The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Pleurobemini genus Hemistena Rafinesque, 1820: cladomics
1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total)
analyzed in 25 cladograms in 12 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 103 | 57 | 8 | -- | +----------------------------------------- Elliptoideus
|
| +-- Elliptio_lanceolata
| +-----------------------------------+
| | +-- Parvaspina
| |
| | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
--+ | |
| | +----------------------------------- Hemistena_lata **
| | |
| | | +-- Elliptio_complanata
| | | |
| | | +-- Elliptio_hopetonensis
| | | |
| | | +-- Elliptio_producta__1
| | | |
| | | +-- Elliptio_shepardiana
+--+ | |
| | +-- Elliptio_angustata
| | |
| | +-- Elliptio_arca
| | |
| | +-- Elliptio_arctata
| | |
| | +-- Elliptio_jayensis
| | |
| | +-- Elliptio_crassidens
| | |
| +--------------------------------+-- Elliptio_dariensis
| | |
| | +-- Elliptio_fisheriana
| | |
| | +-- Elliptio_folliculata
| | |
+--+ +-- Elliptio_fumata
| |
| +-- Elliptio_icterina
| |
| +-- Elliptio_mcmichaeli
| |
| +-- Elliptio_producta__2
| |
| +-- Elliptio_pullata
| |
| +-- Elliptio_purpurella
| |
| +-- Elliptio_congaraea
|
| +-------------------------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
| | +--------------------------+
| | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Smith et al., 2020 [Publ 4136], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 18 | 8 | -- | +-------------- Elliptoideus
|
--+ +----------- Elliptio
| |
| +----------- Plethobasus
+--+
| +-------- Parvaspina
| |
| | +-- Hemistena_lata **
+--+-----+
| +-- Eurynia
|
| +----- Fusconaia
+--+
| +-- Pleuronaia
+--+
+-- Pleurobema
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 40 | 8 | -- | +-------------------------- Elliptoideus
|
| +----------------------- Pleurobema_stabile
| |
| | +----- Pleurobema_riddellii
--+ | +-----------+
| | | | +-- Pleurobema_sintoxia
| | | +--+
| | +--+ +-- Pleurobema_cordatum
| | | |
+--+ | | +-------------- Fusconaia
| | | |
| | +--+ +----------- Pleuronaia
| | | |
| | +--+ +-- Eurynia
| | | +-----+
| | | | +-- Parvaspina
| | +--+
+--+ | +----- Elliptio
| +--+
| | +-- Hemistena_lata **
| +--+
| +-- Plethobasus
|
| +-- Pleurobema_clava
| +-----------+
| | +-- Pleurobema_oviforme
| |
+-----+ +----------- Pleurobema_pyriforme
| |
| | +-- Pleurobema_athearni
| | +--+
+--+ | +-- Pleurobema_georgianum
| +--+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_hanleyianum
+--+
| +----- Pleurobema_beadleianum
| |
| +----- Pleurobema_strodeanum
+--+
+----- Pleurobema_rubellum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Campbell & Lydeard, 2012 [Publ 3308], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------------------------- Elliptoideus
|
--+ +----------------------- Pleurobema_stabile
| |
+--+ +-------------------- Hemistena_lata **
| |
+--+ +----------------- Eurynia
| |
| | +-- Pleurobema_cordatum
+--+ +-----------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleurobema_clava
+--+ +-----+
| | | +-- Pleurobema_decisum
| | +--+
| | +-- Pleurobema_pyriforme
+--+
| +-- Parvaspina
| +-----+
| | +-- Pleuronaia
+--+
| +----- Fusconaia
+--+
| +-- Plethobasus
+--+
+-- Elliptio
