The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini genus Megalonaias Utterback, 1915: cladomics
2 Recent valid species (2 total species) Cladomics Summarynumber of species = 2 (2 total)
analyzed in 30 cladograms in 20 publications
maximum number of terminals in an analysis =
2 (mean = 1.07, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 2
maximum number of included terminals = 2, (mean =
2 median = 2)
proportion of cladograms that support basic monophyly
= 1
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 628 | 19 | 2 | -- | +-------- Uniomerus
+-----------+
| | +----- Megalonaias_nickliniana__2 **
| +--+
| | +-- Megalonaias_nervosa **
| +--+
| +-- Megalonaias_nickliniana__1 **
|
| +-- Theliderma_johnsoni
| +--------+
--+ | +-- Theliderma_metanevra
| |
| +-----+ +-- Quadrula_nobilis
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Theliderma_cylindrica
| | +--+
| | | +-- Quadrula_fragosa
+--+ +--+
| +-- Quadrula_quadrula
|
| +----- Pustulosa_kieneriana
| +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
Neemuchwala et al., 2023 [Publ 4292], fig. 2 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 21 | 2 | -- | +-------------- Megalonaias_nervosa **
--+
| +----------- Uniomerus
+--+
| +-------- Theliderma
+--+
| +----- Cyclonaias
+--+
| +-- Quadrula
+--+
+-- Pustulosa
Pfeiffer et al., 2018 [Publ 3835], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 2 | 2 | -- | +-------- Quadrula
--+
| +----- Uniomerus
+--+
| +-- Megalonaias_nervosa **
+--+
+-- Megalonaias_nickliniana **
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 6 | 1 | -- | -- Megalonaias_nervosa **
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 13 | 1 | -- | +-------- Megalonaias_nervosa **
--+
| +----- Theliderma
+--+
| +-- Quadrula
+--+
+-- Pustulosa
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 3 | 2 | -- | +----- Megalonaias_nervosa **
--+
| +-- Quadrula
+--+
+-- Theliderma
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | -- Megalonaias_nervosa **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 18 | 2 | -- | -- Megalonaias_nervosa **
Szumowski et al., 2012 [Publ 3569], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 13 | 1 | -- | +----- Megalonaias_nervosa **
|
--+----- Theliderma
|
| +-- Quadrula
+--+
+-- Pustulosa
Mulvey et al., 1997 [Publ 3390], fig. 1_3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | -- Megalonaias_nervosa **
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 3 | 2 | -- | -- Megalonaias_nervosa **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 3 | 2 | -- | -- Megalonaias_nervosa **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 3 | 2 | -- | +----- Quadrula
--+
| +-- Megalonaias_nervosa **
+--+
+-- Uniomerus
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 3 | 2 | -- | -- Megalonaias_nervosa **
Johnson et al., 2018 [Publ 3844], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 13 | 2 | -- | +-------------------------- Megalonaias_nervosa **
|
--+ +----------------------- Uniomerus
| |
+--+ +-------------------- Theliderma
| |
+--+ +----------------- Quadrula
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 15 | 2 | -- | +----------------------------- Megalonaias_nervosa **
|
--+ +-------------------------- Uniomerus
| |
+--+ +----------------------- Theliderma
| |
+--+ +-------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +--------------+
| | +-- Quadrula_nobilis
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 13 | 1 | -- | +----------- Megalonaias_nervosa **
|
--+ +-- Quadrula_verrucosa
| +-----+
| | +-- Quadrula_nobilis
+--+
| +----- Quadrula_quadrula
+--+
| +-- Theliderma
+--+
+-- Pustulosa
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 10 | 2 | -- | +-------------------- Megalonaias_nervosa **
|
--+ +----------------- Uniomerus
| |
+--+ +-------------- Theliderma
| |
+--+ +----------- Quadrula_quadrula
| |
| | +-- Quadrula_nobilis
+--+ +-----+
| | +-- Quadrula_verrucosa
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 3 | 2 | -- | -- Megalonaias_nervosa **
Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 3 | 1 | -- | -- Megalonaias_nervosa **
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Uniomerus
--+
+-- Megalonaias_nervosa **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 17 | 2 | -- | +-- Megalonaias_nervosa **
+-----------------------+
| +-- Uniomerus
|
--+ +----------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +-----------------+
| | +-- Quadrula_nobilis
| |
+--+ +----------------- Theliderma
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 6 | 1 | -- | +----------------- Megalonaias_nervosa **
--+
| +-------------- Quadrula_quadrula
+--+
| +----------- Quadrula_verrucosa
+--+
| +-------- Theliderma
+--+
| +----- Pustulosa_pustulosa
+--+
| +-- Cyclonaias
+--+
+-- Pustulosa_nodulata
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 4 | 2 | -- | +----------- Megalonaias_nervosa **
--+
| +-------- Uniomerus
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Cyclonaias
+--+
+-- Quadrula_quadrula
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | -- Megalonaias_nervosa **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 10 | 2 | -- | +-- Megalonaias_nervosa **
+-----------+
| +-- Uniomerus
--+
| +----------- Theliderma
| |
+--+ +-------- Quadrula
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 3 | 2 | -- | +----- Quadrula
--+
| +-- Megalonaias_nervosa **
+--+
+-- Uniomerus
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 3 | 2 | -- | +----- Megalonaias_nervosa **
--+
| +-- Quadrula
+--+
+-- Theliderma
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 2 | 1 | -- | +----- Megalonaias_nervosa **
--+
| +-- Cyclonaias
+--+
+-- Pustulosa
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 3 | 2 | -- | +-------- Megalonaias_nervosa **
--+
| +----- Quadrula_quadrula__2
+--+
| +-- Quadrula_quadrula__1
+--+
+-- Theliderma
|
