The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Pleurobemini genus Elliptio Rafinesque, 1819: cladomics
30 Recent valid species (45 total species) Cladomics Summarynumber of species = 30 (45 total)
analyzed in 76 cladograms in 47 publications
maximum number of terminals in an analysis =
19 (mean = 1.84, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 20
maximum number of included terminals = 19, (mean =
4.2 median = 2)
proportion of cladograms that support basic monophyly
= 0.85
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 57 | 49 | 20 | 85 | 39 | 8 | 0 | +----------------------------------------- Elliptoideus
|
| +-- Elliptio_lanceolata **
| +-----------------------------------+
| | +-- Parvaspina
| |
| | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
--+ | |
| | +----------------------------------- Hemistena
| | |
| | | +-- Elliptio_complanata **
| | | |
| | | +-- Elliptio_hopetonensis **
| | | |
| | | +-- Elliptio_producta__1 **
| | | |
| | | +-- Elliptio_shepardiana **
+--+ | |
| | +-- Elliptio_angustata **
| | |
| | +-- Elliptio_arca **
| | |
| | +-- Elliptio_arctata **
| | |
| | +-- Elliptio_jayensis **
| | |
| | +-- Elliptio_crassidens **
| | |
| +--------------------------------+-- Elliptio_dariensis **
| | |
| | +-- Elliptio_fisheriana **
| | |
| | +-- Elliptio_folliculata **
| | |
+--+ +-- Elliptio_fumata **
| |
| +-- Elliptio_icterina **
| |
| +-- Elliptio_mcmichaeli **
| |
| +-- Elliptio_producta__2 **
| |
| +-- Elliptio_pullata **
| |
| +-- Elliptio_purpurella **
| |
| +-- Elliptio_congaraea **
|
| +-------------------------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
| | +--------------------------+
| | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Perkins et al., 2017 [Publ 3777], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 12 | 14 | 23 | 21 | 6 | 148 | +-------------------------------------- Pleurobema
|
--+ +----------------------------------- Pleuronaia
| |
+--+ +-------------------------------- Fusconaia
| |
+--+ +----------------------------- Parvaspina
| |
+--+ +-- Eurynia
| +-----------------------+
| | +-- Hemistena
+--+
| +----------------------- Elliptio_producta__2 **
| |
+--+ +-------------------- Elliptio_producta__1 **
| |
+--+ +----------------- Elliptio_spinosa **
| |
| | +-------------- Elliptio_icterina **
+--+ |
| | +-------- Elliptio_roanokensis **
| | +--+
+--+ | | +----- Elliptio_complanata **
| | +--+
| | | +-- Elliptio_crassidens **
| | +--+
+--+ +-- Elliptio_pullata **
|
| +-- Elliptio_congaraea **
| +-----+
| | +-- Elliptio_hopetonensis **
| |
+--+ +-- Elliptio_folliculata **
| +--+
| | +-- Elliptio_jayensis **
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_arctata **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 8 | 65 | 33 | 8 | 145 | +----------------- Elliptoideus
|
| +-- Eurynia
| +-----------+
| | +-- Parvaspina
--+ |
| | +-- Hemistena
| +-----------+
| | +-- Plethobasus
| |
| | +----- Pleuronaia
+--+--------+
| | +-- Fusconaia
| +--+
| +-- Pleurobema
|
| +----------- Elliptio_shepardiana **
| |
| | +----- Elliptio_crassidens **
+--+ +--+
| | | +-- Elliptio_mcmichaeli **
| | +--+
+--+ +-- Elliptio_pullata **
|
| +----- Elliptio_complanata **
| |
+--+ +-- Elliptio_purpurella **
| |
+--+-- Elliptio_arca **
|
+-- Elliptio_arctata **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 8 | 65 | 33 | 8 | 144 | +----------------------------------- Elliptoideus
|
| +-------------------------------- Pleurobema_stabile
| |
| | +-- Pleurobema_clava
--+ | +-----------+
| | | +-- Pleurobema_oviforme
| | |