Perkins et al., 2017 [Publ 3777], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 6 | -- | +----------------- Pleurobema
--+
| +-------------- Pleuronaia
+--+
| +----------- Fusconaia
+--+
| +-------- Parvaspina
+--+
| +----- Elliptio
+--+
| +-- Eurynia
+--+
+-- Hemistena_lata **
Serb et al., 2003 [Publ 2659], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | +----- Hemistena_lata **
--+
| +-- Elliptio
+--+
+-- Fusconaia
Perkins et al., 2017 [Publ 3777], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 21 | 6 | -- | +----------------- Parvaspina
|
| +-------------- Elliptio_arca
| |
--+ +-------------- Hemistena_lata **
| |
| | +-- Elliptio_arctata
| +-----------+
| | +-- Elliptio_crassidens
+--+
| +----------- Eurynia
| |
| | +-------- Pleurobema_oviforme
| | |
| | +-------- Pleurobema_clava
| | |
+--+ +-------- Pleurobema_pyriforme
| |
| +-------- Pleurobema_georgianum
| |
| +-------- Pleurobema_beadleianum
| |
+--+-------- Pleurobema_hanleyianum
|
+-------- Pleurobema_decisum
|
| +-- Pleurobema_sintoxia
+-----+
| +-- Pleurobema_rubellum
|
| +----- Elliptio_spinosa
| |
+--+ +-- Fusconaia_escambia
| |
| +-- Fusconaia_flava
+--+
+-- Fusconaia_cerina
|
+-- Pleuronaia
Serb et al., 2003 [Publ 2659], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 4 | 3 | -- | +----- Hemistena_lata **
--+
| +-- Elliptio
+--+
+-- Fusconaia
Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 20 | 8 | -- | +----------------- Elliptio
|
| +-------------- Parvaspina
--+ |
| +-------------- Plethobasus
| |
| | +-- Hemistena_lata **
+--+-----------+
| +-- Eurynia
|
| +----------- Fusconaia
| |
| | +----- Pleurobema_riddellii
+--+ +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +----- Pleuronaia
| |
+--+ +-- Pleurobema_pyriforme
| |
+--+-- Pleurobema_strodeanum
|
+-- Pleurobema_clava
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +-------- Pleuronaia
|
--+-------- Pleurobema
|
| +----- Fusconaia
+--+
| +-- Hemistena_lata **
+--+
+-- Eurynia
Campbell et al., 2008 [Publ 2675], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 23 | 7 | -- | +----------------- Parvaspina
|
--+ +-------------- Hemistena_lata **
| |
+--+ +-- Plethobasus
| +--------+
| | +-- Elliptio
+--+
| +-------- Fusconaia
+--+
| +----- Pleurobema
+--+
| +-- Eurynia
+--+
+-- Pleuronaia
Campbell & Lydeard, 2012 [Publ 3308], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------- Elliptoideus
|
| +----- Hemistena_lata **
| |
--+ +----- Plethobasus
| |
| +----- Elliptio
| |
+--+ +-- Eurynia
+--+
| +-- Parvaspina
|
| +-- Pleuronaia
| |
+--+-- Pleurobema
|
+-- Fusconaia
Campbell & Lydeard, 2012 [Publ 3308], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 42 | 24 | 8 | -- | +----------------------- Elliptio
|
--+ +-------------------- Eurynia
| |
| | +----------------- Parvaspina
+--+ |
| | +-- Hemistena_lata **
| +--------------+
+--+ +-- Plethobasus
|
| +-------------- Pleuronaia
| |
| | +-- Pleurobema_cordatum
+--+ +--------+
| | +-- Fusconaia
| |
+--+ +-- Pleurobema_stabile
| +-----+
| | +-- Pleurobema_sintoxia
+--+
| +----- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
+--+
+-- Pleurobema_pyriforme
Burlakova et al., 2012 [Publ 3486], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 34 | 7 | -- | +----------------------------------- Parvaspina
|
| +-------------------------------- Pleurobema_plenum
