The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina genus Nodularia Conrad, 1853: cladomics
14 Recent valid species (15 total species) Cladomics Summarynumber of species = 14 (15 total)
analyzed in 130 cladograms in 79 publications
maximum number of terminals in an analysis =
11 (mean = 2.72, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 75
maximum number of included terminals = 11, (mean =
3.97 median = 3)
proportion of cladograms that support basic monophyly
= 0.99
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 10 | 12 | 619 | 21 | 6 | 0 | +-- Tchangsinaia
+-----+
| +-- Inversiunio
+-----------------+
| | +----- Pseudocuneopsis
| +--+
| | +-- Diaurora
| +--+
--+ +-- Cuneopsis
|
| +----------------------- Schistodesmus
| |
| | +-------- Nodularia_dorri__2 **
+--+ +-----------+
| | | +----- Nodularia_dorri__1 **
| | +--+
| | | +-- Nodularia_hanensis **
+--+ +--+
| +-- Nodularia_micheloti **
|
| +----------------- Nodularia_breviconcha **
| |
+--+ +-------------- Nodularia_guiensis **
| |
+--+ +----------- Nodularia_nuxpersicae **
| |
+--+ +-- Nodularia_fusiformans **
| +-----+
| | +-- Nodularia_huana **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Hou et al., 2025 [Publ 4401], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 0 | 0 | 0 | 11 | +----- Nodularia_dorri **
+-----------+
| | +-- Nodularia_hanensis **
| +--+
| +-- Nodularia_micheloti **
--+
| +-------------- Nodularia_guiensis **
| |
| | +-- Nodularia_douglasiae **
+--+ +--------+
| | +-- Nodularia_dualobtusa **
| |
+--+ +-- Nodularia_nipponensis **
| +-----+
| | +-- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_huana **
+--+
| +-- Nodularia_breviconcha **
+--+
+-- Nodularia_fusiformans **
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 17 | 14 | 4 | 9 | +-------------------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +-----------------+
| | | +-- Tchangsinaia
| | +--+
| | +-- Cuneopsis
+--+
| +----- Nodularia_dorri **
| +--------------+
| | | +-- Nodularia_hanensis **
| | +--+
+--+ +-- Nodularia_micheloti **
|
| +----------------- Nodularia_breviconcha **
| |
+--+ +-------------- Nodularia_huana **
| |
+--+ +----------- Nodularia_fusiformans **
| |
+--+ +-------- Nodularia_guiensis **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 18 | 15 | 4 | 8 | +----------------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +--------------+
| | | +-- Tchangsinaia
| | +--+
| | +-- Cuneopsis
+--+
| +----- Nodularia_dorri **
| +-----------+
| | | +-- Nodularia_hanensis **
| | +--+
+--+ +-- Nodularia_micheloti **
|
| +-------------- Nodularia_breviconcha **
| |
+--+ +----------- Nodularia_huana **
| |
+--+ +-------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 13 | 2 | 1 | 8 | +-------------------- Schistodesmus
|
| +----- Nodularia_dorri **
--+ +-----------+
| | | +-- Nodularia_micheloti **
| | +--+
+--+ +-- Nodularia_hanensis **
|
| +-------------- Nodularia_breviconcha **
| |
+--+ +----------- Nodularia_huana **
| |
+--+ +-------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nuxpersicae **
+--+
+-- Nodularia_nipponensis **
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 43 | 9 | 4 | 8 | +----------------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +--------------+
| | | +-- Tchangsinaia
| | +--+
| | +-- Cuneopsis
+--+
| +----- Nodularia_dorri **
| +-----------+
| | | +-- Nodularia_hanensis **
| | +--+
+--+ +-- Nodularia_micheloti **
|
| +-------------- Nodularia_breviconcha **
| |
+--+ +----------- Nodularia_huana **
| |
+--+ +-------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 42 | 12 | 4 | 8 | +----------------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +--------------+
| | | +-- Tchangsinaia
| | +--+
| | +-- Cuneopsis
+--+
| +----- Nodularia_dorri **
| +-----------+
| | | +-- Nodularia_micheloti **
| | +--+
+--+ +-- Nodularia_hanensis **
|
| +-------------- Nodularia_breviconcha **
| |
+--+ +----------- Nodularia_huana **
| |
+--+ +-------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_nipponensis **
| +--+
| | +-- Nodularia_nuxpersicae **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Wu et al., 2024 [Publ 4379], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 3 | 3 | 3 | 6 | +-- Cuneopsis
|
+--------------+-- Tchangsinaia
| |
| +-- Pseudocuneopsis
|
--+ +----- Nodularia_dorri **
| +--------+