| | +--------------+ +----------- Pleurobema_pyriforme
| | | | |
| | | | | +-- Pleurobema_athearni
+--+ | | | +--+
| | +--+ | +-- Pleurobema_georgianum
| | | +--+
| | | | | +-- Pleurobema_decisum
| | | | +--+
| | | | +-- Pleurobema_hanleyianum
| | +--+
| | | +----- Pleurobema_beadleianum
| | | |
+--+ | +----- Pleurobema_strodeanum
| +--+
| +----- Pleurobema_rubellum
| |
| | +-- Pleurobema_perovatum
| +--+
| +-- Pleurobema_taitianum
|
| +----- Pleurobema_riddellii
| +--------------------+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +----------------------- Fusconaia
| |
+--+ +-------------------- Pleuronaia
| |
| | +-- Eurynia
+--+ +--------------+
| | +-- Parvaspina
| |
+--+ +-- Hemistena
| +-----------+
| | +-- Plethobasus
| |
+--+ +----------- Elliptio_shepardiana **
| |
| | +-------- Elliptio_complanata **
+--+ |
| | +----- Elliptio_purpurella **
| +--+
| | | +-- Elliptio_arca **
+--+ +--+
| +-- Elliptio_arctata **
|
| +-- Elliptio_mcmichaeli **
| |
+-----+-- Elliptio_pullata **
|
+-- Elliptio_crassidens **
Perkins et al., 2017 [Publ 3777], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 20 | 7 | 4 | 20 | 18 | 6 | 145 | +----------------- Parvaspina
|
| +-------------- Elliptio_arca **
| |
--+ +-------------- Hemistena
| |
| | +-- Elliptio_arctata **
| +-----------+
| | +-- Elliptio_crassidens **
+--+
| +----------- Eurynia
| |
| | +-------- Pleurobema_oviforme
| | |
| | +-------- Pleurobema_clava
| | |
+--+ +-------- Pleurobema_pyriforme
| |
| +-------- Pleurobema_georgianum
| |
| +-------- Pleurobema_beadleianum
| |
+--+-------- Pleurobema_hanleyianum
|
+-------- Pleurobema_decisum
|
| +-- Pleurobema_sintoxia
+-----+
| +-- Pleurobema_rubellum
|
| +----- Elliptio_spinosa **
| |
+--+ +-- Fusconaia_escambia
| |
| +-- Fusconaia_flava
+--+
+-- Fusconaia_cerina
|
+-- Pleuronaia
Campbell et al., 2008 [Publ 2675], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 22 | 19 | 7 | 146 | +----- Elliptio_arca **
--+
| +-- Elliptio_crassidens **
+--+
+-- Elliptio_arctata **
Campbell et al., 2008 [Publ 2675], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 23 | 5 | 3 | 22 | 19 | 7 | 147 | +-- Elliptio_arctata **
+-----+
| +-- Elliptio_crassidens **
|
| +----- Pleurobema_decisum
| |
| +----- Pleurobema_rubellum
| |
| +----- Pleurobema_georgianum__2
| |
| +----- Pleurobema_stabile
| |
| +----- Pleurobema_pyriforme
| |
--+ +----- Pleurobema_oviforme__2
| |
| +----- Pleurobema_hanleyianum__2
| |
| +----- Pleuronaia
| |
| +----- Pleurobema_georgianum__3
| |
| +----- Pleurobema_cordatum
| |
| +----- Pleurobema_beadleianum
| |
| +----- Hemistena
+--+
+----- Fusconaia_flava
|
+----- Fusconaia_cerina
|
+----- Eurynia
|
+----- Elliptio_arca **
|
+----- Cyrtonaias
|
| +-- Plethobasus
+--+
| +-- Parvaspina
|
| +-- Pleurobema_sintoxia
| |
+--+-- Pleurobema_oviforme__1
| |
| +-- Pleurobema_clava
|
| +-- Pleurobema_athearni
| |
+--+-- Pleurobema_georgianum__1
|
+-- Pleurobema_hanleyianum__1
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 0 | 0 | -- | +-- Elliptio_crassidens **
--+
+-- Elliptio_complanata **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 41 | 23 | 8 | -- | +-- Elliptio_arca **
+--------------------+
| +-- Elliptio_crassidens **
|
--+ +-------------------- Eurynia
| |
| | +----------------- Parvaspina
+--+ |
| | +-- Hemistena
| +--------------+
+--+ +-- Plethobasus
|
| +-------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
+--+ +--------+
| | +-- Fusconaia
| |
+--+ +-- Pleurobema_stabile
| +-----+
| | +-- Pleurobema_sintoxia
+--+
| +----- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
+--+
+-- Pleurobema_pyriforme
