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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina

genus Cuneopsis Simpson, 1900: cladomics
     7 Recent valid species (10 total species)

Cladomics Summary

number of species = 7 (10 total)
analyzed in 74 cladograms in 45 publications
maximum number of terminals in an analysis = 7 (mean = 2.62, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 52
maximum number of included terminals = 7, (mean = 3.31 median = 3)
proportion of cladograms that support basic monophyly = 0.81


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Hou et al., 2025 [Publ 4432], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
77584220
  +----------- Cuneopsis_rufescens **    
  |                                   
  |     +----- Cuneopsis_celtiformis **  
--+  +--+                             
  |  |  |  +-- Cuneopsis_celtiosimilis **
  |  |  +--+                          
  +--+     +-- Cuneopsis_kiangsiensis ** 
     |                                
     |  +----- Cuneopsis_szechenyii **   
     |  |                             
     +--+  +-- Cuneopsis_heudei **       
        |  |                          
        +--+-- Cuneopsis_demangei__1 **  
           |                          
           +-- Cuneopsis_demangei__2 **  

Chen et al., 2023 [Publ 4302], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
220239126--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii ** 

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
7757211841
  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           |  |     +----- Cuneopsis_szechenyii **   
           +--+  +--+                             
              |  |  |  +-- Cuneopsis_demangei **     
              |  |  +--+                          
              +--+     +-- Cuneopsis_heudei **       
                 |                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_kiangsiensis ** 
                    +--+                          
                       +-- Cuneopsis_celtiosimilis **

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
6646221942
  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+  +----------------- Schistodesmus         
     |  |                                        
     +--+  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_demangei **    
                    +--+                         
                       +-- Cuneopsis_heudei **      

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
6646591742
  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           +--+        +-- Cuneopsis_heudei **       
              |  +-----+                          
              |  |     +-- Cuneopsis_szechenyii **   
              +--+                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_celtiosimilis **
                    +--+                          
                       +-- Cuneopsis_kiangsiensis ** 

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
6131169629
           +----- Cuneopsis_rufescens **   
  +--------+                            
  |        |  +-- Middendorffinaia      
  |        +--+                         
--+           +-- Pseudocuneopsis       
  |                                     
  |  +----------- Tchangsinaia          
  |  |                                  
  +--+        +-- Cuneopsis_kiangsiensis **
     |  +-----+                         
     |  |     +-- Cuneopsis_celtiformis ** 
     +--+                               
        |  +----- Cuneopsis_szechenyii **  
        +--+                            
           |  +-- Cuneopsis_heudei **      
           +--+                         
              +-- Cuneopsis_demangei **    

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
6646701842
  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+  +----------------- Schistodesmus         
     |  |                                        
     +--+  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_demangei **    
                    +--+                         
                       +-- Cuneopsis_heudei **      

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
6646801352
  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+                 +-- Tongchuania           
     |  +--------------+                         
     |  |              +-- Schistodesmus         
     +--+                                        
        |  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_heudei **      
                    +--+                         
                       +-- Cuneopsis_demangei **    

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
55357653
  +----------------------- Nodularia            
  |                                             
--+  +-------------------- Pseudocuneopsis      
  |  |                                          
  +--+  +----------------- Schistodesmus        
     |  |                                       
     +--+              +-- Tchangsinaia         
        |  +-----------+                        
        |  |           +-- Inversiunio          
        +--+                                    
           |  +----------- Cuneopsis_rufescens **  
           +--+                                 
              |  +-------- Cuneopsis_celtiformis **
              +--+                              
                 |  +----- Cuneopsis_szechenyii ** 
                 +--+                           
                    |  +-- Cuneopsis_demangei **   
                    +--+                        
                       +-- Cuneopsis_heudei **     

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4424197834
  +----------------- Schistodesmus        
--+                                       
  |  +-------------- Nodularia            
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **     

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
442443834
  +----------------- Nodularia            
--+                                       
  |  +-------------- Tchangsinaia         
  +--+                                    
     |  +----------- Schistodesmus        
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_heudei **     
              +--+                        
                 +-- Cuneopsis_szechenyii ** 

