The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Lamprotulini genus Potomida Swainson, 1840: cladomics 3 Recent valid species (13 total species) Cladomics Summarynumber of species = 3 (13 total) analyzed in 134 cladograms in 82 publications maximum number of terminals in an analysis =
3 (mean = 1.1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 9 maximum number of included terminals = 3, (mean =
2.44 median = 2) proportion of cladograms that support basic monophyly
= 1
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 628 | 5 | 2 | 0 |
+----------- Lamprotula
--+
| +-------- Pronodularia
+--+
| +----- Potomida_littoralis **
+--+
| +-- Potomida_acarnanica **
+--+
+-- Potomida_semirugata **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 138 | 3 | 1 | 2 |
+-------- Lamprotula
--+
| +----- Potomida_acarnanica **
+--+
| +-- Potomida_littoralis **
+--+
+-- Potomida_semirugata **
Froufe et al., 2016 [Publ 3727], fig. 3_4 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 2 | 2 | 0 |
+----- Potomida_littoralis **
--+
| +-- Potomida_acarnanica **
+--+
+-- Potomida_semirugata **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 22 | 0 | 0 | 0 |
+----- Potomida_littoralis **
--+
| +-- Potomida_semirugata **
+--+
+-- Potomida_acarnanica **
Whelan et al., 2011 [Publ 3380], fig. 1-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 1 | 1 | -- |
+----- Pronodularia
--+
| +-- Potomida_littoralis **
+--+
+-- Potomida_semirugata **
Wu et al., 2026 [Publ 4811], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 93 | 4 | 2 | -- |
+-- Pronodularia
+-----+
| +-- Lamprotula
--+
| +----- Potomida_semirugata__2 **
+--+
| +-- Potomida_acarnanica **
+--+
+-- Potomida_semirugata__1 **
Whelan et al., 2011 [Publ 3380], fig. 1-MP | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 1 | 1 | -- |
+----- Pronodularia
--+
| +-- Potomida_littoralis **
+--+
+-- Potomida_semirugata **
Matos et al., 2024 [Publ 4383], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 3 | 2 | -- |
+-- Potomida_littoralis **
+--+
| +-- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Matos et al., 2024 [Publ 4384], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 3 | 2 | -- |
+-- Potomida_acarnanica **
+--+
| +-- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Jeratthitikul et al., 2021 [Publ 4219], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 2 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Pronodularia
--+
+-- Potomida_littoralis **
Konopleva et al., 2023 [Publ 4262], fig. 8 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 81 | 1 | 1 | -- |
+-- Lamprotula
--+
+-- Potomida_semirugata **
Zieritz et al., 2021 [Publ 4218], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2020 [Publ 4121], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Froufe et al., 2019 [Publ 3936], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Nagel et al., 1998 [Publ 1946], fig. 9 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
Dai et al., 2025 [Publ 4441], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 3 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Wu et al., 2018 [Publ 3836], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Dai et al., 2024 [Publ 4389], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Pronodularia
+--+
+-- Potomida_semirugata **
Sano et al., 2020 [Publ 4138], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Wu et al., 2021 [Publ 4215], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Froufe et al., 2015 [Publ 3664], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Sonowal et al., 2021 [Publ 4221], fig. 3a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- |
+-- Lamprotula
--+
+-- Potomida_littoralis **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 3 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Guerra et al., 2019 [Publ 4168], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- |
-- Potomida_littoralis **
Bolotov et al., 2018 [Publ 3826], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 81 | 2 | 2 | -- |
-- Potomida_littoralis **
Araujo et al., 2009 [Publ 2589], fig. 7C | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2021 [Publ 4366], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Zhang et al., 2024 [Publ 4397], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2020 [Publ 4121], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Wu et al., 2024 [Publ 4350], fig. 8b_S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Wu et al., 2019 [Publ 3864], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Huang et al., 2019 [Publ 3843], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 4 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Bolotov et al., 2017 [Publ 3761], fig. S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Araujo et al., 2009 [Publ 2589], fig. 7B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2022 [Publ 4238], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Wu et al., 2020 [Publ 4121], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Nagel et al., 1998 [Publ 1946], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- |
-- Potomida_littoralis **
Sano et al., 2020 [Publ 4138], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Pronodularia
+--+
+-- Potomida_littoralis **
Reis et al., 2013 [Publ 3580], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Potomida_littoralis **
Bolotov et al., 2017 [Publ 3764], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 123 | 3 | 2 | -- |
-- Potomida_littoralis **
Sonowal et al., 2021 [Publ 4221], fig. 3b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- |
-- Potomida_littoralis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Froufe et al., 2019 [Publ 3936], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Machordom et al., 2003 [Publ 2688], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- |
-- Potomida_littoralis **
Huang et al., 2019 [Publ 3843], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Huang et al., 2019 [Publ 3843], fig. S3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 4 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Lopes-Lima et al., 2018 [Publ 3829], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- |
-- Potomida_littoralis **
Chen et al., 2025 [Publ 4663], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 3 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 4 | 2 | -- |
+----- Pronodularia
--+
| +-- Potomida_littoralis **
+--+
+-- Lamprotula
Bolotov et al., 2020 [Publ 4084], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 2 | -- |
-- Potomida_littoralis **
Guerra et al., 2019 [Publ 4168], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- |
+----- Pronodularia
--+
| +-- Lamprotula
+--+
+-- Potomida_littoralis **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Pronodularia
+--+
+-- Potomida_littoralis **
Liu et al., 2024 [Publ 4380], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Bolotov et al., 2020 [Publ 4100], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 110 | 4 | 2 | -- |
+-- Lamprotula_leaii
|
+-- Lamprotula
|
--+-- Lamprotula_caveata
|
+-- Potomida_semirugata **
|
+-- Pronodularia
Campbell et al., 2005 [Publ 2660], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Bolotov et al., 2022 [Publ 4250], fig. 8 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 1 | 1 | -- |