Burlakova et al., 2012 [Publ 3486], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 33 | 7 | -- | +----------------------- Hemistena
|
+----------------------- Pleurobema_stabile
|
| +-------------------- Eurynia
--+ |
| | +-------- Pleurobema_riddellii
| | |
| | +--------+ +----- Pleurobema_sintoxia__2
| | | | |
+--+ | +--+----- Pleurobema_sintoxia__3
| | |
| | | +-- Pleurobema_cordatum
| | +--+
| | +-- Pleurobema_sintoxia__1
| |
+--+ +-------------- Parvaspina
| |
| +-------------- Pleuronaia
| |
| +-------------- Fusconaia
| |
| | +----- Plethobasus
| +--------+
+--+ | +-- Elliptio_arca **
| +--+
| +-- Elliptio_crassidens **
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
| |
| | +-------- Pleurobema_decisum
+--+ |
| +-------- Pleurobema_pyriforme
| |
| +-------- Pleurobema_hanleyianum
+--+
+-------- Pleurobema_georgianum
|
| +----- Pleurobema_rubellum
| |
| +----- Pleurobema_beadleianum
+--+
+----- Pleurobema_strodeanum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 100 | 28 | 8 | -- | +-------------- Elliptoideus
|
--+ +----------- Hemistena
| |
| | +----- Plethobasus
+--+ +--+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
+--+
| +----- Pleuronaia
| |
| +----- Fusconaia
+--+
+----- Pleurobema
|
| +-- Eurynia
+--+
+-- Parvaspina
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 36 | 19 | 8 | -- | +-- Elliptio_fumata **
+--------------+
| +-- Elliptio_crassidens **
|
| +-------------- Parvaspina
--+ |
| +-------------- Plethobasus
| |
| | +-- Hemistena
+--+-----------+
| +-- Eurynia
|
| +----------- Fusconaia
| |
| | +----- Pleurobema_riddellii
+--+ +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +----- Pleuronaia
| |
+--+ +-- Pleurobema_pyriforme
| |
+--+-- Pleurobema_strodeanum
|
+-- Pleurobema_clava
Campbell et al., 2008 [Publ 2675], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 24 | 22 | 7 | -- | +----------------- Parvaspina
|
--+ +-------------- Hemistena
| |
| | +----- Plethobasus
+--+ +-----+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +-------- Fusconaia
+--+
| +----- Pleurobema
+--+
| +-- Eurynia
+--+
+-- Pleuronaia
Campbell & Lydeard, 2012 [Publ 3308], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 40 | 23 | 8 | -- | +-------- Elliptoideus
|
| +----- Hemistena
| |
--+ +----- Plethobasus
| |
| | +-- Elliptio_arca **
| +--+
| | +-- Elliptio_crassidens **
+--+
| +-- Eurynia
+--+
| +-- Parvaspina
|
| +-- Pleuronaia
| |
+--+-- Pleurobema
|
+-- Fusconaia
Burlakova et al., 2012 [Publ 3486], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 33 | 33 | 7 | -- | +----------------------------------- Parvaspina
|
| +-------------------------------- Pleurobema_plenum
--+ |
| | +-- Plethobasus
| | +-----+
| | | +-- Hemistena
+--+ +--------------------+
| | | +----- Eurynia
| | +--+
| | | +-- Elliptio_arca **
| | +--+
+--+ +-- Elliptio_crassidens **
|
| +-------------------------- Pleurobema_cordatum__2
| |
| | +----------------------- Fusconaia
+--+ |
| | +-------- Pleurobema_stabile
| | |
| | +-----------+ +----- Pleurobema_riddellii
+--+ | | |
| | +--+ +-- Pleurobema_cordatum__1
| | | |
| | | +-- Pleurobema_sintoxia__1
| | +--+
+--+ +-- Pleurobema_sintoxia__2
| |
| +-- Pleurobema_sintoxia__3
|
| +----------------- Pleuronaia
| |
| | +-- Pleurobema_georgianum
+--+ +-----------+
| | +-- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_decisum
| +--------+
| | +-- Pleurobema_hanleyianum
| |
+--+ +-- Pleurobema_clava
| +-----+
| | +-- Pleurobema_oviforme
| |
+--+ +----- Pleurobema_rubellum
| |
| +----- Pleurobema_beadleianum
+--+
