MUSSELpdb | valid gen. Cuneopsis cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Cuneopsis Simpson, 1900: cladomics
7 Recent valid species (10 total species)

Cladomics Summary

number of species = 7 (10 total)
analyzed in 81 cladograms in 49 publications
maximum number of terminals in an analysis = 7 (mean = 2.77, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 57
maximum number of included terminals = 7, (mean = 3.51 median = 3)
proportion of cladograms that support basic monophyly = 0.82

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	7	6	8	625	25	6	0

                          +-- Schistodesmus          
  +-----------------------+                          
  |                       +-- Nodularia              
  |                                                  
--+                       +-- Tchangsinaia           
  |  +--------------------+                          
  |  |                    +-- Inversiunio            
  +--+                                               
     |  +-------------------- Pseudocuneopsis        
     |  |                                            
     +--+  +----------------- Diaurora               
        |  |                                         
        +--+  +-------------- Cuneopsis_rufescens **    
           |  |                                      
           |  |        +----- Cuneopsis_celtiformis **  
           +--+  +-----+                             
              |  |     |  +-- Cuneopsis_kiangsiensis ** 
              |  |     +--+                          
              +--+        +-- Cuneopsis_celtiosimilis **
                 |                                   
                 |  +-------- Cuneopsis_szechenyii **   
                 +--+                                
                    |  +----- Cuneopsis_demangei__2 **  
                    +--+                             
                       |  +-- Cuneopsis_demangei__1 **  
                       +--+                          
                          +-- Cuneopsis_heudei **

Hou et al., 2025 [Publ 4432], fig. 3

tT	bi	riS	oT	roT	roS	iD
7	7	5	8	4	2	2	1

  +----------- Cuneopsis_rufescens **    
  |                                   
  |     +----- Cuneopsis_celtiformis **  
--+  +--+                             
  |  |  |  +-- Cuneopsis_celtiosimilis **
  |  |  +--+                          
  +--+     +-- Cuneopsis_kiangsiensis ** 
     |                                
     |  +----- Cuneopsis_szechenyii **   
     |  |                             
     +--+  +-- Cuneopsis_heudei **       
        |  |                          
        +--+-- Cuneopsis_demangei__1 **  
           |                          
           +-- Cuneopsis_demangei__2 **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	133	14	4	1

              +----- Pseudocuneopsis        
  +-----------+                             
  |           |  +-- Schistodesmus          
  |           +--+                          
--+              +-- Nodularia              
  |                                         
  |  +-------------- Tchangsinaia           
  |  |                                      
  +--+  +----------- Cuneopsis_rufescens **    
     |  |                                   
     |  |     +----- Cuneopsis_celtiformis **  
     +--+  +--+                             
        |  |  |  +-- Cuneopsis_kiangsiensis ** 
        |  |  +--+                          
        +--+     +-- Cuneopsis_celtiosimilis **
           |                                
           |  +----- Cuneopsis_szechenyii **   
           +--+                             
              |  +-- Cuneopsis_heudei **       
              +--+                          
                 +-- Cuneopsis_demangei **

Hou et al., 2025 [Publ 4432], fig. 4

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	21	18	4	1

  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           |  |     +----- Cuneopsis_szechenyii **   
           +--+  +--+                             
              |  |  |  +-- Cuneopsis_demangei **     
              |  |  +--+                          
              +--+     +-- Cuneopsis_heudei **       
                 |                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_kiangsiensis ** 
                    +--+                          
                       +-- Cuneopsis_celtiosimilis **

Wang et al., 2025 [Publ 4488], fig. 3B

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	58	17	4	2

  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           +--+        +-- Cuneopsis_heudei **       
              |  +-----+                          
              |  |     +-- Cuneopsis_szechenyii **   
              +--+                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_celtiosimilis **
                    +--+                          
                       +-- Cuneopsis_kiangsiensis **

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	89	14	4	2

  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           +--+        +-- Cuneopsis_heudei **       
              |  +-----+                          
              |  |     +-- Cuneopsis_szechenyii **   
              +--+                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_celtiosimilis **
                    +--+                          
                       +-- Cuneopsis_kiangsiensis **

Hou et al., 2025 [Publ 4401], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	22	19	4	2

  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+  +----------------- Schistodesmus         
     |  |                                        
     +--+  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_demangei **    
                    +--+                         
                       +-- Cuneopsis_heudei **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	79	13	5	2