| | | +-- Nodularia_hanensis **
| | +--+
| | +-- Nodularia_micheloti **
+--+
| +----------- Nodularia_breviconcha **
| |
| | +-- Nodularia_fusiformans **
+--+ +-----+
| | +-- Nodularia_huana **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nuxpersicae **
+--+
+-- Nodularia_nipponensis **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 65 | 14 | 4 | 8 | +----------------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +--------------+
| | | +-- Tchangsinaia
| | +--+
| | +-- Cuneopsis
+--+
| +----- Nodularia_dorri **
| +-----------+
| | | +-- Nodularia_hanensis **
| | +--+
+--+ +-- Nodularia_micheloti **
|
| +-------------- Nodularia_breviconcha **
| |
+--+ +----------- Nodularia_huana **
| |
+--+ +-------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 5 | 4 | 3 | 12 | +----------------- Nodularia_breviconcha **
|
--+ +-------------- Nodularia_huana **
| |
+--+ +----------- Nodularia_fusiformans **
| |
+--+ +-- Nodularia_nipponensis **
| +-----+
| | +-- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_dualobtusa__2 **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa__1 **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 3 | 0 | 0 | 9 | +----------------- Nodularia_breviconcha **
--+
| +-------------- Nodularia_fusiformans **
+--+
| +----------- Nodularia_huana **
+--+
| +-------- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Choi et al., 2020 [Publ 4169], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 3 | 0 | 0 | 9 | +----------- Nodularia_breviconcha **
|
| +-- Nodularia_fusiformans **
--+ +-----+
| | +-- Nodularia_huana **
| |
+--+ +-- Nodularia_dualobtusa **
| +--+
| | +-- Nodularia_douglasiae **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 116 | 12 | 4 | 8 | +----- Schistodesmus
|
+-----------+----- Cuneopsis
| |
| | +-- Tchangsinaia
--+ +--+
| +-- Inversiunio
|
| +-------------- Nodularia_breviconcha **
+--+
| +----------- Nodularia_huana **
+--+
| +-------- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 6 | 16 | 3 | 3 | 11 | +----- Cuneopsis
+--+
| | +-- Inversiunio
| +--+
| +-- Schistodesmus
|
--+ +-- Nodularia_nuxpersicae **
| +--+
| | +-- Nodularia_nipponensis **
| |
| | +-- Nodularia_huana **
+--+--+
| +-- Nodularia_breviconcha **
|
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 16 | 3 | 3 | 12 | +----------- Cuneopsis
|
| +-- Inversiunio
+--------+
| +-- Schistodesmus
--+
| +-- Nodularia_huana **
| +-----+
| | +-- Nodularia_breviconcha **
| |
+--+ +-- Nodularia_nuxpersicae **
| +--+
| | +-- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 7 | 4 | 1 | 8 | +-------------- Nodularia_breviconcha **
--+
| +----------- Nodularia_huana **
+--+
| +-------- Nodularia_nuxpersicae **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 8 | 2 | 2 | 8 | +----------------- Inversiunio
|
--+ +-------------- Cuneopsis
| |
+--+ +----------- Nodularia_breviconcha **
| |
+--+ +-- Nodularia_fusiformans **
| +-----+
| | +-- Nodularia_huana **
+--+
| +----- Nodularia_nuxpersicae **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nipponensis **
Wu et al., 2024 [Publ 4388], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 4 | 0 | 0 | 9 | +-------- Nodularia_breviconcha **
|
--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_dualobtusa **
+--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_nuxpersicae **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 135 | 16 | 4 | 9 | +-- Tchangsinaia
+-----------------+
| +-- Cuneopsis
--+
| +----------------- Pseudocuneopsis
+--+
| +-------------- Schistodesmus
+--+
| +----------- Nodularia_breviconcha **
+--+
| +-------- Nodularia_fusiformans **
+--+
| +----- Nodularia_nuxpersicae **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 36 | 6 | 4 | 11 | +----------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +--------+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +----------- Nodularia_breviconcha **
+--+
| +-------- Nodularia_fusiformans **
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_nuxpersicae **
+--+
+-- Nodularia_nipponensis **
Mabuchi et al., 2021 [Publ 4233], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 1 | 1 | 1 | 12 | +-------- Tchangsinaia
|
--+ +-- Nodularia **
| +--+
| | +-- Nodularia_nipponensis **
+--+
| +-- Nodularia_fusiformans **
+--+
+-- Nodularia_breviconcha **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 83 | 12 | 4 | 10 | +----- Schistodesmus