+----- Pleurobema_strodeanum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Campbell & Lydeard, 2012 [Publ 3308], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 40 | 23 | 8 | -- | +----------------------- Elliptoideus
|
| +-------------------- Pleurobema_stabile
--+ |
| | +----- Pleurobema_pyriforme
| | +-----------+
+--+ | | +-- Pleurobema_clava
| | +--+
| | +-- Pleurobema_decisum
| |
+--+ +-- Pleurobema_cordatum
| +-----------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleuronaia
+--+ +-----+
| | | +-- Parvaspina
| | +--+
| | +-- Fusconaia
+--+
| +-- Eurynia
| +-----+
| | +-- Hemistena
+--+
| +----- Plethobasus
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_crassidens **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 100 | 28 | 8 | -- | +-------------------------------- Elliptoideus
|
| +----------------------------- Plethobasus
| |
--+ +----------------------------- Parvaspina
| |
| | +-- Elliptio_arca **
| +--------------------------+
+--+ +-- Elliptio_crassidens **
|
| +-------------------------- Eurynia
| |
| +-------------------------- Hemistena
| |
| | +----------------------- Fusconaia
+--+ |
| | +----- Pleurobema_sintoxia__2
| +-----------------+
| | | +-- Pleurobema_sintoxia__1
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +-------------------- Pleuronaia
| |
| | +-------- Pleurobema_rubellum
| | +--------+
+--+ | | +----- Pleurobema_strodeanum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
| | +-- Pleurobema_taitianum
+--+
| +-- Pleurobema_oviforme__1
| |
| +-----------+-- Pleurobema_oviforme__2
| | |
| | +-- Pleurobema_clava
+--+
| +----------- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_athearni
| +-----+
| | +-- Pleurobema_georgianum__2
+--+
| +----- Pleurobema_decisum
| |
+--+ +-- Pleurobema_hanleyianum__1
| |
+--+-- Pleurobema_hanleyianum__2
|
+-- Pleurobema_georgianum__1
Burlakova et al., 2012 [Publ 3486], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 33 | 7 | -- | +----------------------------------- Parvaspina
|
| +----- Plethobasus
--+ +--------------------------+
| | | +-- Elliptio_arca **
| | +--+
| | +-- Elliptio_crassidens **
+--+
| +----------------------------- Fusconaia
| |
| | +-- Eurynia
+--+ +-----------------------+
| | +-- Hemistena
| |
| | +----------- Pleuronaia
+--+ +-----------+
| | | +-------- Pleurobema_riddellii
| | +--+
| | | +----- Pleurobema_sintoxia__2
| | +--+
+--+ | +-- Pleurobema_sintoxia__1
| +--+
| +-- Pleurobema_cordatum
|
| +-------------------- Pleurobema_stabile
| |
+--+ +----------------- Pleurobema_pyriforme
| |
| | +----- Pleurobema_georgianum
+--+ +--------+
| | | +-- Pleurobema_hanleyianum
| | +--+
| | +-- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_beadleianum
| |
+--+ +-- Pleurobema_rubellum
| +--+
| | +-- Pleurobema_strodeanum
+--+
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Campbell et al., 2008 [Publ 2675], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 24 | 22 | 7 | -- | +-- Pleurobema_cordatum
|
+-----------+-- Pleurobema_sintoxia__1
| |
| +-- Pleurobema_sintoxia__2
+-----+
| | +----------- Fusconaia
| | |
| +--+ +-------- Pleuronaia
| | |
| | | +----- Eurynia
| +--+ |
| | +----- Hemistena
| | |
| +--+----- Plethobasus
--+ |
| +----- Parvaspina
| |
| | +-- Elliptio_arca **
| +--+
| +-- Elliptio_crassidens **
|
| +----------------- Pleurobema_stabile
| |
| | +-------------- Pleurobema_pyriforme
| | |
+--+ | +-- Pleurobema_georgianum__2
| +-----------+
| | +-- Pleurobema_athearni
| |
| | +----- Pleurobema_decisum
+--+--------+
| | +-- Pleurobema_hanleyianum
| +--+
| +-- Pleurobema_georgianum__1
