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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Amblemini

genus Amblema Rafinesque, 1820: cladomics
     3 Recent valid species (12 total species)

Cladomics Summary

number of species = 3 (12 total)
analyzed in 124 cladograms in 84 publications
maximum number of terminals in an analysis = 3 (mean = 1.21, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included terminals = 3, (mean = 2.18 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Inoue et al., 2020 [Publ 4128], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
331320210
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_neislerii **

Campbell et al., 2005 [Publ 2660], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
220210032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Patterson et al., 2021 [Publ 4222], fig. 3

iTtTbiriSoTroTroSiD
33131002
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Mulvey et al., 1997 [Publ 3390], fig. 1_3

iTtTbiriSoTroTroSiD
33132000
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_neislerii **
  +--+                    
     +-- Amblema_elliottii **

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
331319212
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Inoue et al., 2020 [Publ 3941], fig. 3

iTtTbiriSoTroTroSiD
22024021--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
220210232--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
22022600--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iTtTbiriSoTroTroSiD
22021300--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
22024032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
22022700--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
22022921--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Inoue et al., 2020 [Publ 3941], fig. 4

iTtTbiriSoTroTroSiD
22023921--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Burlakova et al., 2012 [Publ 3486], fig. 1

iTtTbiriSoTroTroSiD
22023911--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
22024132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
22023621--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
22027132--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
22022400--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
22027132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014500--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112400--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Kandl et al., 2001 [Publ 3384], fig. 5

iTtTbiriSoTroTroSiD
1101800--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
11012600--
-- Amblema_plicata **

Inoue et al., 2014 [Publ 3635], fig. 3B

iTtTbiriSoTroTroSiD
1101700--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
11011111--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Amblema_plicata **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014500--
-- Amblema_plicata **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
11011621--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016700--
-- Amblema_plicata **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
11013411--
-- Amblema_plicata **

Grobler et al., 2011 [Publ 3359], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013211--
-- Amblema_plicata **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
11012611--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
11011011--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119811--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
11015600--
-- Amblema_plicata **

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Amblema_plicata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018700--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015100--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
11013711--
-- Amblema_plicata **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012711--
-- Amblema_plicata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018400--
-- Amblema_plicata **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Szumowski et al., 2012 [Publ 3569], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Perkins et al., 2017 [Publ 3777], fig. 2

iTtTbiriSoTroTroSiD
11013511--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Keogh & Simons, 2019 [Publ 3935], fig. S1

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013100--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011800--
-- Amblema_plicata **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112000--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119611--
-- Amblema_plicata **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018200--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
11011011--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
11011200--
-- Amblema_plicata **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119811--
-- Amblema_plicata **

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
11014022--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Amblema_plicata **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **
 
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