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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio mardinensis Lea, 1865: synonymy
     5 specimens + 17 Genbank vouchers

Unio kullethensis Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 16: 285.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 251 [11], fig. 29, fig. 65.
Type(s): HOLOTYPE USNM_85663, near Mardin, Asiatic Turkey.

Unio (sec. Lymnium) durieui var. kullethensisSimpson (1900), Simpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio mardinensis Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 16: 286.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 252 [12], pl. 30, fig. 66.
Type(s): HOLOTYPE USNM_85664, R. Tigris, near Mardin, Asiatic Turkey.

Unio (sec. Lymnium) durieuiSimpson (1900), Simpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024), Hou et al. (2024)

Unio orphaensis Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 16: 285.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 250 [10], pl. 29, fig. 64.
Type(s): HOLOTYPE USNM_85603, Tigris R., Pashalic of Orpha, Asiatic Turkey.

Unio (sec. Lymnium) durieuiSimpson (1900), Simpson (1914)
Unio (Limnium) [sic] orphænsisPallary (1939)
Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio rothi var. komarowi O. Boettger, 1880
O. Boettger, 1880. Jb. Dtsch. Malak. Ges. 7: 160, pl. 5, figs. 1a-b.
(intro'd as intra-sp. of Unio rothi Bourguignat, 1863)

Psilunio rothi var. komarovi [sic]Zhadin (1938)
Psilunio littoralis komarowiHaas (1940)
Unio mingrelicusZhadin (1965)
Potomida littoralis komarowiHaas (1969), Falkner (1994)
Potomida komarovi [sic]Zatravkin (1983), Zatravkin (1987)
Potomida littoralisGraf (2007)
Potomida komarowiKantor et al. (2010), Vinarski & Kantor (2016)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio araxenus Drouët, 1881
Drouët, 1881. Unionidae Russie: 18.

Unio (s.s.) sieversi var. araxenusZhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969), Graf (2007)
Unio sieversi var. araxenusZhadin (1965)
Crassiana crassa sieversiAkromovski (1976), Kantor & Sysoev (2005)
Eolymnium (Shadininaia) araxenumZatravkin (1983), Zatravkin (1987)
Eolymnium araxenumKantor et al. (2010)
Unio (Eolymnium) araxenusVinarski & Kantor (2016)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio kallethensis Paetel, 1890
‘Lea’ Paetel, 1890. Cat. Conch.-Samml. 3: 159.
(error for Unio kullethensis Lea, 1865)

Unio (sec. Lymnium) durieui var. kullethensisSimpson (1900), Simpson (1914)

Unio (sec. Lymnium) pseudonymus Simpson, 1900
‘Bourg.’ Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: 22, pl. 41, fig. 255 [as ‘U. hueti’].
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 694.
Simpson, 1914. Descr. Cat. Naiades: 560.

Unio (sec. Lymnium) pseudonymusSimpson (1900), Simpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio armeniacus Kobelt, 1912
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 29, pl. 493, fig. 2631.

Psilunio rothi var. armeniacusZhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio mingrelicusZhadin (1965)
Potomida armeniacaZatravkin (1983), Kantor et al. (2010), Vinarski & Kantor (2016)
Potomida armenica [sic]Zatravkin (1987)
Potomida littoralisGraf (2007), IUCN (2014)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio modiola Preston, 1912
Preston, 1912. Rec. Ind. Mus. 7: 286, pl. 8, figs. 1-2.

Unio modiolaPreston (1912)
Unio (sec. Lymnium) modiolaSimpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Falkner (1994)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio orfaensis Kobelt, 1912
‘Lea’ Kobelt, 1912. Icon. Land- und Süssw.-Moll. 19: 37, pl. 530, figs. 2734-2735.
(error for Unio orphaensis Lea, 1865)

Unio (sec. Lymnium) durieuiSimpson (1914)

Unio bitlisensis Kobelt, 1913
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 35, pl. 538, fig. 2731.

Simpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio ciconius Kobelt, 1913
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 11, pl. 517, fig. 2697.

Unio (sec. Lymnium) tigridisSimpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021), Bogan et al. (2021)
Unio crassusJihad (2023)
Unio mardinensisLopes-Lima et al. (2024)

Unio diarbekirianus Kobelt, 1913
‘Bourguignat mss’ Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 15, pl. 519, fig. 2702.

Simpson (1914)
Unio (Limnium) [sic] diabekirianus [sic]Pallary (1939)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021), Bogan et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio medicus Kobelt, 1913
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 17, pl. 520, fig. 2707.

Unio (sec. Lymnium) tigridisSimpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio specialis Kobelt, 1913
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 8, pl. 515, fig. 2694.

Simpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio subbatavus Kobelt, 1913
Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 317. [nomen nudum]
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 48, pl. 534, fig. 2749.

