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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Margaritiferidae | Gibbosulinae

genus Gibbosula Simpson, 1900: cladomics
     5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 85 cladograms in 50 publications
maximum number of terminals in an analysis = 4 (mean = 1.38, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 23
maximum number of included terminals = 4, (mean = 2.39 median = 2)
proportion of cladograms that support basic monophyly = 0.83

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Chen et al., 2025 [Publ 4455], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
442422000
     +-- Gibbosula_laosensis **     
  +--+                           
  |  +-- Gibbosula_sinensis **      
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Chen et al., 2026 [Publ 4827], fig. 2

iTtTbiriSoTroTroSiD
44248002
  +-------- Gibbosula_crassa **        
--+                                 
  |  +----- Gibbosula_rochechouartii **
  +--+                              
     |  +-- Gibbosula_laosensis **     
     +--+                           
        +-- Gibbosula_sinensis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
331325003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3313137003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
3313629001
  +----- Gibbosula_laosensis **     
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Lyubas et al., 2019 [Publ 3963], fig. 10

iTtTbiriSoTroTroSiD
331313003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
331325003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
22027100--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
2202800--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
22023200--
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
22027100--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
22026700--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
22029300--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
22026700--
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
22026700--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
22029300--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
213021970013
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
213021990013
  +-------- Gibbosula_rochechouartii **
  |                                 
--+  +----- Gibbosula_laosensis **     
  |  |                              
  +--+  +-- Cumberlandia            
     |  |                           
     +--+-- Pseudunio               
        |                           
        +-- Margaritifera

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
21202320012
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
22026600--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Chen et al., 2026 [Publ 4833], fig. 1

iTtTbiriSoTroTroSiD
22022500--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_sinensis **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022700--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
213021990013
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Cumberlandia            
     +--+                              
        |  +-- Pseudunio               
        +--+                           
           +-- Margaritifera

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Gibbosula_crassa **

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
11011000--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012100--
-- Gibbosula_laosensis **

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
1101900--
-- Gibbosula_laosensis **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
11016300--
-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014300--
-- Gibbosula_rochechouartii **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
11019400--
-- Gibbosula_rochechouartii **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
11015000--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_crassa **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_laosensis **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
1101600--
-- Gibbosula_crassa **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_crassa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Gibbosula_laosensis **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_crassa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Gibbosula_laosensis **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
11014500--
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018400--
-- Gibbosula_rochechouartii **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014500--
-- Gibbosula_rochechouartii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017200--
-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014600--
-- Gibbosula_laosensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111600--
-- Gibbosula_laosensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Gibbosula_rochechouartii **

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014700--
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015500--
-- Gibbosula_rochechouartii **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012900--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014500--
-- Gibbosula_laosensis **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_crassa **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113800--
-- Gibbosula_laosensis **

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
1101700--
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
1101500--
-- Gibbosula_crassa **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Gibbosula_laosensis **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012900--
-- Gibbosula_laosensis **

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
1101900--
-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019000--
-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019000--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012100--
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014300--
-- Gibbosula_rochechouartii **

Wu et al., 2026 [Publ 4825], fig. 6B

iTtTbiriSoTroTroSiD
11016400--
-- Gibbosula_rochechouartii **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Gibbosula_rochechouartii **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014100--
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015200--
-- Gibbosula_rochechouartii **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
11011600--
-- Gibbosula_rochechouartii **

Kashiri et al., 2024 [Publ 4822], fig. 3

iTtTbiriSoTroTroSiD
1101900--
-- Gibbosula_laosensis **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
11014800--
-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
11014600--
-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018600--
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Gibbosula_rochechouartii **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_crassa **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
11014700--
-- Gibbosula_laosensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113800--
-- Gibbosula_laosensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015300--
-- Gibbosula_crassa **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_laosensis **
 
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