MUSSELpdb | valid gen. Lanceolaria cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Lanceolaria Conrad, 1853: cladomics
14 Recent valid species (16 total species)

Cladomics Summary

number of species = 14 (16 total)
analyzed in 109 cladograms in 65 publications
maximum number of terminals in an analysis = 10 (mean = 2.55, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 80
maximum number of included terminals = 10, (mean = 3.11 median = 3)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Wu et al., 2026 [Publ 4825], fig. 3

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
10	10	8	10	2	0	0	0

  +----------------- Lanceolaria_oxyrhyncha **   
  |                                           
  |              +-- Lanceolaria_gladiola **     
--+  +-----------+                            
  |  |           +-- Lanceolaria_bilirata **     
  |  |                                        
  +--+        +----- Lanceolaria_acrorrhyncha ** 
     |  +-----+                               
     |  |     |  +-- Lanceolaria_grayii **       
     |  |     +--+                            
     +--+        +-- Lanceolaria_danzhouensis ** 
        |                                     
        |  +-------- Lanceolaria_kihirai **      
        +--+                                  
           |  +----- Lanceolaria_triformis **    
           +--+                               
              |  +-- Lanceolaria_lanceolata **   
              +--+                            
                 +-- Lanceolaria_helianthoides **

Wu et al., 2026 [Publ 4825], fig. 5

tT	bi	riS	oT	roT	roS	iD
10	10	8	10	9	0	0	2

  +-------------------- Lanceolaria_oxyrhyncha **   
  |                                              
  |                 +-- Lanceolaria_bilirata **     
--+  +--------------+                            
  |  |              +-- Lanceolaria_gladiola **     
  |  |                                           
  +--+              +-- Lanceolaria_grayii **       
     |  +-----------+                            
     |  |           +-- Lanceolaria_danzhouensis ** 
     +--+                                        
        |  +----------- Lanceolaria_acrorrhyncha ** 
        +--+                                     
           |  +-------- Lanceolaria_kihirai **      
           +--+                                  
              |  +----- Lanceolaria_triformis **    
              +--+                               
                 |  +-- Lanceolaria_helianthoides **
                 +--+                            
                    +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	114	0	0	4

  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_danzhouensis **
     |  +--------+                           
     |  |        +-- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	4	0	0	4

  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_grayii **      
     |  +--------+                           
     |  |        +-- Lanceolaria_danzhouensis **
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	625	0	0	5

  +----------------- Lanceolaria_oxyrhyncha **  
--+                                          
  |  +-------------- Lanceolaria_gladiola **    
  +--+                                       
     |  +----------- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_triformis **   
              +--+                           
                 +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4825], fig. 6B

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	58	0	0	3

           +-- Lanceolaria_bilirata **     
  +--------+                            
  |        +-- Lanceolaria_gladiola **     
  |                                     
--+        +-- Lanceolaria_grayii **       
  |  +-----+                            
  |  |     +-- Lanceolaria_danzhouensis ** 
  +--+                                  
     |  +----- Lanceolaria_triformis **    
     +--+                               
        |  +-- Lanceolaria_helianthoides **
        +--+                            
           +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	41	0	0	5

  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	194	0	0	5

  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	2	5	196	0	0	6

  +-------- Lanceolaria_gladiola **    
  |                                 
--+  +----- Lanceolaria_grayii **      
  |  |                              
  +--+----- Lanceolaria_danzhouensis **
     |                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	2	5	42	0	0	6

  +-------- Lanceolaria_gladiola **    
  |                                 
--+  +----- Lanceolaria_danzhouensis **
  |  |                              
  +--+----- Lanceolaria_grayii **      
     |                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	196	0	0	5

  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	49	0	0	6

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	77	0	0	8

  +-------- Lanceolaria_gladiola **    
--+                                 
  |  +----- Lanceolaria_danzhouensis **
  +--+                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	50	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	70	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	22	5	4	31	0	0	37

  +----- Lanceolaria_oxyrhyncha__1 **
  |                               
  +----- Lanceolaria_oxyrhyncha__2 **
  |                               
  +----- Cristaria_plicata__1     
  |                               
  +----- Cristaria_plicata__2     
  |                               
  +----- Sinanodonta_lauta        
  |                               
  +----- Beringiana_japonica      
  |                               
  +----- Beringiana_fukuharai__1  
  |                               
  +----- Beringiana_beringiana    
  |                               
  +----- Beringiana_fukuharai__2  
  |                               
  +----- Beringiana_fukuharai__3  
  |                               
  +----- Buldowskia_iwakawai__1   
  |                               
  +----- Buldowskia_iwakawai__2   
  |                               
  +----- Sinanodonta_tumens__1    
--+                               
  +----- Sinanodonta_tumens__2    
  |                               
  +----- Pletholophus             
  |                               
  +----- Anemina                  
  |                               
  +----- Lanceolaria_lanceolata **   
  |                               
  +----- Cristaria_plicata__3     
  |                               
  +----- Lanceolaria_gladiola **     
  |                               
  +----- Sinanodonta_angula       
  |                               
  +----- Lepidodesma              
  |                               
  |  +-- Scriptolamprotula        
  +--+                            
  |  +-- Acuticosta               
  |                               
  |  +-- Cuneopsis                
  +--+                            
  |  +-- Nodularia                
  |                               
  |  +-- Schistodesmus            
  +--+                            
     +-- Inversiunio

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	92	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	82	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	70	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	70	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	66	0	0	7

  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	92	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	66	0	0	7

  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	0	0	7

  +----- Lanceolaria_gladiola **    
--+                              
  |  +-- Lanceolaria_grayii **      
  +--+                           
     +-- Lanceolaria_danzhouensis **

