The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Pleurobemini genus Elliptoideus Frierson, 1927: cladomics
1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total)
analyzed in 27 cladograms in 15 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 630 | 64 | 8 | -- | +-------------------------------------------------------------- Elliptoideus_sloatianus **
|
| +----------------------------------------------------------- Parvaspina
| |
| | +-- Elliptio_fisheriana
--+ | +-----------------------------------------------------+
| | | +-- Elliptio_nasutidus
| | |
| | | +----------------------------------------------------- Elliptio_spinosa
| | | |
+--+ | | +----------------------------- Elliptio_dariensis__5
| | | |
| | | | +----- Elliptio_arctata__3
| | | | |
| | | | +----- Elliptio_crassidens__4
| | | | +--------------------+
| | | | | +----- Elliptio_mcmichaeli__2
| | | | | |
| | | | | | +-- Elliptio_crassidens__5
+--+ | | | +--+
| | +--------------------+ | +-- Elliptio_roanokensis
| | | | |
| | | | | +----- Elliptio_complanata__2
| | | | | |
| | | | | +----- Elliptio_pullata__2
| | | | | |
| | | | | +----- Elliptio_icterina__3
| | | | | |
| | | | +--------------------+ +-- Elliptio_dariensis__4
| | | | | +--+
| | | | | | +-- Elliptio_jayensis__2
| | | +--+ |
| | | | | +-- Elliptio_congaraea__2
+--+ | | | |
| | | | +-- Elliptio_crassidens__3
| | | +--+
| | | +-- Elliptio_fumata__2
| | | |
| | | +-- Elliptio_mcmichaeli__1
| | |
| | | +----------------------- Elliptio_crassidens__2
| | | |
| | | | +-- Elliptio_arca__2
| | | +--------------------+
| | | | +-- Elliptio_crassidens__1
| | | |
| | | | +-------- Elliptio_icterina__2
| | | | +-----------+
| | +--+ | | +----- Elliptio_dariensis__3
| | | | +--+
| | | | | +-- Elliptio_dariensis__2
| | | | +--+
| | | | +-- Elliptio_mediocris
| | | |
| | | | +-- Elliptio_folliculata__2
| | | | +-----------+
| | | | | +-- Elliptio_purpurella__2
| | +--+ |
| | | | +-- Elliptio_jayensis__1
+--+ | +--+-----------+
| | | | +-- Elliptio_pullata__1
| | | |
| | | | +----------- Elliptio_arca__1
| | | +--+
| | | | +-------- Elliptio_arctata__2
| | | +--+
| | | | +----- Elliptio_fumata__1
| | | +--+
| +--+ | +-- Elliptio_arctata__1
| | +--+
| | +-- Elliptio_purpurella__1
| |
| | +-- Elliptio_congaraea__1
| | |
| | +--------+-- Elliptio__1
| | | |
| | | +-- Elliptio__2
| | |
| +-----+ +-- Elliptio_complanata__1
| | +-----+
| | | +-- Elliptio_hopetonensis__1
| | |
| +--+ +----- Elliptio_dariensis__1
| | |
| | +----- Elliptio_producta
| +--+
| | +-- Elliptio_hopetonensis__2
| | |
| +--+-- Elliptio_hopetonensis__3
| |
| +-- Elliptio_icterina__1
|
| +----- Elliptio_shepardiana
| +-----------------------------------------+
| | | +-- Elliptio_angustata
| | +--+
| | +-- Elliptio_folliculata__1
| |
+--+ +-------------------------------------------- Eurynia
| |
| | +-- Hemistena
| | +--------------------------------------+
| | | +-- Plethobasus_cyphyus
+--+ |
| | +-------------------------------------- Elliptio_fraterna
| | |
| | | +-------------------------------- Pleurobema_plenum
| | | |
+--+ | +--+ +----------------------------- Pleurobema_stabile
| | | | |
| | | | | +-------------------------- Pleuronaia
| | | +--+ |
| | | | | +-- Pleurobema_decisum
| | | | | +-----+
| | | | | | +-- Pleurobema_hanleyianum
| | | +--+ +--------------+
+--+ | | | | +----- Pleurobema_pyriforme
| | | | +--+
| | | | | +-- Pleurobema_athearni
| | | | +--+
| | | | +-- Pleurobema_georgianum
| | +--+
| | | +-- Pleurobema_clava__1
| | | +--+
| | | | +-- Pleurobema_oviforme__1
| | | +--------------+
| | | | | +-- Pleurobema_clava__2
| | | | +--+
| | +--+ +-- Pleurobema_oviforme__2
+--+ |
| | +----------------- Pleurobema_beadleianum__2
| +--+
| | +-------------- Pleurobema_rubellum
| +--+
| | +----------- Pleurobema_strodeanum__2
| +--+
| | +-------- Pleurobema_beadleianum__1
| +--+
| | +----- Pleurobema_taitianum
