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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina genus Pustulosa Frierson, 1927: synonymy
8 Recent valid species (9 total species) Bullata Frierson, 1927
Frierson, 1927. Check List N. Amer. Naiades: 48.
(homonym of Jousseaume, 1875 [Gastr.]; intro'd as sub-gen. of Quadrula Rafinesque, 1820; renamed Pustulosa Frierson, 1927)
Type species: Obliquaria (Quadrula) bullata Rafinesque, 1820 (original designation) = Pustulosa pustulosa Pustulosa Frierson, 1927
Frierson, 1927. Check List N. Amer. Naiades: errata et corrigenda.
Thiele, 1934. Handb. Syst. Weichtierk. 3: 827 [1260].
(intro'd as sub-gen. of Quadrula Rafinesque, 1820; new name for Bullata Frierson, 1927) Taxonomic History (since 1900)Obovaria — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) — Simpson (1900), Simpson (1914)
Unio (s.s.) — Call (1900)
Quadrula (sec. Fusconaia) — Simpson (1900), Simpson (1914)
Quadrula — Ihering (1901), Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ... Inoue et al. (2018), Marshall et al. (2018), Chase et al. (2018), Klymus et al. (2020), Gasparini et al. (2020)
Unio — de Gregorio (1914), Henderson (1935)
Fusconaia — Ortmann (1923), Ortmann (1925), Clench & Turner (1956), Haas (1969), Burch (1975), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Quadrula (Bullata) — Frierson (1927)
Quadrula (Quincuncina) — Frierson (1927)
Quadrula (Fusconia [sic]) — Frierson (1927)
Quadrula (Striata) — Frierson (1927)
Quadrula (s.s.) — F.C. Baker (1928)
Quincuncina — Clench & Turner (1956), Haas (1969), Burch (1975), Davis & Fuller (1981), Davis (1983), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Pustulosa — Modell (1957), Neemuchwala et al. (2023), FMCS (2025), Campbell & Lydeard (2026)
Quadrula (Pustulosa) — Haas (1969)
Pleurobema — Morrison (1969), Vidrine (1993)
Obliquaria — Burch (1975)
Amphinaias — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011), ITIS (2013), Roe & Boyer (2015)
Rotundaria — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias — Williams et al. (2017), Johnson et al. (2018), Burlakova et al. (2018), Pieri et al. (2018), Lopes-Lima et al. (2019), Inoue et al. (2020), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Hauck et al. (2023), Freeman et al. (2025)
Cyclonaias (Quadrula) — Porto-Hannes et al. (2021)
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