Unio crassus gontieriHaas (1940), Haas (1969)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio (orfaensis var. ?) ehrmanni Kobelt, 1913
Kobelt, 1913. Icon. Land- und Süssw.-Moll. 19: 38, pl. 530, fig. 2736.
(intro'd as intra-sp. of Unio orphaensis Lea, 1865)

Unio (sec. Lymnium) durieuiSimpson (1914)
Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Unio bruguierianusLopes-Lima et al. (2021)
Unio crassusJihad (2023)
Unio mardinensisLopes-Lima et al. (2024)

Unio thospiensis Kobelt, 1915
Kobelt, 1915. Icon. Land- und Süssw.-Moll. 20: 28, pl. 553, figs. 2810-2811.

Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Unio (Rhombunio) wenzii Kobelt, 1915
Kobelt, 1915. Icon. Land- und Süssw.-Moll. 20: 29, pl. 553, figs. 2812-2814.

Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Unio bruguierianusLopes-Lima et al. (2021)
Unio mardinensisLopes-Lima et al. (2024)

Psilunio rothi var. komarovi Zhadin, 1938
‘Boettg.’ Zhadin, 1938. Faune USSR Mollusques 4(1): 102.
(error for Unio rothi var. komarowi O. Boettger, 1880)

Potomida komarovi [sic]Zatravkin (1987)

Unio bruguierianus Lopes-Lima et al., 2021
‘Bourguignat’ of Authors.
Lopes-Lima et al., 2021. MPE 163: Table S1. [in part]
(misidentified reference to Unio bruguierianus Bourguignat, 1853)

Unio mardinensisLopes-Lima et al. (2024)

Unio kullenthenensis Paetel, 1890
‘Lea’ Paetel, 1890. Cat. Conch.-Samml. 3: 156.
(error for Unio kullethensis Lea, 1865)

Unio (sec. Lymnium) durieui var. kullethensisSimpson (1900), Simpson (1914)

Unio diarbekerianus Simpson, 1914
Simpson, 1914. Descr. Cat. Naiades: 732.
(error for Unio diarbekirianus Kobelt, 1913)

Unio (Limnium) diabekirianus Pallary, 1939
‘Kobelt’ Pallary, 1939. Mém. Inst. Égypte 39: 120.
(error for Unio diarbekirianus Kobelt, 1913)

Unio diarbekrianus Haas, 1940
‘Kobelt’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 132.
(error for Unio diarbekirianus Kobelt, 1913)

Potomida armenica Zatravkin, 1987
‘Kobelt’ Zatravkin, 1987. Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 48.
(error for Unio armeniacus Kobelt, 1912)

Unio arazenus Zatravkin, 1987
‘Drouet’ Zatravkin, 1987. Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 49.
(error for Unio araxenus Drouët, 1881)

Taxonomic History (since 1900)

Unio (sec. Lymnium) durieui var. kullethensisSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) durieuiSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) pseudonymusSimpson (1900), Simpson (1914)
Unio modiolaPreston (1912)
Unio (sec. Lymnium) modiolaSimpson (1914)
Simpson (1914), Falkner (1994)
Unio (sec. Lymnium) tigridisSimpson (1914)
Psilunio rothi var. komarovi [sic]Zhadin (1938)
Unio (s.s.) sieversi var. araxenusZhadin (1938)
Psilunio rothi var. armeniacusZhadin (1938)
Unio (Limnium) [sic] orphænsisPallary (1939)
Unio (Limnium) [sic] diabekirianus [sic]Pallary (1939)
Unio crassus bruguierianusHaas (1940), Haas (1969), Schütt (1983)
Psilunio littoralis komarowiHaas (1940)
Unio crassus gontieriHaas (1940), Haas (1969), Graf (2007)
Unio mingrelicusZhadin (1965)
Unio sieversi var. araxenusZhadin (1965)
Potomida littoralis komarowiHaas (1969), Falkner (1994)
Crassiana crassa sieversiAkromovski (1976), Kantor & Sysoev (2005)
Potomida komarovi [sic]Zatravkin (1983), Zatravkin (1987)
Eolymnium (Shadininaia) araxenumZatravkin (1983), Zatravkin (1987)
Potomida armeniacaZatravkin (1983), Kantor et al. (2010), Vinarski & Kantor (2016)
Potomida armenica [sic]Zatravkin (1987)
Potomida littoralisGraf (2007), IUCN (2014)
Potomida komarowiKantor et al. (2010), Vinarski & Kantor (2016)
Eolymnium araxenumKantor et al. (2010)
Unio (Eolymnium) araxenusVinarski & Kantor (2016)
Unio bruguierianusLopes-Lima et al. (2021), Bogan et al. (2021)
Unio crassusJihad (2023)
Unio mardinensisLopes-Lima et al. (2024), Hou et al. (2024)

 
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