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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Gonideini

genus Gonidea Conrad, 1857: cladomics
     1 Recent valid species (2 total species)

Cladomics Summary

number of species = 1 (2 total)
analyzed in 80 cladograms in 56 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11016312012--
  +-------------- Sinosolenaia_carinata 
  |                                     
  |        +----- Gonidea_angulata **      
--+  +-----+                            
  |  |     |  +-- Microcondylaea        
  |  |     +--+                         
  +--+        +-- Leguminaia            
     |                                  
     |  +-------- Inversidens           
     +--+                               
        |  +----- Sinosolenaia_recognita
        +--+                            
           |  +-- Sinosolenaia_iridinea 
           +--+                         
              +-- Sinosolenaia_oleivora

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012100--
-- Gonidea_angulata **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
11018111--
  +-- Gonidea_angulata **
--+                   
  +-- Leguminaia

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
11013587--
  +----------------- Inversidens     
  |                                  
--+  +-------------- Sinosolenaia    
  |  |                               
  +--+           +-- Gonidea_angulata **
     |  +--------+                   
     |  |        +-- Microcondylaea  
     +--+                            
        |  +-------- Koreosolenaia   
        +--+                         
           |  +----- Parvasolenaia   
           +--+                      
              |  +-- Postolata       
              +--+                   
                 +-- Ptychorhynchus

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
11011422--
  +----- Sinosolenaia    
--+                      
  |  +-- Leguminaia      
  +--+                   
     +-- Gonidea_angulata **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011411--
-- Gonidea_angulata **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
110110100--
-- Gonidea_angulata **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015555--
  +----------- Sinosolenaia    
  |                            
--+        +-- Ptychorhynchus  
  |  +-----+                   
  |  |     +-- Parvasolenaia   
  +--+                         
     |  +----- Gonidea_angulata **
     +--+                      
        |  +-- Microcondylaea  
        +--+                   
           +-- Leguminaia

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
110110400--
-- Gonidea_angulata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113763--
  +-------- Sinosolenaia    
--+                         
  |  +----- Gonidea_angulata **
  +--+                      
     |  +-- Microcondylaea  
     +--+                   
        +-- Leguminaia

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014411--
  +-- Gonidea_angulata **
--+                   
  +-- Microcondylaea

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
11012511--
-- Gonidea_angulata **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
11012376--
           +-- Ptychorhynchus  
  +--------+                   
  |        +-- Parvasolenaia   
--+                            
  |  +-------- Inversidens     
  +--+                         
     |  +----- Gonidea_angulata **
     +--+                      
        |  +-- Microcondylaea  
        +--+                   
           +-- Leguminaia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Gonidea_angulata **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Gonidea_angulata **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013500--
-- Gonidea_angulata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119786--
  +----------- Sinosolenaia    
  |                            
  |     +----- Gonidea_angulata **
--+  +--+                      
  |  |  |  +-- Leguminaia      
  |  |  +--+                   
  +--+     +-- Microcondylaea  
     |                         
     |  +----- Ptychorhynchus  
     +--+                      
        |  +-- Parvasolenaia   
        +--+                   
           +-- Koreosolenaia

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014500--
-- Gonidea_angulata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015600--
-- Gonidea_angulata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1101121118--
     +-------- Sinosolenaia    
  +--+                         
  |  |  +----- Gonidea_angulata **
  |  +--+                      
  |     |  +-- Leguminaia      
--+     +--+                   
  |        +-- Microcondylaea  
  |                            
  |  +-------- Inversidens     
  |  |                         
  +--+     +-- Obovalis        
     |  +--+                   
     |  |  +-- Ptychorhynchus  
     +--+                      
        |  +-- Parvasolenaia   
        +--+                   
           +-- Koreosolenaia

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1101461511--
  +-------------------- Sinosolenaia    
  |                                     
  |              +----- Gonidea_angulata **
--+  +-----------+                      
  |  |           |  +-- Leguminaia      
  |  |           +--+                   
  +--+              +-- Microcondylaea  
     |                                  
     |  +-------------- Pseudopostolata 
     |  |                               
     +--+           +-- Koreosolenaia   
        |  +--------+                   
        |  |        +-- Parvasolenaia   
        +--+                            
           |  +-------- Postolata       
           |  |                         
           +--+     +-- Guiunio         
              |  +--+                   
              |  |  +-- Cosmopseudodon  
              +--+                      
                 |  +-- Obovalis        
                 +--+                   
                    +-- Ptychorhynchus

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
11012600--
-- Gonidea_angulata **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011311--
  +-- Gonidea_angulata **
--+                   
  +-- Microcondylaea

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014500--
-- Gonidea_angulata **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia    
  |                               
--+  +----------- Gonidea_angulata **
  |  |                            
  +--+----------- Microcondylaea  
     |                            
     |  +-------- Koreosolenaia   
     +--+                         
        |  +----- Parvasolenaia   
        +--+                      
           |  +-- Postolata       
           +--+                   
              +-- Ptychorhynchus

