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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Etherioidea | Iridinidae | Aspathariinae

genus Chambardia Bourguignat in Servain, 1890: cladomics
     11 Recent valid species (12 total species)

Cladomics Summary

number of species = 11 (12 total)
analyzed in 37 cladograms in 22 publications
maximum number of terminals in an analysis = 2 (mean = 1.05, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 2
maximum number of included terminals = 2, (mean = 2 median = 2)
proportion of cladograms that support basic monophyly = 1

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
220362921--
  +-------- Aspatharia             
--+                                
  |  +----- Chambardia_wahlbergi__2 **
  +--+                             
     |  +-- Chambardia_rubens **      
     +--+                          
        +-- Chambardia_wahlbergi__1 **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
22022600--
  +-- Chambardia_rubens **   
--+                       
  +-- Chambardia_wahlbergi **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119911--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013500--
-- Chambardia_rubens **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013511--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Chambardia_wahlbergi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110114000--
-- Chambardia_rubens **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012911--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Chambardia_rubens **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112111--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Chambardia_rubens **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014511--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014611--
-- Chambardia_wahlbergi **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113711--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014511--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019011--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11013000--
-- Chambardia_wahlbergi **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Chambardia_wahlbergi **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
11013311--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
11011411--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119911--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014611--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019011--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012711--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013511--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017911--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012511--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012911--
  +-- Chambardia_wahlbergi **
--+                       
  +-- Aspatharia

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
11013311--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016711--
  +-- Aspatharia          
--+                       
  +-- Chambardia_wahlbergi **
 
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