MUSSELpdb | valid gen. Lanceolaria cladomics

	Home \| About \| Database \| Mussel of the Month \| Publications & Presentations


MUSSELpdb
	Introduction Browse d'basing


Taxonomy
	all genera Unionidae Margaritiferidae Hyriidae Etheriidae Mycetopodidae Iridinidae Species & Genus Richness by Taxonomy


Geography
	Summary North America Central America South America Afrotropics North Eurasia East Asia Australasia Species Richness & Endemism by Geography

Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

genus Lanceolaria Conrad, 1853: cladomics
15 Recent valid species (17 total species)

Cladomics Summary

number of species = 15 (17 total)
analyzed in 106 cladograms in 64 publications
maximum number of terminals in an analysis = 8 (mean = 2.27, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 77
maximum number of included terminals = 8, (mean = 2.75 median = 2)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	4	12	42	0	0	0

  +----------- Lanceolaria_gladiola **       
  |                                       
  |  +-------- Lanceolaria_grayii__5 **      
--+  |                                    
  |  |     +-- Lanceolaria_lanceolata **     
  |  +-----+                              
  |  |     +-- Lanceolaria_triformis **      
  +--+                                    
     |  +----- Lanceolaria_grayii__4 **      
     |  |                                 
     |  |  +-- Lanceolaria_eucylindrica__1 **
     |  |  |                              
     +--+  +-- Lanceolaria_eucylindrica__2 **
        |  |                              
        |  +-- Lanceolaria_eucylindrica__3 **
        |  |                              
        +--+-- Lanceolaria_eucylindrica__4 **
           |                              
           +-- Lanceolaria_grayii__1 **      
           |                              
           +-- Lanceolaria_grayii__2 **      
           |                              
           +-- Lanceolaria_grayii__3 **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

tT	bi	riS	oT	roT	roS	iD
8	8	7	9	624	0	0	8

  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  |  |        +----- Lanceolaria_grayii__2 **   
  +--+  +-----+                              
     |  |     |  +-- Lanceolaria_eucylindrica **
     |  |     +--+                           
     +--+        +-- Lanceolaria_grayii__1 **   
        |                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_triformis **   
              +--+                           
                 +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	114	0	0	9

  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_bilirata **    
     |  +--------+                           
     |  |        +-- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	4	0	0	9

  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_grayii **      
     |  +--------+                           
     |  |        +-- Lanceolaria_bilirata **    
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	41	0	0	8

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	194	0	0	8

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	196	0	0	8

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	49	0	0	8

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	5	196	0	0	8

  +-------- Lanceolaria_gladiola **  
  |                               
--+  +----- Lanceolaria_grayii__1 ** 
  |  |                            
  +--+----- Lanceolaria_grayii__2 ** 
     |                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	51	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	66	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	0	0	9

  +----- Lanceolaria_gladiola ** 
--+                           
  |  +-- Lanceolaria_grayii **   
  +--+                        
     +-- Lanceolaria_triformis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	93	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	71	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	71	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	20	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
3	3	2	4	77	0	0	9

  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii__2 ** 
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_grayii__1 **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	51	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	93	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	93	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Xiang et al., 2025 [Publ 4433], fig. 7

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	10	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	89	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	71	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Kim et al., 2021 [Publ 4197], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	15	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	36	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	89	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	14	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	85	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	22	5	4	31	0	0	30

  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Scriptolamprotula      
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
3	21	14	3	21	0	0	22

  +-------------- Lanceolaria_grayii **    
  |                                     
  +-------------- Lanceolaria_gladiola **  
  |                                     
  +-------------- Lanceolaria_lanceolata **
  |                                     
--+     +-------- Lasmigona             
  |  +--+                               
  |  |  |  +----- Pseudanodonta         
  |  |  +--+                            
  |  |     |  +-- Pyganodon             
  |  |     +--+                         
  +--+        +-- Strophitus            
     |                                  
     |  +-------- Unio                  
     |  |                               
     |  +-------- Scriptolamprotula     
     |  |                               
     +--+  +----- Cuneopsis             
        |  |                            
        |  |  +-- Nodularia             
        |  +--+                         
        +--+  +-- Schistodesmus         
           |                            
           |  +-- Tchangsinaia          
           |  |                         
           +--+-- Inversiunio           
              |                         
              +-- Pseudocuneopsis

