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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Margaritiferidae | Gibbosulinae

genus Gibbosula Simpson, 1900: cladomics
     5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 81 cladograms in 46 publications
maximum number of terminals in an analysis = 4 (mean = 1.35, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 4, (mean = 2.33 median = 2)
proportion of cladograms that support basic monophyly = 0.81

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Chen et al., 2025 [Publ 4455], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
442422000
     +-- Gibbosula_laosensis **     
  +--+                           
  |  +-- Gibbosula_sinensis **      
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3313138003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
3313628001
  +----- Gibbosula_laosensis **     
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_rochechouartii **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
331325003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **

Lyubas et al., 2019 [Publ 3963], fig. 10

iTtTbiriSoTroTroSiD
331313003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
331325003
  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
22029400--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
22026600--
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
22027200--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
22023200--
  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
213021980013
  +-------- Gibbosula_rochechouartii **
  |                                 
--+  +----- Gibbosula_laosensis **     
  |  |                              
  +--+  +-- Cumberlandia            
     |  |                           
     +--+-- Pseudunio               
        |                           
        +-- Margaritifera

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
21202320012
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
22026600--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
213021960013
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
22029400--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
213021980013
  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Cumberlandia            
     +--+                              
        |  +-- Pseudunio               
        +--+                           
           +-- Margaritifera

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
22027200--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022700--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
2202800--
  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
22026600--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
22026700--
  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012100--
-- Gibbosula_laosensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014800--
-- Gibbosula_rochechouartii **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014600--
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014500--
-- Gibbosula_laosensis **

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
1101700--
-- Gibbosula_laosensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Gibbosula_laosensis **

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
1101900--
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014500--
-- Gibbosula_rochechouartii **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
11019400--
-- Gibbosula_rochechouartii **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113700--
-- Gibbosula_laosensis **

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
1101900--
-- Gibbosula_laosensis **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017300--
-- Gibbosula_rochechouartii **

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
1101500--
-- Gibbosula_crassa **

Dai et al., 2026 [Publ 4757], fig. 3

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
1101600--
-- Gibbosula_crassa **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015500--
-- Gibbosula_rochechouartii **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012900--
-- Gibbosula_laosensis **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
11014700--
-- Gibbosula_laosensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015300--
-- Gibbosula_crassa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_crassa **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
11014800--
-- Gibbosula_laosensis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019000--
-- Gibbosula_laosensis **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012100--
-- Gibbosula_laosensis **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
11014600--
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_laosensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_laosensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014100--
-- Gibbosula_rochechouartii **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_rochechouartii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Gibbosula_laosensis **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_crassa **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
11011600--
-- Gibbosula_rochechouartii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Gibbosula_rochechouartii **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015300--
-- Gibbosula_crassa **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012900--
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
11013000--
-- Gibbosula_crassa **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
11015000--
-- Gibbosula_laosensis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018700--
-- Gibbosula_rochechouartii **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111600--
-- Gibbosula_laosensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
11012000--
-- Gibbosula_rochechouartii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Gibbosula_crassa **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_laosensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113700--
-- Gibbosula_laosensis **

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
11011000--
-- Gibbosula_laosensis **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Gibbosula_rochechouartii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018500--
-- Gibbosula_rochechouartii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015200--
-- Gibbosula_rochechouartii **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
11016400--
-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
11014400--
-- Gibbosula_rochechouartii **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019000--
-- Gibbosula_laosensis **

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
11011200--
-- Gibbosula_crassa **
 
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