The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina genus Theliderma Swainson, 1840: cladomics
6 Recent valid species (6 total species) Cladomics Summarynumber of species = 6 (6 total)
analyzed in 24 cladograms in 16 publications
maximum number of terminals in an analysis =
5 (mean = 1.88, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 9
maximum number of included terminals = 5, (mean =
3.33 median = 4)
proportion of cladograms that support basic monophyly
= 0.89
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 24 | 12 | 3 | 0 | +----------- Theliderma_cylindrica **
+-----------+
| | +-------- Theliderma_sparsa **
| +--+
| | +----- Theliderma_intermedia **
| +--+
--+ | +-- Theliderma_metanevra **
| +--+
| +-- Theliderma_johnsoni **
|
| +-------------------- Quadrula
| |
+--+ +----------------- Pustulosa_kieneriana
| |
| | +-- Pustulosa_infucata
+--+ +-----------+
| | +-- Pustulosa_kleiniana
| |
+--+ +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
Neemuchwala et al., 2023 [Publ 4292], fig. 2 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 23 | 13 | 3 | 3 | +----- Cyclonaias
+-----+
| | +-- Quadrula
| +--+
--+ +-- Pustulosa
|
| +-------- Theliderma_cylindrica **
+--+
| +----- Theliderma_metanevra **
+--+
| +-- Theliderma_sparsa **
+--+
+-- Theliderma_johnsoni **
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 25 | 9 | 2 | 3 | +-- Quadrula
+--------+
| +-- Pustulosa
--+
| +-------- Theliderma_cylindrica **
+--+
| +----- Theliderma_intermedia **
+--+
| +-- Theliderma_sparsa **
+--+
+-- Theliderma_metanevra **
Szumowski et al., 2012 [Publ 3569], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 10 | 9 | 2 | 3 | +-------- Theliderma_cylindrica **
--+
| +----- Theliderma_intermedia **
+--+
| +-- Theliderma_metanevra **
+--+
+-- Theliderma_sparsa **
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 25 | 9 | 2 | 1 | +-- Quadrula_verrucosa
+--------------+
| +-- Quadrula_nobilis
--+
| +-------------- Quadrula_quadrula
+--+
| +----------- Pustulosa
+--+
| +-------- Theliderma_cylindrica **
+--+
| +----- Theliderma_sparsa **
+--+
| +-- Theliderma_intermedia **
+--+
+-- Theliderma_metanevra **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 7 | 5 | 3 | 629 | 13 | 3 | 6 | +-- Theliderma_johnsoni **
+--------+
| +-- Theliderma_metanevra **
|
+-----+ +-- Quadrula_nobilis
| | +-----+
| | | +-- Quadrula_verrucosa
| +--+
| | +----- Theliderma_cylindrica **
| +--+
| | +-- Quadrula_fragosa
--+ +--+
| +-- Quadrula_quadrula
|
| +----- Pustulosa_kieneriana
| +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 12 | 3 | -- | +----------------------- Quadrula_quadrula
|
| +-- Quadrula_verrucosa
--+ +-----------------+
| | +-- Quadrula_nobilis
| |
+--+ +-- Theliderma_metanevra **
| +--------------+
| | +-- Theliderma_johnsoni **
| |
+--+ +----- Pustulosa_kieneriana
| +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 24 | 12 | 3 | -- | +-- Theliderma_metanevra **
+--------------------+
| +-- Theliderma_johnsoni **
|
--+ +-------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +--------------+
| | +-- Quadrula_nobilis
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 5 | 2 | -- | +-- Theliderma_intermedia **
+-----------+
| +-- Theliderma_sparsa **
--+
| +----------- Quadrula_quadrula
+--+
| +-------- Pustulosa_infucata
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Pustulosa_kieneriana
+--+
+-- Pustulosa_succissa
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 5 | 3 | -- | +-------------- Quadrula_quadrula
--+
| +----------- Quadrula_verrucosa
+--+
| +-------- Theliderma_metanevra **
+--+
| +----- Pustulosa_pustulosa
+--+
| +-- Cyclonaias
+--+
+-- Pustulosa_nodulata
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 1 | 1 | -- | +-- Quadrula
--+
+-- Theliderma_metanevra **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 8 | 3 | -- | +-------------- Theliderma_metanevra **
|
--+ +----------- Quadrula_quadrula
| |
| | +-- Quadrula_nobilis
+--+ +-----+
| | +-- Quadrula_verrucosa
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
Johnson et al., 2018 [Publ 3844], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 11 | 3 | -- | +-------------------- Theliderma_metanevra **
|
--+ +----------------- Quadrula
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 5 | 3 | -- | +----------- Theliderma_metanevra **
--+
| +-------- Quadrula
+--+
| +----- Pustulosa_pustulosa
+--+
| +-- Cyclonaias
+--+
+-- Pustulosa_nodulata
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 1 | 1 | -- | +----- Quadrula_quadrula__2
--+
| +-- Quadrula_quadrula__1
+--+
+-- Theliderma_metanevra **
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Quadrula
--+
+-- Theliderma_metanevra **
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | -- Theliderma_cylindrica **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 4 | 3 | -- | +-- Cyclonaias
|
+-- Pustulosa_pustulosa__1
|
+-- Theliderma_metanevra **
--+
+-- Pustulosa_pustulosa__2
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 8 | 3 | -- | +-- Quadrula_verrucosa
+-----------+
| +-- Quadrula_nobilis
|
--+ +----------- Quadrula_quadrula
| |
| +----------- Theliderma_metanevra **
+--+
| +-- Pustulosa_infucata
| +-----+
| | +-- Pustulosa_kleiniana
+--+
| +----- Cyclonaias
+--+
| +-- Pustulosa_pustulosa
+--+
+-- Pustulosa_succissa
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | -- Theliderma_cylindrica **
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | -- Theliderma_cylindrica **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Theliderma_metanevra **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 8 | 3 | -- | +----------------- Theliderma_metanevra **
|
--+ +-------------- Quadrula_verrucosa
| |
| | +-- Quadrula_quadrula
+--+ +--------+
| | +-- Quadrula_nobilis
| |
+--+ +-- Pustulosa_infucata
| +-----+
| | +-- Pustulosa_kleiniana
+--+
| +----- Cyclonaias
+--+
| +-- Pustulosa_pustulosa
+--+
+-- Pustulosa_succissa
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 8 | 3 | -- | +----------- Theliderma_metanevra **
|
--+ +-------- Quadrula
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
|