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
11013588--
  +-------------------- Sinosolenaia    
  |                                     
--+                 +-- Gonidea_angulata **
  |  +--------------+                   
  |  |              +-- Microcondylaea  
  +--+                                  
     |  +-------------- Koreosolenaia   
     +--+                               
        |  +----------- Parvasolenaia   
        +--+                            
           |  +-------- Postolata       
           +--+                         
              |  +----- Cosmopseudodon  
              +--+                      
                 |  +-- Ptychorhynchus  
                 +--+                   
                    +-- Obovalis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Gonidea_angulata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113763--
  +-------- Sinosolenaia_carinata 
  |                               
  |     +-- Sinosolenaia_oleivora 
--+-----+                         
  |     +-- Sinosolenaia_recognita
  |                               
  |  +----- Gonidea_angulata **      
  +--+                            
     |  +-- Microcondylaea        
     +--+                         
        +-- Leguminaia

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119996--
  +-------- Sinosolenaia_carinata 
  |                               
  |  +----- Gonidea_angulata **      
  +--+                            
  |  |  +-- Leguminaia            
  |  +--+                         
  |     +-- Microcondylaea        
--+                               
  |  +----- Ptychorhynchus        
  +--+                            
  |  |  +-- Parvasolenaia         
  |  +--+                         
  |     +-- Koreosolenaia         
  |                               
  |  +----- Sinosolenaia_recognita
  +--+                            
     |  +-- Sinosolenaia_iridinea 
     +--+                         
        +-- Sinosolenaia_oleivora

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017954--
  +-------- Sinosolenaia    
--+                         
  |  +----- Gonidea_angulata **
  +--+                      
     |  +-- Microcondylaea  
     +--+                   
        +-- Leguminaia

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
11011800--
-- Gonidea_angulata **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1101451410--
  +-------------------- Sinosolenaia    
  |                                     
  |              +----- Gonidea_angulata **
--+  +-----------+                      
  |  |           |  +-- Leguminaia      
  |  |           +--+                   
  +--+              +-- Microcondylaea  
     |                                  
     |  +-------------- Pseudopostolata 
     |  |                               
     +--+           +-- Parvasolenaia   
        |  +--------+                   
        |  |        +-- Koreosolenaia   
        +--+                            
           |  +-------- Postolata       
           +--+                         
              |  +----- Ptychorhynchus  
              +--+                      
                 |  +-- Cosmopseudodon  
                 +--+                   
                    +-- Obovalis

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014600--
-- Gonidea_angulata **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
11011600--
-- Gonidea_angulata **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
110144139--
  +-------------- Sinosolenaia    
  |                               
  |        +----- Gonidea_angulata **
--+  +-----+                      
  |  |     |  +-- Leguminaia      
  |  |     +--+                   
  |  |        +-- Microcondylaea  
  +--+                            
     |        +-- Koreosolenaia   
     |  +-----+                   
     |  |     +-- Parvasolenaia   
     +--+                         
        |  +----- Postolata       
        +--+                      
           |  +-- Ptychorhynchus  
           +--+                   
              +-- Obovalis

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
11011900--
-- Gonidea_angulata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014600--
-- Gonidea_angulata **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
11011300--
-- Gonidea_angulata **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
11014822--
-- Gonidea_angulata **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012100--
-- Gonidea_angulata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
1101471612--
  +----------------------- Sinosolenaia    
  |                                        
  |                 +----- Gonidea_angulata **
--+  +--------------+                      
  |  |              |  +-- Leguminaia      
  |  |              +--+                   
  +--+                 +-- Microcondylaea  
     |                                     
     |  +----------------- Pseudopostolata 
     |  |                                  
     +--+              +-- Koreosolenaia   
        |  +-----------+                   
        |  |           +-- Parvasolenaia   
        +--+                               
           |  +----------- Postolata       
           |  |                            
           +--+        +-- Guiunio         
              |  +-----+                   
              |  |     +-- Cosmopseudodon  
              +--+                         
                 |  +----- Obovalis        
                 +--+                      
                    |  +-- Tenuiunio       
                    +--+                   
                       +-- Ptychorhynchus

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013522--
-- Gonidea_angulata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112363--
-- Gonidea_angulata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019076--
        +----- Sinosolenaia    
  +-----+                      
  |     |  +-- Ptychorhynchus  
  |     +--+                   
--+        +-- Parvasolenaia   
  |                            
  |  +-------- Inversidens     
  +--+                         
     |  +----- Gonidea_angulata **
     +--+                      
        |  +-- Microcondylaea  
        +--+                   
           +-- Leguminaia

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013011--
  +-- Gonidea_angulata **
--+                   
  +-- Microcondylaea