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_rubellum
| |
+--+-------- Pleurobema_strodeanum
|
| +----- Pleurobema_beadleianum
+--+
| +-- Pleurobema_taitianum
+--+
+-- Pleurobema_perovatum
Campbell & Lydeard, 2012 [Publ 3308], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 40 | 23 | 8 | -- | +----------------------------- Elliptoideus
|
--+ +-------------------------- Pleurobema_stabile
| |
+--+ +----------------------- Hemistena
| |
+--+ +-------------------- Eurynia
| |
| | +-- Pleurobema_cordatum
+--+ +--------------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleurobema_clava
+--+ +--------+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_pyriforme
+--+
| +-- Parvaspina
| +--------+
| | +-- Pleuronaia
+--+
| +-------- Fusconaia
+--+
| +----- Plethobasus
+--+
| +-- Elliptio_arca **
+--+
+-- Elliptio_crassidens **
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 84 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptio_crassidens **
Metzger et al., 2018 [Publ 3952], fig. 1C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Elliptio_complanata **
Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Elliptio_complanata **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 124 | 2 | 2 | -- | -- Elliptio_complanata **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- | -- Elliptio_complanata **
Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 3 | 3 | -- | +-- Fusconaia
+--+
| +-- Pleurobema
--+
| +-- Eurynia
+--+
+-- Elliptio_crassidens **
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 4 | -- | +-------------- Elliptoideus
--+
| +----------- Elliptio_fumata **
+--+
| +-------- Pleuronaia
+--+
| +----- Pleurobema_riddellii
+--+
| +-- Fusconaia
+--+
+-- Pleurobema_strodeanum
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptio_complanata **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | -- Elliptio_complanata **
Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 1 | 1 | -- | +-- Elliptio_crassidens **
--+
+-- Pleurobema
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 4 | 4 | -- | +-------- Elliptoideus
|
--+ +-- Elliptio_complanata **
| +--+
| | +-- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Elliptoideus
--+
| +-- Elliptio_complanata **
+--+
+-- Pleurobema
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 0 | 0 | -- | -- Elliptio_complanata **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 4 | 4 | -- | +----------- Elliptoideus
--+
| +-------- Elliptio_complanata **
+--+
| +----- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Kandl et al., 2001 [Publ 3384], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 4 | 1 | -- | Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Elliptio_complanata **
Kandl et al., 2001 [Publ 3384], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 6 | 2 | -- | +----- Elliptio_crassidens **
--+
| +-- Fusconaia
+--+
+-- Pleurobema
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 2 | 2 | -- | +----- Pleurobema
--+
| +-- Elliptio_complanata **
+--+
+-- Eurynia
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | +----- Hemistena
--+
| +-- Elliptio_complanata **
+--+
+-- Fusconaia
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- | +----- Pleurobema
--+
| +-- Eurynia
+--+
+-- Elliptio_complanata **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 2 | 2 | -- | +----- Elliptio_complanata **
--+
| +-- Pleurobema
+--+
+-- Eurynia
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptio_complanata **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptio_complanata **