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	21	19	3	21	0	0	25

                          +-- Lanceolaria_lanceolata **
  +-----------------------+                         
  |                       +-- Lanceolaria_grayii **    
  |                                                 
--+  +----------------------- Lanceolaria_gladiola **  
  |  |                                              
  |  |              +-------- Pseudanodonta         
  +--+  +-----------+                               
     |  |           |  +----- Lasmigona             
     |  |           +--+                            
     |  |              |  +-- Strophitus            
     +--+              +--+                         
        |                 +-- Pyganodon             
        |                                           
        |  +----------------- Unio                  
        |  |                                        
        +--+  +-------------- Scriptolamprotula     
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              +--+        +-- Nodularia             
                 |  +-----+                         
                 |  |     +-- Schistodesmus         
                 +--+                               
                    |  +----- Tchangsinaia          
                    +--+                            
                       |  +-- Inversiunio           
                       +--+                         
                          +-- Pseudocuneopsis

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	72	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	66	0	0	7

  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	0	0	7

  +----- Lanceolaria_gladiola ** 
--+                           
  |  +-- Lanceolaria_grayii **   
  +--+                        
     +-- Lanceolaria_triformis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	92	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	21	14	3	21	0	0	26

  +-------------- Lanceolaria_grayii **    
  |                                     
  +-------------- Lanceolaria_gladiola **  
  |                                     
  +-------------- Lanceolaria_lanceolata **
  |                                     
--+     +-------- Lasmigona             
  |  +--+                               
  |  |  |  +----- Pseudanodonta         
  |  |  +--+                            
  |  |     |  +-- Pyganodon             
  |  |     +--+                         
  +--+        +-- Strophitus            
     |                                  
     |  +-------- Unio                  
     |  |                               
     |  +-------- Scriptolamprotula     
     |  |                               
     +--+  +----- Cuneopsis             
        |  |                            
        |  |  +-- Nodularia             
        |  +--+                         
        +--+  +-- Schistodesmus         
           |                            
           |  +-- Tchangsinaia          
           |  |                         
           +--+-- Inversiunio           
              |                         
              +-- Pseudocuneopsis

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	65	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	18	0	3	122	0	0	23

  +-- Alasmidonta             
  |                           
  +-- Anodonta_anatina        
  |                           
  +-- Anodonta_cygnea         
  |                           
  +-- Brachyanodon            
  |                           
  +-- Lasmigona               
  |                           
  +-- Pseudanodonta           
  |                           
  +-- Pyganodon               
  |                           
  +-- Simpsonaias             
  |                           
  +-- Strophitus              
--+                           
  +-- Sinanodonta_lauta       
  |                           
  +-- Cristaria_plicata       
  |                           
  +-- Cristaria_radiata       
  |                           
  +-- Pletholophus            
  |                           
  +-- Sinanodonta_lucida      
  |                           
  +-- Sinanodonta_jourdyi     
  |                           
  +-- Lanceolaria_gladiola **    
  |                           
  +-- Lanceolaria_grayii **      
  |                           
  +-- Lanceolaria_danzhouensis **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	20	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Kim et al., 2021 [Publ 4197], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	15	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	35	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	84	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Xiang et al., 2025 [Publ 4433], fig. 7

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	10	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	0	0	7

  +----- Lanceolaria_gladiola **    
--+                              
  |  +-- Lanceolaria_danzhouensis **
  +--+                           
     +-- Lanceolaria_grayii **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	14	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	51	0	0	7

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Ouyang et al., 2015 [Publ 4493], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Ouyang et al., 2011 [Publ 3408], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	4	0	2	11	0	0	10

  +----- Lanceolaria_grayii **  
  |                          
--+----- Acuticosta          
  |                          
  |  +-- Lanceolaria_gladiola **
  +--+                       
     +-- Scriptolamprotula

Wang et al., 2018 [Publ 3846], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	26	0	2	42	0	0	32

  +-------------------------- Lanceolaria_gladiola **  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha **
  |                                                 
--+-------------------------- Cristaria             
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Scriptolamprotula     
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Riccardi et al., 2020 [Publ 4089], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	17	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	27	0	2	39	0	0	33

  +-------------------- Lanceolaria_gladiola **  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha **
  |                                           
  +-------------------- Cristaria             
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Scriptolamprotula     
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	0	0	--

  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	0	0	--

  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Ouyang et al., 2015 [Publ 4493], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	138	0	0	--

  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	19	0	0	--

  +-- Lanceolaria_triformis ** 
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	37	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	12	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	80	0	0	--

  +-- Lanceolaria_gladiola **
--+                       
  +-- Lanceolaria_grayii **

Fonseca et al., 2016 [Publ 3775], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	30	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Xue et al., 2016 [Publ 3659], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4811], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	15	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_lanceolata **

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	38	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Sano et al., 2017 [Publ 3799], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	3	31	0	0	--

  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Wang et al., 2018 [Publ 3865], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2020 [Publ 4084], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	0	0	--

  +-- Lanceolaria_grayii **      
--+                           
  +-- Lanceolaria_danzhouensis **

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2e

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	0	0	--

-- Lanceolaria_lanceolata **

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lanceolaria_grayii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	0	0	--

-- Lanceolaria_grayii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	0	0	--

-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Kano et al., 2019 [Publ 4194], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Lanceolaria_grayii **

Liu et al., 2017 [Publ 3939], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_grayii **

Cha et al., 2018 [Publ 3916], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_gladiola **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Lanceolaria_lanceolata **

Sano et al., 2017 [Publ 3799], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	0	0	--

-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Lanceolaria_lanceolata **

Zhou et al., 2017 [Publ 3788], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	0	0	--

-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Lanceolaria_lanceolata **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lanceolaria_lanceolata **

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