--+ |
| | +-- Plethobasus
| | +--+
+--+ | +-- Hemistena_lata **
| +-----------------------+
| | | +-- Eurynia
| | +--+
| | +-- Elliptio
+--+
| +-------------------------- Pleurobema_cordatum__2
| |
| | +----------------------- Fusconaia
+--+ |
| | +-------- Pleurobema_stabile
| | |
| | +-----------+ +----- Pleurobema_riddellii
+--+ | | |
| | +--+ +-- Pleurobema_cordatum__1
| | | |
| | | +-- Pleurobema_sintoxia__1
| | +--+
+--+ +-- Pleurobema_sintoxia__2
| |
| +-- Pleurobema_sintoxia__3
|
| +----------------- Pleuronaia
| |
| | +-- Pleurobema_georgianum
+--+ +-----------+
| | +-- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_decisum
| +--------+
| | +-- Pleurobema_hanleyianum
| |
+--+ +-- Pleurobema_clava
| +-----+
| | +-- Pleurobema_oviforme
| |
+--+ +----- Pleurobema_rubellum
| |
| +----- Pleurobema_beadleianum
+--+
+----- Pleurobema_strodeanum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Burlakova et al., 2012 [Publ 3486], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 34 | 7 | -- | +----------------------- Hemistena_lata **
|
+----------------------- Pleurobema_stabile
|
--+ +-------------------- Eurynia
| |
| | +-------- Pleurobema_riddellii
| | |
| | +--------+ +----- Pleurobema_sintoxia__2
+--+ | | |
| | +--+----- Pleurobema_sintoxia__3
| | |
| | | +-- Pleurobema_cordatum
| | +--+
| | +-- Pleurobema_sintoxia__1
+--+
| +-------------- Parvaspina
| |
| +-------------- Pleuronaia
| |
| +-------------- Fusconaia
| |
| | +-- Plethobasus
+--+-----------+
| +-- Elliptio
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
| |
| | +-------- Pleurobema_decisum
+--+ |
| +-------- Pleurobema_pyriforme
| |
| +-------- Pleurobema_hanleyianum
+--+
+-------- Pleurobema_georgianum
|
| +----- Pleurobema_rubellum
| |
| +----- Pleurobema_beadleianum
+--+
+----- Pleurobema_strodeanum
|
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 7 | 5 | -- | +----------- Parvaspina
|
--+ +-- Eurynia
| +-----+
| | +-- Hemistena_lata **
+--+
| +----- Pleurobema
+--+
| +-- Pleuronaia
+--+
+-- Fusconaia
Campbell & Lydeard, 2012 [Publ 3308], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 24 | 8 | -- | +-------------------- Elliptoideus
|
| +----------------- Pleurobema_stabile
--+ |
| | +----- Pleurobema_pyriforme
| | +--------+
+--+ | | +-- Pleurobema_clava
| | +--+
| | +-- Pleurobema_decisum
| |
+--+ +-- Pleurobema_cordatum
| +--------+
| | +-- Pleurobema_sintoxia
| |
| | +----- Pleuronaia
+--+ +--+
| | | +-- Parvaspina
| | +--+
| | +-- Fusconaia
+--+
| +-- Eurynia
| +--+
| | +-- Hemistena_lata **
+--+
| +-- Plethobasus
+--+
+-- Elliptio
Campbell et al., 2008 [Publ 2675], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 21 | 7 | -- | Burlakova et al., 2012 [Publ 3486], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 34 | 7 | -- | +----------------------------------- Parvaspina
|
| +-- Plethobasus
--+ +-----------------------------+
| | +-- Elliptio
| |
+--+ +----------------------------- Fusconaia
| |
| | +-- Eurynia
+--+ +-----------------------+
| | +-- Hemistena_lata **
| |
| | +----------- Pleuronaia
+--+ +-----------+
| | | +-------- Pleurobema_riddellii
| | +--+
| | | +----- Pleurobema_sintoxia__2
| | +--+
+--+ | +-- Pleurobema_sintoxia__1
| +--+
| +-- Pleurobema_cordatum
|