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017522--
  +----- Sinosolenaia    
--+                      
  |  +-- Leguminaia      
  +--+                   
     +-- Gonidea_angulata **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
110111085--
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
--+                            
  +-- Gonidea_angulata **         
  |                            
  +-- Ptychorhynchus           
  |                            
  +-- Parvasolenaia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Gonidea_angulata **

Lopes-Lima et al., 2021 [Publ 4217], fig. 14

iTtTbiriSoTroTroSiD
1101442--
  +----- Leguminaia      
--+                      
  |  +-- Gonidea_angulata **
  +--+                   
     +-- Microcondylaea

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
11013322--
  +----- Sinosolenaia    
--+                      
  |  +-- Leguminaia      
  +--+                   
     +-- Gonidea_angulata **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Gonidea_angulata **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia    
  |                               
  |           +-- Gonidea_angulata **
--+  +--------+                   
  |  |        +-- Microcondylaea  
  |  |                            
  +--+        +-- Koreosolenaia   
     |  +-----+                   
     |  |     +-- Parvasolenaia   
     +--+                         
        |  +----- Postolata       
        +--+                      
           |  +-- Obovalis        
           +--+                   
              +-- Ptychorhynchus

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
11011631--
  +-- Gonidea_angulata **
--+                   
  +-- Inversidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
11013422--
  +----- Sinosolenaia    
--+                      
  |  +-- Gonidea_angulata **
  +--+                   
     +-- Leguminaia

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
11011196--
  +-------------- Sinosolenaia_carinata
  |                                    
  |        +----- Inversidens          
--+  +-----+                           
  |  |     |  +-- Gonidea_angulata **     
  |  |     +--+                        
  +--+        +-- Leguminaia           
     |                                 
     |  +-------- Sinosolenaia_oleivora
     +--+                              
        |  +----- Postolata            
        +--+                           
           |  +-- Obovalis             
           +--+                        
              +-- Ptychorhynchus

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015600--
-- Gonidea_angulata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015243--
  +-------- Sinosolenaia_carinata
  |                              
--+  +----- Gonidea_angulata **     
  |  |                           
  +--+----- Sinosolenaia_oleivora
     |                           
     |  +-- Parvasolenaia        
     +--+                        
        +-- Ptychorhynchus

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019076--
        +----- Gonidea_angulata **
  +-----+                      
  |     |  +-- Leguminaia      
  |     +--+                   
--+        +-- Microcondylaea  
  |                            
  |  +-------- Inversidens     
  +--+                         
     |  +----- Sinosolenaia    
     +--+                      
        |  +-- Parvasolenaia   
        +--+                   
           +-- Ptychorhynchus

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
11013000--
-- Gonidea_angulata **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
11015822--
  +----- Sinosolenaia    
--+                      
  |  +-- Gonidea_angulata **
  +--+                   
     +-- Leguminaia

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia    
  |                               
--+           +-- Gonidea_angulata **
  |  +--------+                   
  |  |        +-- Microcondylaea  
  +--+                            
     |  +-------- Koreosolenaia   
     +--+                         
        |  +----- Parvasolenaia   
        +--+                      
           |  +-- Postolata       
           +--+                   
              +-- Ptychorhynchus

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018122--
-- Gonidea_angulata **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Gonidea_angulata **

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
1101400--
-- Gonidea_angulata **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Gonidea_angulata **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
110144139--
  +-------------- Sinosolenaia    
  |                               
  |        +----- Gonidea_angulata **
--+  +-----+                      
  |  |     |  +-- Leguminaia      
  |  |     +--+                   
  |  |        +-- Microcondylaea  
  +--+                            
     |        +-- Koreosolenaia   
     |  +-----+                   
     |  |     +-- Parvasolenaia   
     +--+                         
        |  +----- Postolata       
        +--+                      
           |  +-- Ptychorhynchus  
           +--+                   
              +-- Obovalis

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119996--
  +----------- Sinosolenaia    
  |                            
  |     +----- Gonidea_angulata **
--+  +--+                      
  |  |  |  +-- Leguminaia      
  |  |  +--+                   
  +--+     +-- Microcondylaea  
     |                         
     |     +-- Ptychorhynchus  
     |     |                   
     +-----+-- Parvasolenaia   
           |                   
           +-- Koreosolenaia

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia    
  |                               
--+           +-- Gonidea_angulata **
  |  +--------+                   
  |  |        +-- Microcondylaea  
  +--+                            
     |  +-------- Parvasolenaia   
     +--+                         
        |  +----- Koreosolenaia   
        +--+                      
           |  +-- Postolata       
           +--+                   
              +-- Ptychorhynchus

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011411--
-- Gonidea_angulata **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111685--
-- Gonidea_angulata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017522--
  +----- Sinosolenaia    
--+                      
  |  +-- Gonidea_angulata **
  +--+                   
     +-- Leguminaia

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
11014922--
  +----- Sinosolenaia    
--+                      
  |  +-- Gonidea_angulata **
  +--+                   
     +-- Leguminaia

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
110110100--
-- Gonidea_angulata **
 
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