  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+                 +-- Tongchuania           
     |  +--------------+                         
     |  |              +-- Schistodesmus         
     +--+                                        
        |  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_heudei **      
                    +--+                         
                       +-- Cuneopsis_demangei **

Hou et al., 2024 [Publ 4400], fig. 1

tT	bi	riS	oT	roT	roS	iD
6	13	11	6	9	6	2	9

           +----- Cuneopsis_rufescens **   
  +--------+                            
  |        |  +-- Middendorffinaia      
  |        +--+                         
--+           +-- Pseudocuneopsis       
  |                                     
  |  +----------- Tchangsinaia          
  |  |                                  
  +--+        +-- Cuneopsis_kiangsiensis **
     |  +-----+                         
     |  |     +-- Cuneopsis_celtiformis ** 
     +--+                               
        |  +----- Cuneopsis_szechenyii **  
        +--+                            
           |  +-- Cuneopsis_heudei **      
           +--+                         
              +-- Cuneopsis_demangei **

Hou et al., 2024 [Publ 4400], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	69	18	4	2

  +----------------------- Pseudocuneopsis       
  |                                              
--+  +-------------------- Nodularia             
  |  |                                           
  +--+  +----------------- Schistodesmus         
     |  |                                        
     +--+  +-------------- Tchangsinaia          
        |  |                                     
        +--+  +----------- Cuneopsis_rufescens **   
           |  |                                  
           +--+        +-- Cuneopsis_celtiformis ** 
              |  +-----+                         
              |  |     +-- Cuneopsis_kiangsiensis **
              +--+                               
                 |  +----- Cuneopsis_szechenyii **  
                 +--+                            
                    |  +-- Cuneopsis_demangei **    
                    +--+                         
                       +-- Cuneopsis_heudei **

Wu et al., 2026 [Publ 4825], fig. 6B

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	59	14	4	2

  +----------------------- Pseudocuneopsis        
  |                                               
--+  +-------------------- Nodularia              
  |  |                                            
  +--+  +----------------- Schistodesmus          
     |  |                                         
     +--+  +-------------- Tchangsinaia           
        |  |                                      
        +--+  +----------- Cuneopsis_rufescens **    
           |  |                                   
           +--+        +-- Cuneopsis_heudei **       
              |  +-----+                          
              |  |     +-- Cuneopsis_szechenyii **   
              +--+                                
                 |  +----- Cuneopsis_celtiformis **  
                 +--+                             
                    |  +-- Cuneopsis_celtiosimilis **
                    +--+                          
                       +-- Cuneopsis_kiangsiensis **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	7	6	5	3

  +----------------------- Nodularia            
  |                                             
--+  +-------------------- Pseudocuneopsis      
  |  |                                          
  +--+  +----------------- Schistodesmus        
     |  |                                       
     +--+              +-- Tchangsinaia         
        |  +-----------+                        
        |  |           +-- Inversiunio          
        +--+                                    
           |  +----------- Cuneopsis_rufescens **  
           +--+                                 
              |  +-------- Cuneopsis_celtiformis **
              +--+                              
                 |  +----- Cuneopsis_szechenyii ** 
                 +--+                           
                    |  +-- Cuneopsis_demangei **   
                    +--+                        
                       +-- Cuneopsis_heudei **

Wu et al., 2024 [Publ 4379], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	49	15	4	4

  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_demangei **   
                 +--+                        
                    +-- Cuneopsis_heudei **

Hou et al., 2025 [Publ 4445], fig. 5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	48	18	4	4

  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_demangei **   
                 +--+                        
                    +-- Cuneopsis_heudei **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	118	14	4	4

  +-------------- Nodularia            
  |                                    
--+  +----------- Schistodesmus        
  |  |                                 
  |  |        +-- Tchangsinaia         
  +--+--------+                        
     |        +-- Inversiunio          
     |                                 
     |  +-------- Cuneopsis_rufescens **  
     +--+                              
        |  +----- Cuneopsis_celtiformis **
        +--+                           
           |  +-- Cuneopsis_szechenyii ** 
           +--+                        
              +-- Cuneopsis_heudei **

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	5	3	4	42	8	3	5

  +----------------- Schistodesmus        
--+                                       
  |  +-------------- Nodularia            
  +--+                                    
     |  +----------- Cuneopsis_rufescens **  
     +--+                                 
        |  +-------- Tchangsinaia         
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	87	8	4	4