| +-------------------- Pleurobema_stabile
| |
+--+ +----------------- Pleurobema_pyriforme
| |
| | +----- Pleurobema_georgianum
+--+ +--------+
| | | +-- Pleurobema_hanleyianum
| | +--+
| | +-- Pleurobema_decisum
+--+
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_beadleianum
| |
+--+ +-- Pleurobema_rubellum
| +--+
| | +-- Pleurobema_strodeanum
+--+
| +-- Pleurobema_perovatum
+--+
+-- Pleurobema_taitianum
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 7 | 5 | -- | +----------- Parvaspina
|
--+ +-- Eurynia
| +-----+
| | +-- Hemistena_lata **
+--+
| +----- Pleurobema
+--+
| +-- Pleuronaia
+--+
+-- Fusconaia
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 29 | 8 | -- | +-------------------------------- Elliptoideus
|
| +----------------------------- Plethobasus
--+ |
| +----------------------------- Parvaspina
| |
| +----------------------------- Elliptio
+--+
| +-------------------------- Eurynia
| |
| +-------------------------- Hemistena_lata **
| |
| | +----------------------- Fusconaia
+--+ |
| | +----- Pleurobema_sintoxia__2
| +-----------------+
| | | +-- Pleurobema_sintoxia__1
| | +--+
+--+ +-- Pleurobema_cordatum
|
| +-------------------- Pleuronaia
| |
| | +-------- Pleurobema_rubellum
| | +--------+
+--+ | | +----- Pleurobema_strodeanum
| | +--+
| | | +-- Pleurobema_perovatum
| | +--+
| | +-- Pleurobema_taitianum
+--+
| +-- Pleurobema_oviforme__1
| |
| +-----------+-- Pleurobema_oviforme__2
| | |
| | +-- Pleurobema_clava
+--+
| +----------- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_athearni
| +-----+
| | +-- Pleurobema_georgianum__2
+--+
| +----- Pleurobema_decisum
| |
+--+ +-- Pleurobema_hanleyianum__1
| |
+--+-- Pleurobema_hanleyianum__2
|
+-- Pleurobema_georgianum__1
Campbell et al., 2008 [Publ 2675], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 21 | 7 | -- | Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 40 | 8 | -- | +----------- Elliptoideus
|
| +-------- Elliptio
--+ |
| | +-- Eurynia
| +-----+
| | +-- Parvaspina
+--+
| +-- Hemistena_lata **
+-----+
| +-- Plethobasus
|
| +----- Pleuronaia
+--+
| +-- Fusconaia
+--+
+-- Pleurobema
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 29 | 8 | -- | +-------------- Elliptoideus
|
--+ +----------- Hemistena_lata **
| |
| | +-- Plethobasus
+--+ +-----+
| | +-- Elliptio
| |
+--+ +----- Pleuronaia
| |
| +----- Fusconaia
+--+
+----- Pleurobema
|
| +-- Eurynia
+--+
+-- Parvaspina
Campbell et al., 2008 [Publ 2675], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 23 | 7 | -- | +-- Pleurobema_cordatum
|
+--------+-- Pleurobema_sintoxia__1
| |
| +-- Pleurobema_sintoxia__2
+--------+
| | +-------- Fusconaia
| | |
| +--+ +----- Pleuronaia
| | |
| | | +-- Eurynia
| +--+ |
| | +-- Hemistena_lata **
| | |
--+ +--+-- Plethobasus
| |
| +-- Parvaspina
| |
| +-- Elliptio
|
| +----------------- Pleurobema_stabile
| |
| | +-------------- Pleurobema_pyriforme
| | |
+--+ | +-- Pleurobema_georgianum__2
| +-----------+
| | +-- Pleurobema_athearni
| |
| | +----- Pleurobema_decisum
+--+--------+
| | +-- Pleurobema_hanleyianum
| +--+
| +-- Pleurobema_georgianum__1
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_rubellum
| |
+--+-------- Pleurobema_strodeanum
|
| +----- Pleurobema_beadleianum
+--+
| +-- Pleurobema_taitianum
+--+
+-- Pleurobema_perovatum
|