| +--+
| | +-- Pleurobema_strodeanum__1
| +--+
| +-- Pleurobema_perovatum
|
| +----- Pleurobema_riddellii
| +-----------------+
| | | +-- Plethobasus_cooperianus
| | +--+
| | +-- Pleurobema_sintoxia
| |
+-----------+ +-------- Pleurobema_cordatum
| +-----------+
| | | +----- Fusconaia_subrotunda
| | +--+
| | | +-- Fusconaia_cor
| | +--+
+--+ +-- Fusconaia_cuneolus
|
| +----------------- Fusconaia_ozarkensis
| |
| | +----- Fusconaia_escambia__2
| | +--------+
+--+ | | +-- Fusconaia_burkei
| | +--+
| | +-- Fusconaia_escambia__1
| |
+--+ +----- Fusconaia_masoni
| |
| +-----+ +-- Fusconaia_cerina__1
| | | |
| | +--+-- Fusconaia_cerina__2
| | |
+--+ +-- Fusconaia_flava__2
|
| +-- Fusconaia_askewi
| +-----+
| | +-- Fusconaia_chunii
+--+
| +----- Fusconaia_flava__1
+--+
| +-- Fusconaia_iheringi
+--+
+-- Fusconaia_mitchelli
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 20 | 8 | -- | -- Elliptoideus_sloatianus **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------------------- Elliptoideus_sloatianus **
|
| +----------------- Pleurobema_stabile
--+ |
| | +----- Pleurobema_pyriforme
| | +--------+
+--+ | | +-- Pleurobema_clava
| | +--+
| | +-- Pleurobema_decisum
| |
+--+ +-- Pleurobema_cordatum
| +--------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleuronaia
+--+ +--+
| | | +-- Parvaspina
| | +--+
| | +-- Fusconaia
+--+
| +-- Eurynia
| +--+
| | +-- Hemistena
+--+
| +-- Plethobasus
+--+
+-- Elliptio
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 58 | 8 | -- | +----------------------------------------- Elliptoideus_sloatianus **
|
| +-------------------------------------- Parvaspina
| |
--+ | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
| | |
| | +----------------------------------- Hemistena
+--+ |
| +----------------------------------- Elliptio
| |
| | +-------------------------------- Pleuronaia
| | |
| | | +-- Pleurobema_cordatum
| | | +--------------------------+
+--+ | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 4 | 4 | -- | +----------- Elliptoideus_sloatianus **
--+
| +-------- Elliptio
+--+
| +----- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Campbell & Lydeard, 2012 [Publ 3308], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------- Elliptoideus_sloatianus **
|
| +----- Hemistena
| |
--+ +----- Plethobasus
| |
| +----- Elliptio
| |
+--+ +-- Eurynia
+--+
| +-- Parvaspina
|
| +-- Pleuronaia
| |
+--+-- Pleurobema
|
+-- Fusconaia
Smith et al., 2020 [Publ 4129], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptoideus_sloatianus **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 4 | 4 | -- | +-------- Elliptoideus_sloatianus **
|
--+ +-- Elliptio
| +--+
| | +-- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | -- Elliptoideus_sloatianus **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 29 | 8 | -- | +-------------- Elliptoideus_sloatianus **
|
--+ +----------- Hemistena
| |
| | +-- Plethobasus
+--+ +-----+
| | +-- Elliptio
| |
+--+ +----- Pleuronaia
| |
| +----- Fusconaia
+--+
+----- Pleurobema
|
| +-- Eurynia
+--+
+-- Parvaspina
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 4 | -- | +-------------- Elliptoideus_sloatianus **
--+
| +----------- Elliptio
+--+
| +-------- Pleuronaia
+--+
| +----- Pleurobema_riddellii
+--+
| +-- Fusconaia
+--+
+-- Pleurobema_strodeanum
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 29 | 8 | -- | +-------------------------------- Elliptoideus_sloatianus **
|
| +----------------------------- Plethobasus
--+ |
| +----------------------------- Parvaspina
| |
| +----------------------------- Elliptio
+--+
| +-------------------------- Eurynia
| |
| +-------------------------- Hemistena
| |
| | +----------------------- Fusconaia
+--+ |
| | +----- Pleurobema_sintoxia__2
| +-----------------+
| | | +-- Pleurobema_sintoxia__1
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +-------------------- Pleuronaia