-- Potomida_semirugata **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Dai et al., 2023 [Publ 4285], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 3 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Zieritz et al., 2024 [Publ 4381], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 1 | 1 | -- |
-- Potomida_littoralis **
Wu et al., 2021 [Publ 4366], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- |
+----- Pronodularia
--+
| +-- Potomida_littoralis **
+--+
+-- Lamprotula
Burzynski et al., 2017 [Publ 3774], fig. 1-top | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Cha et al., 2018 [Publ 3916], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Froufe et al., 2017 [Publ 3802], fig. 7 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
-- Potomida_littoralis **
Machordom et al., 2003 [Publ 2688], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
-- Potomida_littoralis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 1 | 1 | -- |
+-- Lamprotula
--+
+-- Potomida_littoralis **
Aunins et al., 2018 [Publ 3863], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 1 | 1 | -- |
+-- Lamprotula
--+
+-- Potomida_littoralis **
Araujo et al., 2016 [Publ 3746], fig. 1_2b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- |
-- Potomida_littoralis **
Wang et al., 2018 [Publ 3865], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Jeratthitikul et al., 2021 [Publ 4219], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Araujo et al., 2017 [Publ 3801], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Araujo et al., 2009 [Publ 2589], fig. 7A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Potomida_littoralis **
Dai et al., 2023 [Publ 4285], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 3 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Fonseca et al., 2016 [Publ 3775], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Guerra et al., 2019 [Publ 4168], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Huang et al., 2019 [Publ 3843], fig. S4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 4 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Huang et al., 2018 [Publ 3828], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Froufe et al., 2015 [Publ 3664], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Wu et al., 2021 [Publ 4366], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_semirugata **
+--+
+-- Pronodularia
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Froufe et al., 2019 [Publ 3936], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Huang et al., 2018 [Publ 3828], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
-- Potomida_littoralis **
Khalloufi et al., 2011 [Publ 2748], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Potomida_littoralis **
Välilä et al., 2017 [Publ 3793], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Sano et al., 2020 [Publ 4138], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Pronodularia
+--+
+-- Potomida_littoralis **
Froufe et al., 2019 [Publ 3936], fig. 3-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Wu et al., 2024 [Publ 4350], fig. 5a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Dai et al., 2023 [Publ 4285], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Dai et al., 2024 [Publ 4386], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Jeratthitikul et al., 2024 [Publ 4387], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 58 | 2 | 2 | -- |
+----- Pronodularia
--+
| +-- Lamprotula
+--+
+-- Potomida_semirugata **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 4 | 2 | -- |
+----- Pronodularia
--+
| +-- Potomida_littoralis **
+--+
+-- Lamprotula
Wu et al., 2024 [Publ 4388], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 4 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Lopes-Lima et al., 2018 [Publ 3829], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- |
-- Potomida_littoralis **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Sonowal et al., 2021 [Publ 4221], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- |
-- Potomida_littoralis **
Chen et al., 2025 [Publ 4455], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- |
-- Potomida_littoralis **
Wu et al., 2021 [Publ 4366], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Wu et al., 2024 [Publ 4350], fig. 5c | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
-- Potomida_littoralis **
Dai et al., 2025 [Publ 4440], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Bolotov et al., 2020 [Publ 4100], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 116 | 4 | 2 | -- |
-- Potomida_semirugata **
Guerra et al., 2019 [Publ 4168], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- |
+----- Pronodularia
--+
| +-- Lamprotula
+--+
+-- Potomida_littoralis **
Wu et al., 2024 [Publ 4379], fig. 4_6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Jeratthitikul et al., 2025 [Publ 4423], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 49 | 2 | 2 | -- |
+----- Potomida_semirugata **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
-- Potomida_littoralis **
Campbell et al., 2005 [Publ 2660], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Pronodularia
Bolotov et al., 2017 [Publ 3761], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- |
+----- Lamprotula
--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia
Wu et al., 2020 [Publ 4147], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 3 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Chen et al., 2025 [Publ 4448], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 85 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Dai et al., 2024 [Publ 4358], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Wu et al., 2024 [Publ 4350], fig. 6-7 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Wu et al., 2024 [Publ 4350], fig. 5b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
-- Potomida_littoralis **
Wang et al., 2018 [Publ 3846], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Prié & Puillandre, 2014 [Publ 3529], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
-- Potomida_littoralis **
Kongim et al., 2023 [Publ 4290], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 2 | 2 | -- |
+----- Lamprotula
--+
| +-- Pronodularia
+--+
+-- Potomida_semirugata **
Huang et al., 2019 [Publ 3843], fig. 3-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Reis et al., 2013 [Publ 3580], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
-- Potomida_littoralis **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Huang et al., 2018 [Publ 3828], fig. 6-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Wu et al., 2021 [Publ 4366], fig. 2_I | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Pronodularia
+--+
+-- Lamprotula
Fonseca et al., 2016 [Publ 3775], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- |
+----- Potomida_littoralis **
--+
| +-- Lamprotula
+--+
+-- Pronodularia
Wu et al., 2024 [Publ 4388], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 2 | 1 | -- |
+-- Potomida_littoralis **
--+
+-- Lamprotula
Konopleva et al., 2021 [Publ 4213], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 1 | 1 | -- |
-- Potomida_semirugata **
|