amurensis — Haas (1969) Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularia (s.s.) schrencki — Bogatov (2012) Nodularidia douglasiae — He & Zhuang (2013) | |
Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973 Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1. ‘Moskv. et Star.’ Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 41. (incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973) | |
Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973 Moskvicheva & Starobogatov, 1973. Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п. (correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)
Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983) Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010) Unio crassus mongolicus — Graf (2007) Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016) Middendorffinaia mongolica — Klishko et al. (2019) Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020) Middendorffinaia mongolica ussuriensis — Bogatov (2022) | |
Nodularia sujfunica Moskvicheva, 1973 Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 с-т.
Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983) Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987) Nodularia (s.s.) vladivostokensis — Starobogatov et al. (2004) Nodularia vladivostokensis — Kantor & Sysoev (2005) Nodularidia douglasiae — He & Zhuang (2013) Nodularia suifunica [sic] — Klishko et al. (2018) Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularia douglasiae schrencki — Bogatov (2022) | |
Nodularia vladivostokensis Moskvicheva, 1973 Moskvicheva, 1973. Zool. Zhur. 52(10): 1463, figs. 1 п-р.
Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018) Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae douglasiae — Bogatov (2022) | |
Nodularia lebedevi Zatravkin & Starobogatov, 1984 Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1787, figs. 5-8.
Nodularia sp.n. — Zatravkin (1983) Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018) Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularia (s.s.) middendorffi — Bogatov (2012) Nodularidia douglasiae — He & Zhuang (2013) Nodularia middendorffi — Vinarski & Kantor (2016) Nodularia douglasiae middendorffi — Bogatov (2022) | |
Scabies longata Liu et al., 1991 Liu et al., 1991. Acta Zootax. Sinica 16(3): 279 [281], fig’d.
Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017) Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021) Nodularia douglasiae — Dai et al. (2024) | |
Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992 Martynov & Chernyshev, 1992. Zool. Zhur. 71: 19, figs. 1-3.
Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004) Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae schrencki — Bogatov (2022) | |
Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992 Bogatov & Starobogatov, 1992. Zool. Zhur. 71: 134, fig’d.
Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004) Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia douglasiae — Graf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularia douglasiae middendorffi — Bogatov (2022) | |
Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000 Bogatov, 2000. Zool. Zhur. 79: 861, text fig. a-r.
Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017) Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010) Unio crassus mongolicus — Graf (2007) Middendorffinaia mongolica — Klishko et al. (2019) Nodularia douglasiae — Lopes-Lima et al. (2020), Bolotov et al. (2020) Middendorffinaia mongolica ochotica — Bogatov (2022) | |
Nodularia sakhalinensis Bogatov, 2001 Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 75.
Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004) Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018) Nodularia douglasiae — Graf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae schrencki — Bogatov (2022) | |
Acuticosta jianghanensis He & Zhuang, 2013 He & Zhuang, 2013. Freshwater Bivalves China: 23, fig. 47-48.
Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020) Nodularia douglasiae — Bogatov & Prozorova (2017) | |
Nodularia douglasiae nipponensis Sano et al., 2017 ‘Martens’ of Authors. Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part] (misidentified reference to Unio nipponensis Martens, 1877)
Nodularia douglasiae — Sano et al. (2020), Choi et al. (2020) | |
Nodularia suifunica Klishko et al., 2018 ‘Moskvicheva’ of Authors. Klishko et al., 2018. Systemat. Biodiv. 16(3): 288. Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 59. (error for Nodularia sujfunica Moskvicheva, 1973) | |
Unio ezonis of Authors, unknown year ‘Kuroda, MS’ in ANSP coll. of Authors. | |
Unio föttgeri of Authors, unknown year of Authors.