  +----------------- Nodularia            
  |                                       
--+----------------- Pseudocuneopsis      
  |                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	69	8	4	4

  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	87	8	4	4

  +----------------- Nodularia            
  |                                       
--+----------------- Pseudocuneopsis      
  |                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	91	8	4	4

                 +-- Nodularia            
  +--------------+                        
  |              +-- Pseudocuneopsis      
--+                                       
  |  +-------------- Tchangsinaia         
  +--+                                    
     |  +----------- Schistodesmus        
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_demangei **   
              +--+                        
                 +-- Cuneopsis_heudei **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	91	8	4	4

  +-------------------- Pseudocuneopsis          
  |                                              
--+  +----------------- Nodularia                
  |  |                                           
  |  |           +----- Tchangsinaia             
  +--+  +--------+                               
     |  |        |  +-- Schistodesmus_xinyuensis 
     |  |        +--+                            
     +--+           +-- Schistodesmus_spinosus   
        |                                        
        |  +----------- Schistodesmus_lampreyanus
        +--+                                     
           |  +-------- Cuneopsis_rufescens **      
           +--+                                  
              |  +----- Cuneopsis_celtiformis **    
              +--+                               
                 |  +-- Cuneopsis_demangei **       
                 +--+                            
                    +-- Cuneopsis_heudei **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	195	7	3	4

  +----------------- Nodularia            
--+                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	197	8	3	4

  +----------------- Schistodesmus        
--+                                       
  |  +-------------- Nodularia            
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_szechenyii ** 
              +--+                        
                 +-- Cuneopsis_heudei **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	49	10	4	4

  +-------------------- Pseudocuneopsis      
--+                                          
  |  +----------------- Nodularia            
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Tchangsinaia         
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	43	8	3	4

  +----------------- Nodularia            
--+                                       
  |  +-------------- Tchangsinaia         
  +--+                                    
     |  +----------- Schistodesmus        
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_heudei **     
              +--+                        
                 +-- Cuneopsis_szechenyii **

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	34	11	5	4

  +-------------------- Pseudocuneopsis      
  |                                          
--+  +----------------- Nodularia            
  |  |                                       
  +--+  +-------------- Schistodesmus        
     |  |                                    
     +--+           +-- Tchangsinaia         
        |  +--------+                        
        |  |        +-- Inversiunio          
        +--+                                 
           |  +-------- Cuneopsis_rufescens **  
           +--+                              
              |  +----- Cuneopsis_celtiformis **
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_demangei **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	83	12	4	4

                 +-- Nodularia            
  +--------------+                        
  |              +-- Pseudocuneopsis      
--+                                       
  |  +-------------- Schistodesmus        
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Cuneopsis_rufescens **  
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Cuneopsis_heudei **     
              +--+                        
                 +-- Cuneopsis_demangei **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
4	8	3	4	197	8	3	8

  +----- Cuneopsis_celtiformis **
  |                           
  +----- Tchangsinaia         
  |                           
--+----- Cuneopsis_rufescens **  
  |                           
  +----- Nodularia            
  |                           
  |  +-- Cuneopsis_szechenyii ** 
  +--+                        
     +-- Cuneopsis_heudei **

Ouyang et al., 2015 [Publ 4493], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	6	4	3	28	4	3	8

        +-- Tchangsinaia         
  +-----+                        
  |     +-- Cuneopsis_rufescens **  
--+                              
  |  +----- Nodularia            
  +--+                           
     |  +-- Cuneopsis_szechenyii ** 
     +--+                        
        +-- Cuneopsis_celtiformis **

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	50	4	3	5

           +-- Nodularia            
  +--------+                        
  |        +-- Schistodesmus        
--+                                 
  |  +-------- Tchangsinaia         
  +--+                              
     |  +----- Cuneopsis_rufescens **  
     +--+                           
        |  +-- Cuneopsis_celtiformis **
        +--+                        
           +-- Cuneopsis_szechenyii **

Ouyang et al., 2015 [Publ 4493], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	5	6	6	28	4	3	11

  +----------------- Nodularia               
  |                                          
  |           +----- Cuneopsis_rufescens__2 **  
--+  +--------+                              
  |  |        |  +-- Schistodesmus           
  |  |        +--+                           
  +--+           +-- Cuneopsis_celtiformis__2 **
     |                                       
     |  +----------- Tchangsinaia            
     +--+                                    
        |  +-------- Cuneopsis_celtiformis__1 **
        +--+                                 
           |  +----- Cuneopsis_szechenyii__2 ** 
           +--+                              
              |  +-- Cuneopsis_szechenyii__1 ** 
              +--+                           
                 +-- Cuneopsis_rufescens__1 **

