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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Lamprotulini

genus Lamprotula Simpson, 1900: cladomics
     17 Recent valid species (26 total species)

Cladomics Summary

number of species = 17 (26 total)
analyzed in 149 cladograms in 93 publications
maximum number of terminals in an analysis = 4 (mean = 1.66, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 74
maximum number of included terminals = 4, (mean = 2.32 median = 2)
proportion of cladograms that support basic monophyly = 0.93

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
332644000
  +-------- Lamprotula_leaii **          
  |                                   
--+  +----- Lamprotula_cornuumlunae__1 **
  |  |                                
  |  +----- Lamprotula_cornuumlunae__3 **
  +--+                                
     +----- Lamprotula_cornuumlunae__4 **
     |                                
     |  +-- Lamprotula_cornuumlunae__2 **
     +--+                             
        +-- Lamprotula_caveata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4435628424
           +-- Pronodularia           
  +--------+                          
  |        +-- Potomida               
  |                                   
--+        +-- Lamprotula_caveata **     
  |  +-----+                          
  |  |     +-- Lamprotula_cornuumlunae **
  +--+                                
     |  +----- Lamprotula_leaii__2 **    
     +--+                             
        |  +-- Lamprotula_leaii__1 **    
        +--+                          
           +-- Lamprotula_seomjinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
3325119224
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_caveata **     
     |  |                                
     +--+  +----- Lamprotula_leaii__2 **    
        |  |                             
        +--+----- Lamprotula_leaii__3 **    
           |                             
           |  +-- Lamprotula_seomjinensis **
           +--+                          
              +-- Lamprotula_leaii__1 **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
331392323
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
331443004
  +----- Lamprotula_leaii **       
  |                             
--+  +-- Lamprotula_caveata__1 **  
  |  |                          
  +--+-- Lamprotula_cornuumlunae **
     |                          
     +-- Lamprotula_caveata__2 **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
331388223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
316031142216
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
  |                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus           
  |                            
  +-- Parvasolenaia            
  |                            
--+-- Lamprotula_leaii **         
  |                            
  +-- Lamprotula **               
  |                            
  +-- Lamprotula_caveata **       
  |                            
  +-- Potomida                 
  |                            
  +-- Pronodularia             
  |                            
  +-- Sinohyriopsis            
  |                            
  +-- Chamberlainia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3324137314
  +----------- Potomida               
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata__2 **  
     +--+                             
        |  +-- Lamprotula_cornuumlunae **
        +--+                          
           +-- Lamprotula_caveata__1 **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
342318004
  +-------- Lamprotula_leaii **       
--+                                
  |  +----- Lamprotula_caveata **     
  +--+                             
     |  +-- Gibbosula              
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
331388223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331315003
  +----- Lamprotula_leaii **       
--+                             
  |  +-- Lamprotula_caveata **     
  +--+                          
     +-- Lamprotula_cornuumlunae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
331392223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
331392223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3313196223
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3313198223
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
331324223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
3503108225
  +-- Lamprotula_leaii **  
  |                     
  +-- Lamprotula **        
  |                     
--+-- Lamprotula_caveata **
  |                     
  +-- Potomida          
  |                     
  +-- Pronodularia