+--------+
| | +-- Tchangsinaia
| +--+
--+ +-- Cuneopsis
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia_breviconcha **
+--+
| +----- Nodularia_fusiformans **
+--+
| +-- Nodularia_nuxpersicae **
+--+
+-- Nodularia_douglasiae **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 14 | 3 | 2 | 8 | +-- Cuneopsis
+--------+
| +-- Schistodesmus
--+
| +-------- Nodularia_breviconcha **
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_dualobtusa **
+--+
+-- Nodularia_douglasiae **
Choi et al., 2020 [Publ 4169], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 8 | 2 | 1 | 1 | 11 | +----------- Nodularia_breviconcha **
|
--+ +-------- Nodularia_nipponensis__3 **
| |
+--+ +----- Nodularia_douglasiae__3 **
| |
| | +-- Nodularia_dualobtusa **
+--+ |
| +-- Nodularia_nipponensis__1 **
| |
+--+-- Nodularia_nipponensis__2 **
|
+-- Nodularia_douglasiae__1 **
|
+-- Nodularia_douglasiae__2 **
Klishko et al., 2018 [Publ 3945], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 4 | 0 | 0 | 8 | +-------- Nodularia_breviconcha **
--+
| +----- Nodularia_huana **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nipponensis **
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 91 | 16 | 4 | 10 | +-------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +-------- Nodularia_hanensis **
+--+
| +----- Nodularia_fusiformans **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_guiensis **
Wang et al., 2025 [Publ 4488], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 60 | 19 | 4 | 10 | +-------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +-------- Nodularia_hanensis **
+--+
| +----- Nodularia_fusiformans **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_guiensis **
Wu et al., 2026 [Publ 4825], fig. 6B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 61 | 16 | 4 | 10 | +-------------- Pseudocuneopsis
|
| +----- Schistodesmus
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +-------- Nodularia_hanensis **
+--+
| +----- Nodularia_fusiformans **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_guiensis **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 6 | 11 | 3 | 2 | 10 | +-------------- Schistodesmus
|
+-------------- Cuneopsis
--+
| +----------- Nodularia_breviconcha **
| |
+--+ +-------- Nodularia_douglasiae__3 **
| |
+--+ +----- Nodularia_douglasiae__2 **
| |
+--+ +-- Nodularia_douglasiae__1 **
| |
+--+-- Nodularia_nuxpersicae **
|
+-- Nodularia_nipponensis **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 198 | 9 | 3 | 11 | +-------- Cuneopsis_celtiformis
|
+-------- Tchangsinaia
|
+-------- Cuneopsis_rufescens
--+
| +-- Cuneopsis_szechenyii
+-----+
| +-- Cuneopsis_heudei
|
| +----- Nodularia_huana **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nuxpersicae **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 196 | 8 | 3 | 11 | +----- Nodularia_douglasiae **
+--+
| | +-- Nodularia_huana **
| +--+
--+ +-- Nodularia_nuxpersicae **
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 82 | 16 | 5 | 9 | +----------- Pseudocuneopsis
|
| +----- Nodularia_breviconcha **
--+ +--+
| | | +-- Nodularia_douglasiae **
| | +--+
| | +-- Nodularia_nipponensis **
+--+
| +-- Tongchuania
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 44 | 9 | 3 | 11 | +----- Nodularia_huana **
+--+
| | +-- Nodularia_nuxpersicae **
| +--+
--+ +-- Nodularia_douglasiae **
|
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2026 [Publ 4825], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 16 | 2 | 1 | 9 | +-------- Schistodesmus
--+
| +----- Nodularia_fusiformans **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 5 | 4 | 9 | +-------------- Pseudocuneopsis
|
--+ +-- Cuneopsis
| +--------+
| | +-- Schistodesmus
+--+
| +-------- Tchangsinaia
+--+
| +----- Nodularia_nipponensis **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_dualobtusa **
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 43 | 9 | 3 | 11 | +----------------- Schistodesmus
|
| +----- Nodularia_douglasiae **
--+ +--------+
| | | +-- Nodularia_huana **
| | +--+
+--+ +-- Nodularia_nuxpersicae **
|
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 8 | 0 | 0 | 9 | +----- Nodularia_breviconcha **
--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_huana **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 198 | 9 | 3 | 11 | +----------- Schistodesmus
