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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | ... | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Pustulosa species Pustulosa nodulata (Rafinesque, 1820): synonymy
2 specimens + 12 Genbank vouchers Obliquaria (Quadrula) nodulata Rafinesque, 1820
 Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 307, pl. 81, figs. 17-18 [also as ‘Unio nodulata’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 51.
Type(s): LECTOTYPE ANSP_20225, Kentucky River.
— Simpson (1900), Simpson (1914)
Quadrula nodulata — Utterback (1915), Vanatta (1915), Ortmann & Walker (1922), Murray & Leonard (1962), Burch (1975), ... Boyer et al. (2011), ITIS (2012), Szumowski et al. (2012), Haag (2012), McMurray et al. (2012)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) nodulata — F.C. Baker (1928)
Quadrula (Pustulosa) nodulata — Haas (1969)
Obliquaria nodulata — Burch (1975)
Amphinaias nodulata — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011)
Cyclonaias nodulata — Williams et al. (2017), Burlakova et al. (2018), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
Pustulosa nodulata — Neemuchwala et al. (2023), FMCS (2025), Campbell & Lydeard (2026) | |
Unio pustulatus Lea, 1831
  Lea, 1831. Trans. Amer. Phil. Soc. 4 [O. 1]: 79 [89], pl. 7, fig. 9.
Type(s): SYNTYPE USNM_84200, Cincinnati, Ohio.
Quadrula (sec. Theliderma) pustulata — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulatus — Call (1900)
Quadrula pustulata — Ortmann (1914)
Unio pustulatus, var. — de Gregorio (1914)
Quadrula nodulata — Utterback (1915), Vanatta (1915), Ortmann & Walker (1922), Murray & Leonard (1962), Burch (1975), R.I. Johnson (1980), Oesch (1984), Vidrine (1993), Howells et al. (1996), Strayer & Jirka (1997), Parmalee & Bogan (1998), Watters et al. (2009)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) nodulata — F.C. Baker (1928)
Quadrula (Pustulosa) nodulata — Haas (1969)
Cyclonaias nodulata — Lopes-Lima et al. (2019) | |
Unio nodulatus Say, 1834
‘Rafinesque’ of Authors.
Say, 1834. Amer. Conch. 6: ??.
Conrad, 1837. Monogr. Fam. Unionidae 1(9): 80, pl. 45, fig. 1.
 Reeve, 1864. Conch. Icon. 16: pl. 13, sp. 51.
(reference to Obliquaria (Quadrula) nodulata Rafinesque, 1820)
Quadrula (sec. Theliderma) pustulata — Simpson (1900), Simpson (1914)
Quadrula nodulata — Utterback (1915), Watters et al. (2009) | |
Unio undulatus Lea, 1852
‘Con.’ Lea, 1852. Synopsis Fam. Naiades: 22.
(error for Unio nodulatus Say, 1834) | |
Unio modulatus Küster, 1862
‘Raf.’ Küster, 1862. Conchyl. Cab. 9(2): 311.
(error for Obliquaria (Quadrula) nodulata Rafinesque, 1820) | |
Taxonomic History (since 1900) — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) pustulata — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulatus — Call (1900)
Quadrula pustulata — Ortmann (1914)
Unio pustulatus, var. — de Gregorio (1914)
Quadrula nodulata — Utterback (1915), Vanatta (1915), Ortmann & Walker (1922), Murray & Leonard (1962), Burch (1975), ... Boyer et al. (2011), ITIS (2012), Szumowski et al. (2012), Haag (2012), McMurray et al. (2012)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) nodulata — F.C. Baker (1928)
Quadrula (Pustulosa) nodulata — Haas (1969)
Obliquaria nodulata — Burch (1975)
Amphinaias nodulata — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011)
Cyclonaias nodulata — Williams et al. (2017), Burlakova et al. (2018), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
Pustulosa nodulata — Neemuchwala et al. (2023), FMCS (2025), Campbell & Lydeard (2026)
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