Unio douglasiae — Liu et al. (1979) | |
Margarita (Unio) merchisonianus Lea, 1836 Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata]. (error for Unio murchisonianus Lea, 1834) | |
Unio pictorum var. longirostris Middendorff, 1851 Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3. Schrenck, 1867. Reise Forsch. Amurlande 2: 696. (misidentified reference to Unio longirostris Rossmässler, 1836)
Nodularia middendorffi — Moskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Bogatov (2012) Nodularidia douglasiae — He & Zhuang (2013) Nodularia douglasiae — Lopes-Lima et al. (2020) | |
Unio murchinsonianus Küster, 1862 ‘Lea’ Küster, 1862. Conchyl. Cab. 9(2): 309. (error for Unio murchisonianus Lea, 1834) | |
Unio pictorum Schrenck, 1867 ‘L.’ Schrenck, 1867. Reise Forsch. Amurlande 2: 696-699. (misidentified reference to Mya pictorum Linnaeus, 1758)
Nodularia middendorffi — Moskvicheva (1973) | |
Unio pictorum var. longirostris Westerlund, 1877 Westerlund, 1877. Kongl. Svenska Vet. Akad. Handl. 14(12): 74. (misidentified reference to Unio longirostris Rossmässler, 1836)
Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Unio douglasiae amurensis — Haas (1969) Nodularia amurensis — Vinarski & Kantor (2016) | |
Unio nipponensis Kobelt, 1879 Kobelt, 1879. Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3. (misidentified reference to Unio nipponensis Martens, 1877)
Nodularia (s.s.) douglasiae var. nipponensis — Simpson (1900) Unio (Nodularia) nipponensis — Martens (1905) | |
Unio douglasiae var. osbeckii Heude, 1885 Heude, 1885. Conch. Fluv. Nanking (9): pl. 65, fig. 128i. (reference to Unio osbeckii Philippi, 1844)
Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905) | |
Unio schanghaiensis Neumayr, 1899 ‘Lea’ Neumayr, 1899. Wiss. Erg. Reise Szechény Ost-Asien 2: 645. (error for Unio shanghaiensis Lea, 1859) | |
Unio wrigtii de Gregorio, 1914 ‘Lea’ de Gregorio, 1914. Il Nat. Siciliano 2: 40. (error for Unio wrightii Lea, 1865) | |
Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938 ‘Westerlund’ Zhadin, 1938. Faune USSR Mollusques 4(1): 99. (error for Unio schrencki Westerlund, 1897) | |
Unio taiwanensis Habe, 1977 ‘Pilsbry’ Habe, 1977. Mollusc. Japan: 114. (error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905) | |
Unio firnus Liu et al., 1979 Liu et al., 1979. Econ. Fauna China: 74. (error for Unio douglasiae firmus Heude, 1885) | |
Unio douglasiae chlorens Liu et al., 1979 Liu et al., 1979. Econ. Fauna China: 74. (error for Unio douglasiae chloreus Heude, 1885) | |
Unio douglasiae cleltae Liu et al., 1979 Liu et al., 1979. Econ. Fauna China: 74. (error for Unio douglasiae deltae Heude, 1885) | |
Nodularia doctylina Liu et al., 1980 Liu et al., 1980. Acta Zool. Sinica 26(4): 366. (error for Unio douglasiae dactylinus Heude, 1885) | |
Nodularia sachalinensis Bogatov, 2001 Bogatov, 2001. Byull. Dal'nev. Malak.Obshch. 5: 59. (error for Nodularia sakhalinensis Bogatov, 2001) | |
Unio douglassiae Kantor & Sysoev, 2005 ‘Griffith et Pidgeon’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 337. (error for Unio douglasiae Griffith & Pidgeon, 1833) | |
Nodularia jourdyi Bolotov et al., 2018 Bolotov et al., 2018. Sci. Repts. 8: 4. Pfeiffer et al., 2019. MPE 137: 117. (reference to Unio jourdyi Morlet, 1886)
Nodularia douglasiae — Lopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024) | |
Unio osbreckii Lopes-Lima et al., 2020 ‘Philippi’ Lopes-Lima et al., 2020. MPE 146: Appendix 1. (error for Unio osbeckii Philippi, 1844) | |
Taxonomic History (since 1900)Nodularia (s.s.) douglasiae — Simpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998) Nodularia (s.s.) douglasiae var. shanghaiensis — Simpson (1900), Preston (1912), Simpson (1914) Nodularia (s.s.) biwae — Simpson (1900), Haas (1911), Simpson (1914) Nodularia (s.s.) douglasiae var. nipponensis — Simpson (1900) Nodularia (s.s.) jourdyi — Simpson (1900), Simpson (1914) — Simpson (1900), Pilsbry & Bequaert (1927), Haas (1936), Haas (1969), Petit & Coan (2008) Unio (Nodularia) douglasiae — Martens (1905), Zhadin (1938) Unio (Nodularia) nipponensis — Martens (1905) Unio (Nodularia) douglasiae var. sinuolatus — Martens (1905) Unio jourdyi — Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905) Unio (Nodularia) pliculosus — Martens (1905) Unio (Nodularia) verrucifer — Martens (1905) Unio jourdyi var. corrugata — Dautzenberg & Fischer (1905) Unio jourdyi var. ponderosa — Dautzenberg & Fischer (1905) Nodularia verrucosa — Haas (1910), Simpson (1914) Nodularia (s.s.) douglasiae douglasiae — Haas (1911) Nodularia (s.s.) douglasiae shanghaiensis — Haas (1911) Nodularia (s.s.) pliculosus — Haas (1911) Nodularia (s.s.) verrucifera — Haas (1911), Simpson (1914) Nodularia (s.s.) denserugata — Haas (1911), Simpson (1914) Nodularia (s.s.) douglasiae crassidens — Haas (1911) Oxynaia jourdyi — Haas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Zieritz et al. (2018), Do et al. (2018) Unio wrigtii [sic] — de Gregorio (1914) Nodularia (s.s.) pliculosa — Simpson (1914) Nodularia (s.s.) boettgeri — Simpson (1914) Unio kobelti — Simpson (1914) Nodularia (s.s.) douglasiae var. taiwanica — Simpson (1914) Nodularia (s.s.) jourdyi var. corrugata — Simpson (1914) Nodularia (s.s.) jourdyi var. ponderosa — Simpson (1914) Nodularia (s.s.) douglasiae var. crassidens — Simpson (1914) Nodularia douglasiae — Haas (1920), Prozorova et al. (2005), Graf (2007), Graf & Cummings (2007), Bogan (2010), ... Wu et al. (2024), Hou et al. (2024), Hou et al. (2025), Hou et al. (2025), Dai et al. (2025) Nodularia (s.s.) douglasiae taiwanica — Haas (1924) Unio (Nodularia) douglasiae var. schrenki [sic] — Zhadin (1938) Unio pictorum middendorffi — Haas (1940) Unio douglasiae taiwanicus — Kuroda (1941) Unio douglasiae — Zhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), Liu et al. (1980), ... Zhou et al. (2017), Araujo et al. (2018), Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018) Unio douglasiae var. abbreviatus — Zhadin (1965) Unio douglasiae var. schrencki — Zhadin (1965) Unio douglasiae douglasiae — Haas (1969) Unio douglasiae amurensis — Haas (1969) Unio biwae — Haas (1969), Habe (1977), Habe (1991), Slugina (2006) Unio douglasiae pliculosus — Haas (1969) Unio douglasiae verrucifer — Haas (1969) Unio douglasiae taiwanica — Haas (1969) Unio denserugatus — Haas (1969) Nodularia middendorffi — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia amurensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia abbreviata — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia schrencki — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017) Middendorffinaia (Sujfununio) denserugata — Moskvicheva & Starobogatov (1973) Nodularia flavoviridis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010) Middendorffinaia (s.s.) arsenievi — Moskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia sujfunica — Moskvicheva (1973), Zatravkin (1983) Nodularia vladivostokensis — Moskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018) Nodularia douglasiae crassidens — Dang et al. (1980), Do et al. (2018) Middendorffinaia (s.s.) arsenjevi [sic] — Zatravkin (1983) Nodularia sp.n. — Zatravkin (1983) Nodularia (s.s.) middendorffi — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia (s.s.) amurensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia (s.s.) abbreviata — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia (s.s.) schrencki — Zatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012) Nodularia (s.s.) flavoviridis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia (s.s.) sujfunica — Zatravkin & Bogatov (1987) Nodularia (s.s.) vladivostokensis — Zatravkin & Bogatov (1987), Starobogatov et al. (2004) Nodularia (Amurunio) lebedevi — Zatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018) Nodularia (Magadaninaia) extremalis — Martynov & Chernyshev (1992), Starobogatov et al. (2004) Unio douglasiae biwae — Kondo (1997), Kondo (2008) Unio sculptus — Yu et al. (2001) Nodularia (s.s.) moskvichevae — Starobogatov et al. (2004) Middendorffinaia (s.s.) ochotica — Starobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017) Nodularia (s.s.) sakhalinensis — Starobogatov et al. (2004) Nodularia crassidens — Prozorova et al. (2005) Middendorffinaia arsenievi — Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia lebedevi — Kantor & Sysoev (2005), Kantor et al. (2010) Scabies longata — Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017) Nodularia extremalis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Nodularia moskvichevae — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016) Middendorffinaia ochotica — Kantor & Sysoev (2005), Kantor et al. (2010) Nodularia sakhalinensis — Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018) Nodularia denserugata — Graf & Cummings (2007), Bogan (2010) Unio crassus mongolicus — Graf (2007) Nodularia biwae — Bogan (2010) Nodularia dorri — IUCN (2012) Middendorffinaia (s.s.) ussuriensis — Bogatov (2012), Vinarski & Kantor (2016) Nodularidia douglasiae — He & Zhuang (2013) Ptychorhynchus denserugata — He & Zhuang (2013), Zieritz et al. (2018) Scabies crispata — He & Zhuang (2013), Graf & Cummings (2021) Acuticosta jianghanensis — He & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020) Nodularia douglasiae biwae — Sano et al. (2017), Mabuchi & Nishida (2020) Nodularia jourdyi — Bolotov et al. (2018), Pfeiffer et al. (2019), Sayenko et al. (2021), Graf & Cummings (2021) Nodularia suifunica [sic] — Klishko et al. (2018) Nodularia cf. douglasiae — Huang et al. (2019) Middendorffinaia mongolica — Klishko et al. (2019) Nodularia douglasiae Clade C — Choi et al. (2020) Nodularia douglasiae Clade B — Choi et al. (2020) Nodularia douglasiae Clade A — Choi et al. (2020) Ptychorhynchus denserugatum — Graf & Cummings (2021) Nodularia douglasiae douglasiae — Bogatov (2022) Nodularia douglasiae amurensis — Bogatov (2022) Nodularia douglasiae middendorffi — Bogatov (2022) Nodularia douglasiae schrencki — Bogatov (2022) Middendorffinaia mongolica ussuriensis — Bogatov (2022) Middendorffinaia mongolica ochotica — Bogatov (2022) Nodularia sp. 2 — Dai et al. (2024)
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