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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Amblemini

genus Amblema Rafinesque, 1820: cladomics
     3 Recent valid species (12 total species)

Cladomics Summary

number of species = 3 (12 total)
analyzed in 127 cladograms in 86 publications
maximum number of terminals in an analysis = 3 (mean = 1.22, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 23
maximum number of included terminals = 3, (mean = 2.22 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3313629320
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Mulvey et al., 1997 [Publ 3390], fig. 1_3

iTtTbiriSoTroTroSiD
33132002
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_neislerii **
  +--+                    
     +-- Amblema_elliottii **

Inoue et al., 2020 [Publ 4128], fig. 2

iTtTbiriSoTroTroSiD
331320212
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_neislerii **

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
331319210
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Patterson et al., 2021 [Publ 4222], fig. 3

iTtTbiriSoTroTroSiD
33131000
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
22022600--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
22027132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
22024032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
220210332--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
22022921--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
22022400--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
22024132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
22027132--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
220210032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Inoue et al., 2020 [Publ 3941], fig. 3

iTtTbiriSoTroTroSiD
22024021--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Burlakova et al., 2012 [Publ 3486], fig. 1

iTtTbiriSoTroTroSiD
22023911--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
22023621--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iTtTbiriSoTroTroSiD
22021300--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Inoue et al., 2020 [Publ 3941], fig. 4

iTtTbiriSoTroTroSiD
22023921--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
22022700--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Kandl et al., 2001 [Publ 3384], fig. 5

iTtTbiriSoTroTroSiD
1101800--
-- Amblema_plicata **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112100--
-- Amblema_plicata **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Grobler et al., 2011 [Publ 3359], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014500--
-- Amblema_plicata **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
11015600--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Keogh & Simons, 2019 [Publ 3935], fig. S1

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
11011011--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018400--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011800--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11018000--
-- Amblema_plicata **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Perkins et al., 2017 [Publ 3777], fig. 2

iTtTbiriSoTroTroSiD
11013611--
-- Amblema_plicata **

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110113900--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
11011011--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Amblema_plicata **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018100--
-- Amblema_plicata **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014400--
-- Amblema_plicata **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119811--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110120011--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016700--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112400--
-- Amblema_plicata **

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
11014022--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
11011200--
-- Amblema_plicata **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
11011621--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
11013411--
-- Amblema_plicata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013211--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
11012611--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
11013711--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
11011111--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012711--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
11012600--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Szumowski et al., 2012 [Publ 3569], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018600--
-- Amblema_plicata **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110120011--
-- Amblema_plicata **

Inoue et al., 2014 [Publ 3635], fig. 3B

iTtTbiriSoTroTroSiD
1101700--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Amblema_plicata **
 
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