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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio delphinus Spengler, 1793: synonymy
     13 specimens + 82 Genbank vouchers

Unio delphinus Spengler, 1793
Spengler, 1793. Skr. Naturhist. Selsk. 3: 63, no. 28.

Unio delphinusSimpson (1900), Simpson (1914), Araujo et al. (2009), Khalloufi et al. (2011), IUCN (2011), ... Ma et al. (2024), Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025)
Unio pictorum delphinusHaas (1940), Haas (1969)

Unio hispanus Rossmässler, 1844
‘Moq.-Tand. in litt.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 26, pl. 56, fig. 747.

Unio (sec. Lymnium) turtoniSimpson (1900), Simpson (1914)
Unio requieniGermain (1931)
Unio pictorum delphinusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009), Fauna Europaea (2012)
Unio mancus requieniiPrie (2012)

Unio dactylus Morelet, 1845
Morelet, 1845. Moll. Terr. Fluv. Portugal: 110, pl. 14, fig. 2.
(homonym of Unio dactylus Lea, 1840)
Type(s): HOLOTYPE BMNH_93-2-4-1587, Guadiana, near Castro, Portugal.

Unio (sec. Lymnium) pictorumSimpson (1900), Simpson (1914)
Unio pictorum delphinusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009), Fauna Europaea (2012)

Unio mucidus Morelet, 1845
Morelet, 1845. Moll. Terr. Fluv. Portugal: 111, pl. 14, fig. 3.
Type(s): HOLOTYPE BMNH_92-2-4-1974, le Cavado, Portugal.

Unio (sec. Lymnium) pictorumSimpson (1900), Simpson (1914)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusFauna Europaea (2012), Lopes-Lima et al. (2017), Araujo et al. (2018)
Unio pictorumHe & Zhuang (2013)

Unio hyperephanus da Silva e Castro, 1885
da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 289.
Castro in Locard, 1899. Conchyliol. Portug.: 244.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio nevesi da Silva e Castro, 1885
da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 290.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio simoesi da Silva e Castro, 1885
da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 292.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio (hispanus var.?) sevillensis Kobelt, 1887
Kobelt, 1887. An. Soc. Españ. Hist. Nat. 16: 437.
Kobelt, 1888. Icon. Land- und Süssw.-Moll. 3: 56, pl. 89, fig. 496.
(intro'd as intra-sp. of Unio hispanus Rossmässler, 1844)

Unio pictorum delphinusHaas (1940), Haas (1969)

Unio sousanus Locard, 1889
Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3 [as ‘Unio requienii var. acruata’].
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2 [as ‘Unio requienii var. acruatus’].
‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 197, fig. 211 [as ‘souzanus’].
‘Castro’ Locard, 1899. Conchyliol. Portug.: 258.
Type(s): LECTOTYPE MNHN_22545 (by Graf, 2011: 505), le Rhône, à Valence (Drôme).

Unio (Cyrtusiana) souzanus [sic]Germain (1904)
Unio requieniGermain (1931)
Unio elongatulus requieniiHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)
Unio mancus requieniiPrie (2012)

Unio subhispanus Locard, 1889
‘Castro’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 171 [65], 253 [147].
‘Castro’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 200.
‘Castro’ Locard, 1899. Conchyliol. Portug.: 244.
Type(s): LECTOTYPE MNHN_22562 (by Graf, 2011: 507), La Loire, à Saint-Gemmes, près Angers.

Unio (Hispaniana) subhispanusGermain (1904)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)
Unio mancus requieniiPrie (2012)

Unio hispanus var. sphenoides Westerlund, 1892
Westerlund, 1892. An. Soc. Españ. Hist. Nat. 21: 390.
(intro'd as intra-sp. of Unio hispanus Rossmässler, 1844)

Unio pictorum delphinusHaas (1940), Haas (1969)

Unio cameratus Drouët, 1893
Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 183, pl. 2, fig. 8.

Unio cameratusKobelt (1903)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio decurtatus Drouët, 1893
Drouët, 1893. J. de Conch. 41: 170.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 185, pl. 1, fig. 9.

Unio decurtatusKobelt (1903)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio gravatus Drouët, 1893
Drouët, 1893. Unionidae de l’Espagne: 49.

Unio gravatusKobelt (1903)
Unio pictorum delphinusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio limosellus Drouët, 1893
Drouët, 1893. J. de Conch. 41: 169.
Drouët, 1893. Mém. Acad. Sci. Dijon 4: 184, pl. 12, fig. 4.