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	11	6	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii **

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	12	6	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	74	4	3	7

  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Cuneopsis_szechenyii **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	12	6	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	74	4	3	7

  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Cuneopsis_szechenyii **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens **

Wu et al., 2022 [Publ 4238], fig. 2e

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	16	7	4	--

  +-------------- Pseudocuneopsis      
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Tchangsinaia         
     +--+                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Cuneopsis_celtiformis **
           +--+                        
              +-- Cuneopsis_heudei **

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	11	6	--

              +-- Tchangsinaia         
  +-----------+                        
  |           +-- Inversiunio          
--+                                    
  |  +----------- Pseudocuneopsis      
  |  |                                 
  +--+        +-- Cuneopsis_celtiformis **
     |  +-----+                        
     |  |     +-- Cuneopsis_szechenyii ** 
     +--+                              
        |  +----- Nodularia            
        +--+                           
           |  +-- Diaurora             
           +--+                        
              +-- Schistodesmus

Wu et al., 2025 [Publ 4442], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	33	0	0	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_heudei **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	79	4	3	--

  +----------- Nodularia           
--+                                
  |  +-------- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis_szechenyii **
        +--+                       
           +-- Cuneopsis_rufescens **

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	6	0	2	12	5	4	9

  +-------------- Tchangsinaia       
--+                                  
  |  +----------- Cuneopsis_heudei **   
  +--+                               
     |  +-------- Nodularia          
     +--+                            
        |  +----- Pseudocuneopsis    
        +--+                         
           |  +-- Cuneopsis_rufescens **
           +--+                      
              +-- Schistodesmus

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	7	4	--

  +-------------- Pseudocuneopsis      
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Tchangsinaia         
     +--+                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Cuneopsis_celtiformis **
           +--+                        
              +-- Cuneopsis_heudei **

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	11	6	--

           +-- Tchangsinaia         
  +--------+                        
  |        +-- Inversiunio          
  |                                 
--+     +----- Pseudocuneopsis      
  |  +--+                           
  |  |  |  +-- Cuneopsis_celtiformis **
  |  |  +--+                        
  +--+     +-- Cuneopsis_szechenyii ** 
     |                              
     |  +----- Nodularia            
     +--+                           
        |  +-- Diaurora             
        +--+                        
           +-- Schistodesmus

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	137	4	3	--

  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_rufescens ** 
  |  +--+                       
  |  |  +-- Cuneopsis_szechenyii **
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	22	9	5	--

           +-- Cuneopsis_szechenyii **
  +--------+                       
  |        +-- Cuneopsis_rufescens ** 
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	54	2	2	--

  +-------- Nodularia           
--+                             
  |  +----- Tchangsinaia        
  +--+                          
     |  +-- Cuneopsis_rufescens ** 
     +--+                       
        +-- Cuneopsis_szechenyii **

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	22	9	5	--

     +-- Cuneopsis_rufescens ** 
  +--+                       
  |  +-- Cuneopsis_szechenyii **
  |                          
  |  +-- Nodularia           
--+--+                       
  |  +-- Schistodesmus       
  |                          
  |  +-- Tchangsinaia        
  |  |                       
  +--+-- Inversiunio         
     |                       
     +-- Pseudocuneopsis

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	10	0	2	123	4	3	13

  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Scriptolamprotula    
  |                        
  +-- Cuneopsis_szechenyii ** 
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens **  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus

Choi et al., 2020 [Publ 4169], fig. 2a

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	10	9	3	--

  +-- Cuneopsis_heudei **   
--+                      
  +-- Cuneopsis_rufescens **

Hou et al., 2025 [Publ 4445], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	5	5	1	--

  +----- Schistodesmus        
--+                           
  |  +-- Cuneopsis_heudei **     
  +--+                        
     +-- Cuneopsis_celtiformis **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	4	3	--

  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_szechenyii **
  |  +--+                       
  |  |  +-- Cuneopsis_rufescens ** 
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	4	3	--

  +-------- Nodularia           
  |                             
--+     +-- Cuneopsis_szechenyii **
  |  +--+                       
  |  |  +-- Cuneopsis_rufescens ** 
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus

Liu et al., 2017 [Publ 3939], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	5	2	--