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Nodularia_douglasiae **
+--+
| +-- Nodularia_huana **
+--+
+-- Nodularia_nuxpersicae **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 50 | 4 | 4 | 9 | +----------- Pseudocuneopsis
|
| +----- Nodularia_breviconcha **
--+ +--+
| | | +-- Nodularia_douglasiae **
| | +--+
+--+ +-- Nodularia_nipponensis **
|
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 4 | 30 | 5 | 3 | 10 | +----------- Tchangsinaia
--+
| +-------- Nodularia_nipponensis__1 **
+--+
| +----- Nodularia_breviconcha **
+--+
| +-- Nodularia_nipponensis__2 **
+--+
+-- Nodularia_douglasiae **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 66 | 5 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Nodularia_breviconcha **
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 36 | 13 | 5 | -- | +-------------- Pseudocuneopsis
|
--+ +-- Nodularia_breviconcha **
| +--------+
| | +-- Nodularia_douglasiae **
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Lopes-Lima et al., 2020 [Publ 4080], fig. 12 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 4 | 1 | -- | +----- Inversiunio
--+
| +-- Nodularia_breviconcha **
+--+
+-- Nodularia_douglasiae **
Klishko et al., 2017 [Publ 3946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 0 | 0 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 10 | 0 | 2 | 123 | 4 | 3 | 18 | +-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Scriptolamprotula
|
+-- Cuneopsis_szechenyii
--+
+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae **
|
+-- Nodularia_nuxpersicae **
|
+-- Schistodesmus
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 42 | 11 | 6 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 79 | 4 | 3 | -- | +-- Nodularia_nuxpersicae **
+-----+
| +-- Nodularia_douglasiae **
--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 12 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Nodularia_breviconcha **
| +-----+
| | +-- Nodularia_douglasiae **
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 137 | 4 | 3 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_nuxpersicae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 74 | 4 | 3 | -- | +-- Nodularia_douglasiae **
+--------+
| +-- Nodularia_nuxpersicae **
--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Ouyang et al., 2015 [Publ 4493], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 3 | -- | +-- Tchangsinaia
+-----+
| +-- Cuneopsis_rufescens
|
--+ +-- Cuneopsis_szechenyii
| +--+
| | +-- Cuneopsis_celtiformis
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_huana **
Chen et al., 2025 [Publ 4448], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 7 | 6 | 1 | -- | +----- Pseudocuneopsis
--+
| +-- Nodularia_nipponensis **
+--+
+-- Nodularia_douglasiae **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 7 | 0 | 0 | -- | +-- Nodularia_breviconcha **
--+
+-- Nodularia_douglasiae **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 71 | 5 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Nodularia_breviconcha **
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Froufe et al., 2016 [Publ 3738], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 12 | 0 | 0 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 12 | 6 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 93 | 10 | 4 | -- | +----- Pseudocuneopsis
+--+
| | +-- Nodularia_breviconcha **
| +--+
--+ +-- Nodularia_douglasiae **
|
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 74 | 4 | 3 | -- | +-- Nodularia_douglasiae **
+--------+
| +-- Nodularia_nuxpersicae **
--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 67 | 5 | 3 | -- | +-------------- Inversiunio
|
--+ +-- Schistodesmus
| +--------+
| | +-- Tchangsinaia
+--+
| +-------- Nodularia_nipponensis__1 **
+--+
| +----- Nodularia_nipponensis__2 **
+--+
| +-- Nodularia_nipponensis__3 **
+--+
+-- Nodularia_douglasiae **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 12 | 6 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 71 | 5 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Nodularia_breviconcha **
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Ouyang et al., 2015 [Publ 4493], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 3 | -- | +-- Nodularia_douglasiae **
+--------------+
| +-- Nodularia_huana **