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
442470844
  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **   

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
442492844
  +-------------------- Pseudocuneopsis          
  |                                              
--+  +----------------- Nodularia                
  |  |                                           
  |  |           +----- Tchangsinaia             
  +--+  +--------+                               
     |  |        |  +-- Schistodesmus_xinyuensis 
     |  |        +--+                            
     +--+           +-- Schistodesmus_spinosus   
        |                                        
        |  +----------- Schistodesmus_lampreyanus
        +--+                                     
           |  +-------- Cuneopsis_rufescens **      
           +--+                                  
              |  +----- Cuneopsis_celtiformis **    
              +--+                               
                 |  +-- Cuneopsis_demangei **       
                 +--+                            
                    +-- Cuneopsis_heudei **         

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
442488844
  +----------------- Nodularia            
  |                                       
--+----------------- Pseudocuneopsis      
  |                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **     

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
453441835
  +----------------- Schistodesmus        
--+                                       
  |  +-------------- Nodularia            
  +--+                                    
     |  +----------- Cuneopsis_rufescens **  
     +--+                                 
        |  +-------- Tchangsinaia         
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **     

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4424841244
                 +-- Nodularia            
  +--------------+                        
  |              +-- Pseudocuneopsis      
--+                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_heudei **     
              +--+                        
                 +-- Cuneopsis_demangei **   

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4834197838
  +----- Cuneopsis_celtiformis **
  |                           
  +----- Tchangsinaia         
  |                           
--+----- Cuneopsis_rufescens **  
  |                           
  +----- Nodularia            
  |                           
  |  +-- Cuneopsis_szechenyii ** 
  +--+                        
     +-- Cuneopsis_heudei **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
442492844
                 +-- Nodularia            
  +--------------+                        
  |              +-- Pseudocuneopsis      
--+                                       
  |  +-------------- Tchangsinaia         
  +--+                                    
     |  +----------- Schistodesmus        
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
44241181444
  +-------------- Nodularia            
  |                                    
--+  +----------- Schistodesmus        
  |  |                                 
  |  |        +-- Tchangsinaia         
  +--+--------+                        
     |        +-- Inversiunio          
     |                                 
     |  +-------- Cuneopsis_rufescens **  
     +--+                              
        |  +----- Cuneopsis_celtiformis **
        +--+                           
           |  +-- Cuneopsis_szechenyii ** 
           +--+                        
              +-- Cuneopsis_heudei **     

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
4424491044
  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **   

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
4424351154
  +-------------------- Pseudocuneopsis      
  |                                          
--+  +----------------- Nodularia            
  |  |                                       
  +--+  +-------------- Schistodesmus        
     |  |                                    
     +--+           +-- Tchangsinaia         
        |  +--------+                        
        |  |        +-- Inversiunio          
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **   

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4424195734
  +----------------- Nodularia            
--+                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **     

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
4424501544
  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_demangei **   
                 +--+                        
                    +-- Cuneopsis_heudei **     

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
4424491844
  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_demangei **   
                 +--+                        
                    +-- Cuneopsis_heudei **     

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
442488844
  +----------------- Nodularia            
  |                                       
--+----------------- Pseudocuneopsis      
  |                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **     

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
364328438
        +-- Tchangsinaia         
  +-----+                        
  |     +-- Cuneopsis_rufescens **  
--+                              
  |  +----- Nodularia            
  +--+                           
     |  +-- Cuneopsis_szechenyii ** 
     +--+                        
        +-- Cuneopsis_celtiformis **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
331350435
           +-- Nodularia            
  +--------+                        
  |        +-- Schistodesmus        
--+                                 
  |  +-------- Tchangsinaia         
  +--+                              
     |  +----- Cuneopsis_rufescens **  
     +--+                           
        |  +-- Cuneopsis_celtiformis **
        +--+                        
           +-- Cuneopsis_szechenyii ** 