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 0 | 0 | -- | -- Elliptio_complanata **
Smith et al., 2020 [Publ 4136], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 18 | 8 | -- | +-------------- Elliptoideus
|
--+ +----------- Elliptio_crassidens **
| |
| +----------- Plethobasus
+--+
| +-------- Parvaspina
| |
| | +-- Hemistena
+--+-----+
| +-- Eurynia
|
| +----- Fusconaia
+--+
| +-- Pleuronaia
+--+
+-- Pleurobema
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- | -- Elliptio_complanata **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 2 | 2 | -- | +----- Pleurobema
--+
| +-- Eurynia
+--+
+-- Elliptio_complanata **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 5 | 3 | -- | +-------- Pleurobema
--+
| +----- Fusconaia
+--+
| +-- Elliptio_complanata **
+--+
+-- Eurynia
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 2 | 2 | -- | -- Elliptio_complanata **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 0 | 0 | -- | -- Elliptio_complanata **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Campbell, 2000 [Publ 2724], fig. 1-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Elliptio_complanata **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 0 | 0 | -- | -- Elliptio_complanata **
Burdick & White, 2007 [Publ 2679], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 27 | 6 | -- | +----- Eurynia
+-----------------+
| | +-- Plethobasus
| +--+
| +-- Elliptio_crassidens **
|
| +-------------------- Pleurobema_plenum
--+ |
| +-------------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum__1
| | +-----+
| | | +-- Fusconaia
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1
| | | |
| | +--+ +-- Pleurobema_cordatum__2
| | | |
| | +--+-- Pleurobema_sintoxia__2
| | |
+--+ +-- Pleurobema_sintoxia__3
|
| +-------------- Pleurobema_stabile
| |
| | +-- Pleurobema_rubellum
| | +-----+
| | | +-- Pleurobema_strodeanum
+--+ +--+
| | | +----- Pleurobema_beadleianum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
+--+ +-- Pleurobema_taitianum
|
| +-- Pleurobema_clava
| +-----+
| | +-- Pleurobema_oviforme
| |
+--+ +-- Pleurobema_decisum
| +--+
| | +-- Pleurobema_hanleyianum
+--+
| +-- Pleurobema_pyriforme
+--+
+-- Pleurobema_georgianum
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptio_complanata **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Elliptoideus
--+
| +-- Elliptio_complanata **
+--+
+-- Pleurobema
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 120 | 0 | 0 | -- | -- Elliptio_complanata **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- | +----- Elliptio_complanata **
--+
| +-- Pleurobema
+--+
+-- Eurynia
Johnson et al., 2018 [Publ 3844], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- | -- Elliptio_crassidens **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | -- Elliptio_complanata **
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | +----- Hemistena
--+
| +-- Elliptio_complanata **
+--+
+-- Fusconaia
Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | -- Elliptio_complanata **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 82 | 1 | 1 | -- | -- Elliptio_complanata **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 196 | 4 | 4 | -- | +-------- Elliptoideus
|
--+ +-- Elliptio_complanata **
| +--+
| | +-- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 3 | 3 | -- | +-------- Pleurobema
--+
| +----- Elliptio_crassidens **
+--+
| +-- Fusconaia
+--+
+-- Eurynia
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- | -- Elliptio_complanata **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 1 | 1 | -- | +-- Elliptio_complanata **
--+
+-- Pleurobema
|