| |
| | +-------- Pleurobema_rubellum
| | +--------+
+--+ | | +----- Pleurobema_strodeanum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
| | +-- Pleurobema_taitianum
+--+
| +-- Pleurobema_oviforme__1
| |
| +-----------+-- Pleurobema_oviforme__2
| | |
| | +-- Pleurobema_clava
+--+
| +----------- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_athearni
| +-----+
| | +-- Pleurobema_georgianum__2
+--+
| +----- Pleurobema_decisum
| |
+--+ +-- Pleurobema_hanleyianum__1
| |
+--+-- Pleurobema_hanleyianum__2
|
+-- Pleurobema_georgianum__1
Campbell & Lydeard, 2012 [Publ 3308], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 42 | 24 | 8 | -- | -- Elliptoideus_sloatianus **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Elliptoideus_sloatianus **
--+
| +-- Elliptio
+--+
+-- Pleurobema
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 2 | 2 | -- | -- Elliptoideus_sloatianus **
Smith et al., 2020 [Publ 4136], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 18 | 8 | -- | +-------------- Elliptoideus_sloatianus **
|
--+ +----------- Elliptio
| |
| +----------- Plethobasus
+--+
| +-------- Parvaspina
| |
| | +-- Hemistena
+--+-----+
| +-- Eurynia
|
| +----- Fusconaia
+--+
| +-- Pleuronaia
+--+
+-- Pleurobema
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | -- Elliptoideus_sloatianus **
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 3 | 3 | -- | -- Elliptoideus_sloatianus **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Elliptoideus_sloatianus **
--+
| +-- Elliptio
+--+
+-- Pleurobema
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 40 | 8 | -- | +-------------------------- Elliptoideus_sloatianus **
|
| +----------------------- Pleurobema_stabile
| |
| | +----- Pleurobema_riddellii
--+ | +-----------+
| | | | +-- Pleurobema_sintoxia
| | | +--+
| | +--+ +-- Pleurobema_cordatum
| | | |
+--+ | | +-------------- Fusconaia
| | | |
| | +--+ +----------- Pleuronaia
| | | |
| | +--+ +-- Eurynia
| | | +-----+
| | | | +-- Parvaspina
| | +--+
+--+ | +----- Elliptio
| +--+
| | +-- Hemistena
| +--+
| +-- Plethobasus
|
| +-- Pleurobema_clava
| +-----------+
| | +-- Pleurobema_oviforme
| |
+-----+ +----------- Pleurobema_pyriforme
| |
| | +-- Pleurobema_athearni
| | +--+
+--+ | +-- Pleurobema_georgianum
| +--+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_hanleyianum
+--+
| +----- Pleurobema_beadleianum
| |
| +----- Pleurobema_strodeanum
+--+
+----- Pleurobema_rubellum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Campbell & Lydeard, 2012 [Publ 3308], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------------------------- Elliptoideus_sloatianus **
|
--+ +----------------------- Pleurobema_stabile
| |
+--+ +-------------------- Hemistena
| |
+--+ +----------------- Eurynia
| |
| | +-- Pleurobema_cordatum
+--+ +-----------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleurobema_clava
+--+ +-----+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_pyriforme
+--+
| +-- Parvaspina
| +-----+
| | +-- Pleuronaia
+--+
| +----- Fusconaia
+--+
| +-- Plethobasus
+--+
+-- Elliptio
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | -- Elliptoideus_sloatianus **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 4 | 4 | -- | +-------- Elliptoideus_sloatianus **
|
--+ +-- Elliptio
| +--+
| | +-- Eurynia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 2 | 2 | -- | -- Elliptoideus_sloatianus **
Smith et al., 2020 [Publ 4129], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Pleurobema
--+
+-- Elliptoideus_sloatianus **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 40 | 8 | -- | +----------- Elliptoideus_sloatianus **
|
| +-------- Elliptio
--+ |
| | +-- Eurynia
| +-----+
| | +-- Parvaspina
+--+
| +-- Hemistena
+-----+
| +-- Plethobasus
|
| +----- Pleuronaia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | -- Elliptoideus_sloatianus **
|
