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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Amblemini

genus Amblema Rafinesque, 1820: cladomics
     3 Recent valid species (12 total species)

Cladomics Summary

number of species = 3 (12 total)
analyzed in 123 cladograms in 83 publications
maximum number of terminals in an analysis = 3 (mean = 1.21, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included terminals = 3, (mean = 2.18 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Inoue et al., 2020 [Publ 4128], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
331323210
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_neislerii **

Campbell et al., 2005 [Publ 2660], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
220210032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
331322212
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Patterson et al., 2021 [Publ 4222], fig. 3

iTtTbiriSoTroTroSiD
33131002
  +----- Amblema_neislerii **
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Mulvey et al., 1997 [Publ 3390], fig. 1_3

iTtTbiriSoTroTroSiD
33132000
  +----- Amblema_plicata **  
--+                       
  |  +-- Amblema_neislerii **
  +--+                    
     +-- Amblema_elliottii **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
22022600--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
22024032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
22024032--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
22027132--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
22027132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
22023621--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210032--
  +----- Reginaia         
--+                       
  |  +-- Amblema_elliottii **
  +--+                    
     +-- Amblema_plicata **

Inoue et al., 2020 [Publ 3941], fig. 3

iTtTbiriSoTroTroSiD
22024021--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_elliottii **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
22023321--
  +----- Reginaia         
--+                       
  |  +-- Amblema_plicata **  
  +--+                    
     +-- Amblema_neislerii **

Inoue et al., 2020 [Publ 3941], fig. 4

iTtTbiriSoTroTroSiD
22023921--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iTtTbiriSoTroTroSiD
22021300--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
220210232--
  +-- Amblema_plicata **  
--+                    
  +-- Amblema_elliottii **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
22022700--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
22024132--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Burlakova et al., 2012 [Publ 3486], fig. 1

iTtTbiriSoTroTroSiD
22023911--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
22022400--
  +-- Amblema_elliottii **
--+                    
  +-- Amblema_plicata **

Inoue et al., 2014 [Publ 3635], fig. 3B

iTtTbiriSoTroTroSiD
1101700--
-- Amblema_plicata **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
11015600--
-- Amblema_plicata **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015200--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113800--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
11011111--
-- Amblema_plicata **

Kandl et al., 2001 [Publ 3384], fig. 5

iTtTbiriSoTroTroSiD
1101800--
-- Amblema_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119811--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119811--
-- Amblema_plicata **

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
11012722--
  +----- Reginaia       
--+                     
  |  +-- Amblema_plicata **
  +--+                  
     +-- Plectomerus

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
11014022--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Perkins et al., 2017 [Publ 3777], fig. 2

iTtTbiriSoTroTroSiD
11013511--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014500--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016700--
-- Amblema_plicata **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
11011621--
  +-- Amblema_plicata **
--+                  
  +-- Reginaia

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
11013411--
-- Amblema_plicata **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
11012611--
-- Amblema_plicata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112400--
-- Amblema_plicata **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012111--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
11011011--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119611--
-- Amblema_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018200--
-- Amblema_plicata **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014500--
-- Amblema_plicata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017200--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iTtTbiriSoTroTroSiD
11011700--
-- Amblema_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015622--
  +----- Reginaia       
--+                     
  |  +-- Plectomerus    
  +--+                  
     +-- Amblema_plicata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015500--
-- Amblema_plicata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Amblema_plicata **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
11012600--
-- Amblema_plicata **

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
11013711--
-- Amblema_plicata **

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011800--
-- Amblema_plicata **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Amblema_plicata **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
11011111--
  +-- Amblema_plicata **
--+                  
  +-- Plectomerus

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011900--
-- Amblema_plicata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Amblema_plicata **

Szumowski et al., 2012 [Publ 3569], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1101400--
-- Amblema_plicata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112000--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013211--
-- Amblema_plicata **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
11012011--
-- Amblema_plicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
11011011--
-- Amblema_plicata **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
11011400--
-- Amblema_plicata **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
11011200--
-- Amblema_plicata **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Keogh & Simons, 2019 [Publ 3935], fig. S1

iTtTbiriSoTroTroSiD
1101900--
-- Amblema_plicata **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013100--
-- Amblema_plicata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018700--
-- Amblema_plicata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015100--
-- Amblema_plicata **

Grobler et al., 2011 [Publ 3359], fig. 2

iTtTbiriSoTroTroSiD
11011600--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014600--
-- Amblema_plicata **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
-- Amblema_plicata **

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
11012822--
-- Amblema_plicata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019000--
-- Amblema_plicata **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012711--
-- Amblema_plicata **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012900--
-- Amblema_plicata **

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Amblema_plicata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019400--
-- Amblema_plicata **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Amblema_plicata **

Khalloufi et al., 2011 [Publ 2748], fig. 2

iTtTbiriSoTroTroSiD
11011000--
-- Amblema_plicata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013500--
-- Amblema_plicata **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Amblema_plicata **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011300--
-- Amblema_plicata **
 
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