Unio limosellusKobelt (1903)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio souzanus Locard, 1893
‘Castro’ of Authors.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 197.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 275 [37].
Germain, 1931. Faune de France 22: 737.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 126.
(error for Unio sousanus Locard, 1889)

Unio abrantesianus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 253.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio allenianus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 255.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio barbozanus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 233.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio castroi Locard, 1899
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Bourguignat in Locard, 1899. Conchyliol. Portug.: 234.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio chasmirhynchus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 246.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio chorellinus Locard, 1899
Locard, 1899. Conchyliol. Portug.: 228.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio chorellus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52] [as ‘chlorellus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 227.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio cyrtus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 257.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio euchasmus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 249.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio hypoxanthus Locard, 1899
Casto in Locard, 1899. Conchyliol. Portug.: 226.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio mundanus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 247.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio neothaumus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 259.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio novus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 165 [59]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 258.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio ocresanus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 236.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio oncomensis Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46] [as ‘oncomenus’]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 225.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio paulinoi Locard, 1899
Locard, 1899. Conchyliol. Portug.: 242.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio scalabisianus Locard, 1899
Castro in Locard, 1899. Conchyliol. Portug.: 254.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio schousboei Locard, 1899
Bourguignat in Locard, 1899. Conchyliol. Portug.: 250.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio silvai Locard, 1899
Bourguignat in Locard, 1899. Conchyliol. Portug.: 235.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio submucidus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 160 [54]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 230.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio taganus Locard, 1899
Servain in Locard, 1899. Conchyliol. Portug.: 252.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio tameganus Locard, 1899
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40]. [nomen nudum]
Castro in Locard, 1899. Conchyliol. Portug.: 221.

Unio pictorum mucidusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio requienii taginus Kobelt, 1903
Kobelt, 1903. Icon. Land- und Süssw.-Moll. 10: 28, pl. 280, fig. 1796.
(intro'd as intra-sp. of Unio requienii Michaud, 1831; homonym of Unio littoralis taginus Kobelt, 1903)

Unio requienii taginusKobelt (1903)
Unio pictorum delphinusHaas (1940), Haas (1969)
Unio delphinusAraujo et al. (2009)

Unio subhispanicus Germain, 1931
‘Castro in Locard’ of Authors.
Germain, 1931. Faune de France 22: 737.
Prie, 2012. MalaCo 8: 432.
(error for Unio subhispanus Locard, 1889)

Unio requieniGermain (1931)

Unio hispanicus Graëls, 1846
Graëls, 1846. Catal. Moll. Esp.: 22.
‘Moquin. Rossm.’ Lea, 1852. Synopsis Fam. Naiades: 30.
‘Moquin. Rossm.’ Lea, 1870. Synopsis Fam. Naiades: 48.
(error for Unio hispanus Rossmässler, 1844)

Unio requienii var. arcuata Drouët, 1857
Drouët, 1857. Étud. Naïades. France, Unio: pl. 7, fig. 3.
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 291, pl. 11, fig. 2.
(misidentified reference to Unio arcuata Jacquemin, 1835)

Unio (Cyrtusiana) souzanus [sic]Germain (1904)
Unio requieniGermain (1931)

Unio hispanus Lowe, 1860
Lowe, 1860. unknown publication: 176.
(reference to Unio hispanus Rossmässler, 1844)

Unio elongatulus durieuiDaget (1998)

Unio chlorellus Locard, 1889
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 158 [52].
(error for Unio chorellus Locard, 1899)

Unio oncomenus Locard, 1889
Castro in Locard, 1889. Contr. Faune Malacol. Franc. 13: 152 [46].
(error for Unio oncomensis Locard, 1899)

Taxonomic History (since 1900)

Unio delphinusSimpson (1900), Simpson (1914), Araujo et al. (2009), Khalloufi et al. (2011), IUCN (2011), ... Ma et al. (2024), Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025)
Unio (sec. Lymnium) turtoniSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) pictorumSimpson (1900), Simpson (1914)
Unio cameratusKobelt (1903)
Unio decurtatusKobelt (1903)
Unio gravatusKobelt (1903)
Unio limosellusKobelt (1903)
Unio requienii taginusKobelt (1903)
Unio (Cyrtusiana) souzanus [sic]Germain (1904)
Unio (Hispaniana) subhispanusGermain (1904)
Unio requieniGermain (1931)
Unio pictorum delphinusHaas (1940), Haas (1969)
Unio pictorum mucidusHaas (1940), Haas (1969)
Unio elongatulus requieniiHaas (1940), Haas (1969)
Unio elongatulus durieuiDaget (1998)
Unio mancus requieniiPrie (2012)
Unio pictorumHe & Zhuang (2013)

 
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