The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | ... | Quadrulini | Quadrulina | Pustulosa kieneriana nominal species Unio asperatus Lea, 1861  Lea, 1861. Proc. Acad. Nat. Sci. 13: 41.
Lea, 1862. Jour. Acad. Nat. Sci. 5 [O. 8]: 68 [72], pl. 7, fig. 218.
Type(s): HOLOTYPE USNM_84272, Claiborne, Ala.
= Pustulosa kieneriana Taxonomic Opinions= Cyclonaias asperata — Hauck et al., 2023 + Cyclonaias kieneriana (Lea, 1852) — Graf & Cummings, 2021 + Cyclonaias kieneriana (Lea, 1852) — FMCS, 2021 + Cyclonaias kieneriana (Lea, 1852) — Lopes-Lima et al., 2019 = Cyclonaias asperata — Johnson et al., 2018 = Cyclonaias asperata (Lea, 1861) — Williams et al., 2017 = Quadrula asperata (I. Lea, 1861) — ITIS, 2012 = Quadrula asperata (I. Lea, 1861) — Szumowski et al., 2012 = Quadrula asperata — Haag, 2012 = Quadrula asperata Lea, 1861 — Bogan, 2010 = Amphinaias asperata — Cummings & Graf, 2010 = Quadrula asperata (Lea, 1861) — Williams et al., 2008 = Amphinaias asperata (Lea, 1861) — Graf & Cummings, 2007 = Quadrula asperata — Zhou et al., 2007 = Quadrula asperata (I. Lea) — Jones et al., 2005 = Quadrula asperata — Serb et al., 2003 = Quadrula asperata (Lea) — Lydeard et al., 1999 = Quadrula asperata (Lea, 1861) — Turgeon et al., 1998 = Amphinaias asperata (Lea, 1861) — IUCN, 1996 = Quadrula asperata (I. Lea) — Williams et al., 1993 = Quadrula asperata (Lea, 1861) — Turgeon et al., 1988 + Quadrula pustulosa (Lea, 1831) — R.I. Johnson, 1980 + Quadrula pustulosa (Lea, 1831) — Burch, 1975 + Quadrula (Pustulosa) pustulosa pernodosa (Lea, 1845) — Haas, 1969 = Quadrula (Bullata) pustulosa-asperata (Lea, 1861) — Frierson, 1927 = Quadrula asperata (Lea, 1861) — Ortmann, 1923 = Quadrula pustulosa asperata (Lea) — Utterback, 1915 + Quadrula (sec. Theliderma) pustulosa var. pernodosa (Lea, 1845) — Simpson, 1914 + Quadrula (sec. Theliderma) pustulosa var. pernodosa (Lea, 1845) — Simpson, 1900 + Unio pustulosus Lea, 1830 — Call, 1895 = Unio asperatus Lea, 1861 — Paetel, 1890 = Unio asperatus Lea — B.H. Wright, 1888 = Margaron (Unio) asperatus (Lea) — Lea, 1870 = Unio asperatus Lea, 1861 — Sowerby, 1868
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