|
--+ +----- Cuneopsis_rufescens__2
| +--------+
| | | +-- Schistodesmus
| | +--+
+--+ +-- Cuneopsis_celtiformis__2
|
| +----------- Tchangsinaia
+--+
| +-------- Cuneopsis_celtiformis__1
+--+
| +----- Cuneopsis_szechenyii__2
+--+
| +-- Cuneopsis_szechenyii__1
+--+
+-- Cuneopsis_rufescens__1
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 11 | 6 | -- | +-- Tchangsinaia
+-----------+
| +-- Inversiunio
--+
| +----------- Pseudocuneopsis
| |
+--+ +-------- Cuneopsis
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_breviconcha **
+--+
| +-- Diaurora
+--+
+-- Schistodesmus
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 42 | 11 | 6 | -- | +-- Nodularia_douglasiae **
--+
+-- Nodularia_breviconcha **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 67 | 5 | 3 | -- | +----------- Inversiunio
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Nodularia_nipponensis__1 **
+--+
| +-- Nodularia_nipponensis__2 **
+--+
+-- Nodularia_douglasiae **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 71 | 10 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Nodularia_breviconcha **
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 32 | 4 | 3 | -- | +----- Cuneopsis
|
--+ +-- Nodularia_nipponensis__1 **
| |
+--+-- Nodularia_douglasiae **
|
+-- Nodularia_nipponensis__2 **
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 11 | 6 | -- | +-- Tchangsinaia
+--------+
| +-- Inversiunio
|
--+ +-- Cuneopsis
| +-----+
| | +-- Pseudocuneopsis
| |
+--+ +-- Nodularia_douglasiae **
| +--+
| | +-- Nodularia_breviconcha **
+--+
| +-- Diaurora
+--+
+-- Schistodesmus
Lopes-Lima et al., 2020 [Publ 4080], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 0 | 0 | -- | +-- Nodularia_breviconcha **
--+
+-- Nodularia_douglasiae **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 10 | 4 | -- | +-------- Pseudocuneopsis
|
| +-- Nodularia_breviconcha **
--+-----+
| +-- Nodularia_douglasiae **
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 93 | 10 | 4 | -- | +-------------- Pseudocuneopsis
|
| +-- Nodularia_breviconcha **
--+ +--------+
| | +-- Nodularia_douglasiae **
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_xinyuensis
+--+
+-- Schistodesmus_spinosus
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 80 | 0 | 0 | -- | +----- Nodularia_douglasiae__1 **
--+
| +-- Nodularia_nuxpersicae **
+--+
+-- Nodularia_douglasiae__2 **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 137 | 4 | 3 | -- | +-- Nodularia_douglasiae **
+-----+
| +-- Nodularia_nuxpersicae **
--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 67 | 5 | 3 | -- | +----- Inversiunio
+-----+
| | +-- Schistodesmus
| +--+
--+ +-- Tchangsinaia
|
| +-------- Nodularia_nipponensis__3 **
+--+
| +----- Nodularia_nipponensis__1 **
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Nodularia_nipponensis__2 **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 10 | 4 | -- | +-------- Pseudocuneopsis
|
| +-- Nodularia_breviconcha **
--+-----+
| +-- Nodularia_douglasiae **
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 2 | -- | +----- Schistodesmus
--+
| +-- Tchangsinaia
+--+
+-- Nodularia_douglasiae **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 2 | 2 | -- | +----- Schistodesmus
--+
| +-- Nodularia **
+--+
+-- Tchangsinaia
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | +-- Nodularia_douglasiae **
--+
+-- Tchangsinaia
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 2 | 2 | -- | +----- Schistodesmus
--+
| +-- Tchangsinaia
+--+
+-- Nodularia_douglasiae **
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 3 | 1 | -- | +-- Nodularia_nipponensis **
--+
+-- Inversiunio
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Tchangsinaia
--+
+-- Nodularia_douglasiae **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- | -- Nodularia_douglasiae **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 5 | 4 | -- | +-- Cuneopsis
+-----+
| +-- Schistodesmus
--+
| +----- Pseudocuneopsis
+--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Nodularia_douglasiae **
--+
+-- Tchangsinaia
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- | +-- Nodularia_douglasiae **
--+
+-- Cuneopsis
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | +-- Nodularia_douglasiae **
--+
+-- Tchangsinaia