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
3566284311
  +----------------- Nodularia               
  |                                          
  |           +----- Cuneopsis_rufescens__2 **  
--+  +--------+                              
  |  |        |  +-- Schistodesmus           
  |  |        +--+                           
  +--+           +-- Cuneopsis_celtiformis__2 **
     |                                       
     |  +----------- Tchangsinaia            
     +--+                                    
        |  +-------- Cuneopsis_celtiformis__1 **
        +--+                                 
           |  +----- Cuneopsis_szechenyii__2 ** 
           +--+                              
              |  +-- Cuneopsis_szechenyii__1 ** 
              +--+                           
                 +-- Cuneopsis_rufescens__1 **  

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
230274437
  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Cuneopsis_szechenyii **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens ** 

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
260212549
  +-------------- Tchangsinaia       
--+                                  
  |  +----------- Cuneopsis_heudei **   
  +--+                               
     |  +-------- Nodularia          
     +--+                            
        |  +----- Pseudocuneopsis    
        +--+                         
           |  +-- Cuneopsis_rufescens **
           +--+                      
              +-- Schistodesmus      

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
210021234313
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Aculamprotula        
  |                        
  +-- Cuneopsis_szechenyii ** 
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens **  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus        

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
22028943--
  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_szechenyii **
  |  +--+                       
  |  |  +-- Cuneopsis_rufescens ** 
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus       

Hou et al., 2025 [Publ 4445], fig. 4

iTtTbiriSoTroTroSiD
2202551--
  +----- Schistodesmus        
--+                           
  |  +-- Cuneopsis_heudei **     
  +--+                        
     +-- Cuneopsis_celtiformis **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
22022295--
     +-- Cuneopsis_rufescens ** 
  +--+                       
  |  +-- Cuneopsis_szechenyii **
  |                          
  |  +-- Nodularia           
--+--+                       
  |  +-- Schistodesmus       
  |                          
  |  +-- Tchangsinaia        
  |  |                       
  +--+-- Inversiunio         
     |                       
     +-- Pseudocuneopsis     

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
22021674--
  +-------------- Pseudocuneopsis      
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Tchangsinaia         
     +--+                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Cuneopsis_celtiformis **
           +--+                        
              +-- Cuneopsis_heudei **     

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2302137437
  +----- Cuneopsis_szechenyii **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Cuneopsis_rufescens ** 

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
220243116--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii ** 

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
220239116--
           +-- Tchangsinaia         
  +--------+                        
  |        +-- Inversiunio          
  |                                 
--+     +----- Pseudocuneopsis      
  |  +--+                           
  |  |  |  +-- Cuneopsis_celtiformis **
  |  |  +--+                        
  +--+     +-- Cuneopsis_szechenyii ** 
     |                              
     |  +----- Nodularia            
     +--+                           
        |  +-- Diaurora             
        +--+                        
           +-- Schistodesmus        

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
22021352--
  +----- Schistodesmus       
  |                          
--+----- Nodularia           
  |                          
  |  +-- Cuneopsis_szechenyii **
  +--+                       
     +-- Cuneopsis_rufescens ** 

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
220239126--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii ** 

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
22025422--
  +-------- Nodularia           
--+                             
  |  +----- Tchangsinaia        
  +--+                          
     |  +-- Cuneopsis_rufescens ** 
     +--+                       
        +-- Cuneopsis_szechenyii **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
220213743--
  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_rufescens ** 
  |  +--+                       
  |  |  +-- Cuneopsis_szechenyii **
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus       

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
22022185--
           +-- Cuneopsis_szechenyii **
  +--------+                       
  |        +-- Cuneopsis_rufescens ** 
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
22027843--
  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis_szechenyii **
        +--+                       
           +-- Cuneopsis_rufescens ** 

Choi et al., 2020 [Publ 4169], fig. 2a

iTtTbiriSoTroTroSiD
22021093--
  +-- Cuneopsis_heudei **   
--+                      
  +-- Cuneopsis_rufescens **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
22024094--
  +-------------- Pseudocuneopsis      
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Schistodesmus        
     +--+                              
        |  +----- Tchangsinaia         
        +--+                           
           |  +-- Cuneopsis_heudei **     
           +--+                        
              +-- Cuneopsis_celtiformis **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
22022295--
           +-- Cuneopsis_szechenyii **
  +--------+                       
  |        +-- Cuneopsis_rufescens ** 
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis     