  +----- Schistodesmus       
  |                          
--+----- Nodularia           
  |                          
  |  +-- Cuneopsis_szechenyii **
  +--+                       
     +-- Cuneopsis_rufescens **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	42	11	6	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Cuneopsis_szechenyii **

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	8	5	--

           +-- Cuneopsis_szechenyii **
  +--------+                       
  |        +-- Cuneopsis_rufescens ** 
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	39	9	4	--

  +-------------- Pseudocuneopsis      
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Schistodesmus        
     +--+                              
        |  +----- Tchangsinaia         
        +--+                           
           |  +-- Cuneopsis_heudei **     
           +--+                        
              +-- Cuneopsis_celtiformis **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	137	4	3	7

  +----- Cuneopsis_szechenyii **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Cuneopsis_rufescens **

Ouyang et al., 2011 [Publ 3408], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	1	1	--

  +-- Nodularia       
--+                   
  +-- Cuneopsis_heudei **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	6	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Tchangsinaia

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	8	3	--

  +----- Cuneopsis_celtiformis **
  |                           
--+----- Nodularia            
  |                           
  |  +-- Inversiunio          
  +--+                        
     +-- Schistodesmus

Chen et al., 2026 [Publ 4805], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Cuneopsis_celtiformis **

Wang et al., 2018 [Publ 3846], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +----- Nodularia       
--+                      
  |  +-- Tchangsinaia    
  +--+                   
     +-- Cuneopsis_heudei **

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	5	3	--

  +-- Cuneopsis_celtiformis **
--+                        
  +-- Nodularia

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	6	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	4	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	4	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus

Zhou et al., 2007 [Publ 2658], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	1	1	--

  +-- Nodularia            
--+                        
  +-- Cuneopsis_celtiformis **

Wu et al., 2024 [Publ 4379], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	12	3	--

  +----- Nodularia            
  |                           
--+  +-- Cuneopsis_celtiformis **
  |  |                        
  +--+-- Tchangsinaia         
     |                        
     +-- Pseudocuneopsis

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	4	4	--

        +-- Nodularia            
  +-----+                        
  |     +-- Pseudocuneopsis      
--+                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis_celtiformis **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	6	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Tchangsinaia    
        +--+                   
           +-- Cuneopsis_heudei **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	8	3	--

  +-------- Nodularia            
--+                              
  |  +----- Cuneopsis_celtiformis **
  +--+                           
     |  +-- Inversiunio          
     +--+                        
        +-- Schistodesmus

Choi et al., 2020 [Publ 4169], fig. 3a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	4	1	--

-- Cuneopsis_heudei **

Wu et al., 2020 [Publ 4147], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	5	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Schistodesmus   
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Tchangsinaia

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	3	3	--

  +-------- Nodularia       
--+                         
  |  +----- Tchangsinaia    
  +--+                      
     |  +-- Cuneopsis_heudei **
     +--+                   
        +-- Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	7	2	--

  +----- Inversiunio          
--+                           
  |  +-- Cuneopsis_celtiformis **
  +--+                        
     +-- Nodularia

Kim et al., 2021 [Publ 4197], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	6	2	--

  +----- Nodularia       
--+                      
  |  +-- Cuneopsis_heudei **
  +--+                   
     +-- Schistodesmus

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	4	4	--

        +-- Pseudocuneopsis      
  +-----+                        
  |     +-- Nodularia            
--+                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Cuneopsis_celtiformis **
     +--+                        
        +-- Tchangsinaia

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	6	4	--

  +----------- Pseudocuneopsis 
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Tchangsinaia    
     +--+                      
        |  +-- Cuneopsis_heudei **
        +--+                   
           +-- Schistodesmus

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	4	4	--

        +-- Pseudocuneopsis      
  +-----+                        
  |     +-- Schistodesmus        
--+                              
  |  +----- Nodularia            
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis_celtiformis **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	7	4	--

  +-------- Pseudocuneopsis 
  |                         
--+     +-- Cuneopsis_heudei **
  |  +--+                   
  |  |  +-- Schistodesmus   
  +--+                      
     |  +-- Tchangsinaia    
     +--+                   
        +-- Nodularia

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	5	4	--

        +-- Cuneopsis_heudei **
  +-----+                   
  |     +-- Schistodesmus   
--+                         
  |  +----- Pseudocuneopsis 
  +--+                      
     |  +-- Nodularia       
     +--+                   
        +-- Tchangsinaia

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