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 9 | 5 | -- | +----------- Cuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 6 | 3 | -- | +-- Nodularia_douglasiae **
+--+
| +-- Schistodesmus
--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 4 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 3 | 3 | -- | +-------- Nodularia_douglasiae **
--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | -- Nodularia_douglasiae **
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 2 | 1 | -- | +-- Nodularia_douglasiae **
--+
+-- Inversiunio
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 10 | 5 | -- | +----- Cuneopsis
|
| +-- Nodularia_douglasiae **
+--+
--+ +-- Schistodesmus
|
| +-- Tchangsinaia
| |
+--+-- Inversiunio
|
+-- Pseudocuneopsis
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 4 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 0 | 0 | -- | -- Nodularia_douglasiae **
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- | +-- Nodularia_douglasiae **
--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 5 | 3 | -- | +-------- Nodularia_douglasiae **
--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 10 | 5 | -- | +-------------- Nodularia **
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 2 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 5 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Dai et al., 2024 [Publ 4358], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 5 | 2 | -- | +----- Pseudocuneopsis
--+
| +-- Schistodesmus
+--+
+-- Nodularia_douglasiae **
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 3 | 2 | -- | +-- Tchangsinaia
--+
+-- Nodularia_douglasiae **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | +-- Tchangsinaia
--+
+-- Nodularia_douglasiae **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 5 | 3 | -- | +-------- Nodularia_douglasiae **
--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | +-- Tchangsinaia
--+
+-- Nodularia_dualobtusa **
Klymus et al., 2025 [Publ 4814], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 4 | -- | +-- Nodularia_douglasiae **
+-----+
| +-- Pseudocuneopsis
--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Inversiunio
--+
+-- Nodularia_nipponensis **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 4 | 4 | -- | +-- Pseudocuneopsis
+-----+
| +-- Nodularia_douglasiae **
--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 3 | 2 | -- | +----- Schistodesmus
--+
| +-- Nodularia_nipponensis **
+--+
+-- Tchangsinaia
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 8 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | -- Nodularia_douglasiae **
Klymus et al., 2025 [Publ 4814], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 4 | 4 | -- | +-- Pseudocuneopsis
+-----+
| +-- Schistodesmus
--+
| +----- Nodularia_breviconcha **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- | +----- Schistodesmus
--+
| +-- Nodularia_douglasiae **
+--+
+-- Tchangsinaia
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 1 | 1 | -- | +-- Tchangsinaia
--+
+-- Nodularia_douglasiae **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 3 | 2 | -- | +----- Schistodesmus
--+
| +-- Nodularia_nipponensis **
+--+
+-- Tchangsinaia
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | -- Nodularia_douglasiae **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | -- Nodularia_douglasiae **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 3 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 2 | 2 | -- | +----- Nodularia_douglasiae **
--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 8 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | -- Nodularia_douglasiae **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Nodularia_douglasiae **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 10 | 5 | -- | +----------- Cuneopsis
|
--+ +-- Nodularia_douglasiae **
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 6 | 4 | -- | +-------------- Tchangsinaia
--+
| +----------- Cuneopsis_heudei
+--+
| +-------- Nodularia_douglasiae **
+--+
| +----- Pseudocuneopsis
+--+
| +-- Cuneopsis_rufescens
+--+
+-- Schistodesmus
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | +-- Nodularia_douglasiae **
--+
+-- Tchangsinaia
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | -- Nodularia_douglasiae **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- | +-- Tchangsinaia
--+
+-- Nodularia_douglasiae **
|