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
22028943--
  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_szechenyii **
  |  +--+                       
  |  |  +-- Cuneopsis_rufescens ** 
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus       

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
230274437
  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Cuneopsis_szechenyii **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens ** 

Wu et al., 2025 [Publ 4442], fig. 2

iTtTbiriSoTroTroSiD
22023300--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_heudei **     

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
220243116--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii ** 

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
220239116--
              +-- Tchangsinaia         
  +-----------+                        
  |           +-- Inversiunio          
--+                                    
  |  +----------- Pseudocuneopsis      
  |  |                                 
  +--+        +-- Cuneopsis_celtiformis **
     |  +-----+                        
     |  |     +-- Cuneopsis_szechenyii ** 
     +--+                              
        |  +----- Nodularia            
        +--+                           
           |  +-- Diaurora             
           +--+                        
              +-- Schistodesmus        

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014544--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus   

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016864--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Tchangsinaia    
        +--+                   
           +-- Cuneopsis_heudei **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012183--
  +-------- Nodularia            
--+                              
  |  +----- Cuneopsis_celtiformis **
  +--+                           
     |  +-- Inversiunio          
     +--+                        
        +-- Schistodesmus        

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015354--
        +-- Cuneopsis_heudei **
  +-----+                   
  |     +-- Schistodesmus   
--+                         
  |  +----- Pseudocuneopsis 
  +--+                      
     |  +-- Nodularia       
     +--+                   
        +-- Tchangsinaia    

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
11011674--
  +-------- Pseudocuneopsis 
  |                         
--+     +-- Cuneopsis_heudei **
  |  +--+                   
  |  |  +-- Schistodesmus   
  +--+                      
     |  +-- Tchangsinaia    
     +--+                   
        +-- Nodularia       

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011211--
  +-- Nodularia       
--+                   
  +-- Cuneopsis_heudei **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017364--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus   

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014844--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus   

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
11011711--
  +-- Nodularia            
--+                        
  +-- Cuneopsis_celtiformis **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015244--
        +-- Pseudocuneopsis      
  +-----+                        
  |     +-- Nodularia            
--+                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Cuneopsis_celtiformis **
     +--+                        
        +-- Tchangsinaia         

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013353--
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Nodularia            

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013933--
  +-------- Nodularia       
--+                         
  |  +----- Tchangsinaia    
  +--+                      
     |  +-- Cuneopsis_heudei **
     +--+                   
        +-- Schistodesmus   

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012954--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Tchangsinaia    

Wu et al., 2024 [Publ 4379], fig. 2

iTtTbiriSoTroTroSiD
110112123--
  +----- Nodularia            
  |                           
--+  +-- Cuneopsis_celtiformis **
  |  |                        
  +--+-- Tchangsinaia         
     |                        
     +-- Pseudocuneopsis      

Chen et al., 2026 [Publ 4805], fig. 4

iTtTbiriSoTroTroSiD
11013300--
-- Cuneopsis_celtiformis **

Choi et al., 2020 [Publ 4169], fig. 3a

iTtTbiriSoTroTroSiD
1101541--
-- Cuneopsis_heudei **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
11011762--
  +----- Nodularia       
--+                      
  |  +-- Cuneopsis_heudei **
  +--+                   
     +-- Schistodesmus   

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015264--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus   

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013022--
  +----- Nodularia       
--+                      
  |  +-- Tchangsinaia    
  +--+                   
     +-- Cuneopsis_heudei **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017364--
  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Tchangsinaia    

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012183--
  +----- Cuneopsis_celtiformis **
  |                           
--+----- Nodularia            
  |                           
  |  +-- Inversiunio          
  +--+                        
     +-- Schistodesmus        

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011372--
  +----- Inversiunio          
--+                           
  |  +-- Cuneopsis_celtiformis **
  +--+                        
     +-- Nodularia            
 
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