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
332427114
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata__1 **  
     +--+                             
        |  +-- Lamprotula_cornuumlunae **
        +--+                          
           +-- Lamprotula_caveata__2 **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
331311113
  +-------- Potomida               
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
332528223
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_leaii **       
     |  |                                
     +--+  +----- Lamprotula_caveata__3 **  
        |  |                             
        +--+  +-- Lamprotula_cornuumlunae **
           |  |                          
           +--+-- Lamprotula_caveata__1 **  
              |                          
              +-- Lamprotula_caveata__2 **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
33245115
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_caveata **     
  +--+                                
     |  +----- Lamprotula_leaii__2 **    
     +--+                             
        |  +-- Lamprotula_seomjinensis **
        +--+                          
           +-- Lamprotula_leaii__1 **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3313198223
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
331350223
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
22022900--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
22027122--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
22021832--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
22022911--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
22023411--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
22023411--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
22024222--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
22025111--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2202600--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
22023411--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_leaii **  
  +--+                     
     +-- Lamprotula_caveata **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
22022222--
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
22022111--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
22021000--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
22024322--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
22028322--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
22022900--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
22023211--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
22021300--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
22024622--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
22027122--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
22027922--
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
22027122--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
22024522--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
22025422--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
22024511--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
22024422--
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
220213722--
     +-- Potomida          
  +--+                     
  |  +-- Pronodularia      
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
22023411--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
22024222--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
22025422--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
22023211--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2202400--
  +-- Lamprotula_leaii **       
--+                          
  +-- Lamprotula_seomjinensis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
22026611--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
213021232212
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida              
  |                         
  +-- Pronodularia          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
22023411--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
22021832--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
22028922--
  +-------- Pronodularia      
--+                           
  |  +----- Potomida          
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
22028922--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
22023622--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
22024022--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22021400--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
22022911--
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
22023922--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
22028522--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
22025122--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
23402320033
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
                    |     +-- Anemina                 
  +-----------------+                                 
  |                 |     +-- Lamprotula_caveata **      
  |                 |  +--+                           
  |                 |  |  +-- Chamberlainia           
  |                 +--+                              
  |                    |  +-- Pseudodon               
  |                    +--+                           
  |                       +-- Pilsbryoconcha          
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Scriptolamprotula       
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodonta                
     |                                                
     |                    +-- Utterbackia_peninsularis
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii **        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrula                
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
22024622--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
220213722--
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
22025222--
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
22023411--
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
22024322--
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011200--
-- Lamprotula_caveata **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
11015822--
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Lamprotula_caveata **
--+                     
  +-- Pronodularia

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014311--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
11013322--
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
1101642--
-- Lamprotula_leaii **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
11011122--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015600--
-- Lamprotula_cornuumlunae **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014011--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014022--
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
11014922--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011500--
-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017522--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014011--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011411--
-- Lamprotula_leaii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017522--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
11012800--
-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012222--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
11012511--
-- Lamprotula_leaii **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
11014011--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1101600--
-- Lamprotula_caveata **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014522--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011800--
-- Lamprotula_leaii **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012011--
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013611--
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011411--
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
11013000--
-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016822--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
11013422--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011222--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014722--
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101700--
-- Lamprotula_leaii **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
11011300--
-- Lamprotula_caveata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013822--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
11018111--
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
11014011--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014022--
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016822--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
11013511--
  +-- Lamprotula_caveata **
--+                     
  +-- Potomida

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012300--
-- Lamprotula_leaii **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
11011422--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018122--
-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016822--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Wu et al., 2026 [Publ 4825], fig. 5

iTtTbiriSoTroTroSiD
11011811--
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013022--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012300--
-- Lamprotula_leaii **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011511--
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011622--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011322--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013122--
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011400--
-- Lamprotula_leaii **

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Lamprotula_leaii **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011222--
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014311--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011000--
-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015222--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Lamprotula_leaii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015600--
-- Lamprotula_cornuumlunae **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013911--
  +-- Lamprotula_caveata **
--+                     
  +-- Potomida

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012111--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
11014811--
-- Lamprotula_leaii **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013300--
-- Lamprotula_leaii **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012200--
-- Lamprotula_leaii **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013211--
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
11011911--
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101600--
-- Lamprotula_leaii **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Lamprotula_leaii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011922--
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
  +-- Potomida        
--+                   
  +-- Lamprotula_leaii **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
11013511--
  +-- Lamprotula_caveata **
--+                     
  +-- Potomida

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013922--
  +----- Potomida          
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Lamprotula_caveata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013522--
-- Lamprotula_leaii **
 
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