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	83	0	0	9

  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	21	19	3	21	0	0	21

                          +-- Lanceolaria_lanceolata **
  +-----------------------+                         
  |                       +-- Lanceolaria_grayii **    
  |                                                 
--+  +----------------------- Lanceolaria_gladiola **  
  |  |                                              
  |  |              +-------- Pseudanodonta         
  +--+  +-----------+                               
     |  |           |  +----- Lasmigona             
     |  |           +--+                            
     |  |              |  +-- Strophitus            
     +--+              +--+                         
        |                 +-- Pyganodon             
        |                                           
        |  +----------------- Unio                  
        |  |                                        
        +--+  +-------------- Scriptolamprotula     
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              +--+        +-- Nodularia             
                 |  +-----+                         
                 |  |     +-- Schistodesmus         
                 +--+                               
                    |  +----- Tchangsinaia          
                    +--+                            
                       |  +-- Inversiunio           
                       +--+                         
                          +-- Pseudocuneopsis

Froufe et al., 2019 [Publ 3936], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	136	0	0	--

  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Fonseca et al., 2016 [Publ 3775], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	3	66	0	0	--

  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Froufe et al., 2019 [Publ 3936], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Wang et al., 2018 [Publ 3865], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	12	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Xue et al., 2016 [Publ 3659], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	30	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	136	0	0	--

  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	47	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	27	0	2	43	0	0	30

  +-------------------------- Lanceolaria_gladiola **  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha **
  |                                                 
--+-------------------------- Cristaria             
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Scriptolamprotula     
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	32	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	37	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Sano et al., 2017 [Publ 3799], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	3	31	0	0	--

  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Riccardi et al., 2020 [Publ 4089], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	17	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	17	0	3	122	0	0	19

  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Brachyanodon         
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola ** 
  |                        
  +-- Lanceolaria_grayii__1 **
  |                        
  +-- Lanceolaria_grayii__2 **

Bolotov et al., 2020 [Publ 4084], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	0	0	--

  +-- Lanceolaria_grayii **
--+                     
  +-- Lanceolaria **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	0	0	--

  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	38	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Ouyang et al., 2015 [Publ 4493], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Wang et al., 2018 [Publ 3846], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4811], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	15	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	3	66	0	0	--

  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	27	0	2	39	0	0	30

  +-------------------- Lanceolaria_gladiola **  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha **
  |                                           
  +-------------------- Cristaria             
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Scriptolamprotula     
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	0	0	--

  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	0	0	--

  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Ouyang et al., 2015 [Publ 4493], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	0	0	--

  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Ouyang et al., 2011 [Publ 3408], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	4	0	2	11	0	0	11

  +----- Lanceolaria_grayii **  
  |                          
--+----- Acuticosta          
  |                          
  |  +-- Lanceolaria_gladiola **
  +--+                       
     +-- Scriptolamprotula

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	80	0	0	--

  +-- Lanceolaria_gladiola **
--+                       
  +-- Lanceolaria_grayii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	139	0	0	--

  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	0	0	--

  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	19	0	0	--

  +-- Lanceolaria_triformis ** 
--+                         
  +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	3	66	0	0	--

  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Huang et al., 2019 [Publ 3843], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	0	0	--

-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Liu et al., 2017 [Publ 3939], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Lanceolaria_grayii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	0	0	--

-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Lanceolaria_lanceolata **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_grayii **

Kano et al., 2019 [Publ 4194], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Sano et al., 2017 [Publ 3799], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	20	0	0	--

-- Lanceolaria_grayii **

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lanceolaria_lanceolata **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	0	0	--

-- Lanceolaria_grayii **

Zhou et al., 2017 [Publ 3788], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	0	0	--

-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2e

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_gladiola **

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Lanceolaria_lanceolata **

Cha et al., 2018 [Publ 3916], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Lanceolaria_grayii **

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	0	0	--

-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	0	0	--

-- Lanceolaria_lanceolata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Lanceolaria_lanceolata **

"Making the world a better place, one mollusk at a time."
	The MUSSEL Project — Home Page https://musselpdb.org/. Site developed and maintained by Dan Graf & Kevin Cummings. Funded by The National Science Foundation.