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Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Hoeh et al. (2009): cladograms
     publication summary | cladograms

Complete Citation

Hoeh, W.R., A.E. Bogan, W.H. Heard & E.G. Chapman. 2009. Palaeoheterodont phylogeny, character evolution, diversity and phylogenetic classification: a reflection on methods of analysis. Malacologia 51(2): 307-317. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 3
  +----------------------------------------------- Neotrigonia_margaritacea   
  |                                                                           
  |                                         +----- Velesunio_ambiguus         
  |                                +--------+                                 
  |                                |        |  +-- Velesunio_angasi           
  |                                |        +--+                              
  |  +-----------------------------+           +-- Lortiella_rugata           
  |  |                             |                                          
  |  |                             |  +----------- Echyridella_menziesii      
--+  |                             |  |                                       
  |  |                             +--+        +-- Hyridella_depressa         
  |  |                                |  +-----+                              
  |  |                                |  |     +-- Hyridella_australis        
  |  |                                +--+                                    
  |  |                                   |  +----- Castalia                   
  |  |                                   +--+                                 
  |  |                                      |  +-- Diplodon_chilensis         
  |  |                                      +--+                              
  |  |                                         +-- Diplodon_gratus            
  +--+                                                                        
     |                                      +----- Chambardia_wahlbergi       
     |                                      |                                 
     |                             +--------+----- Aspatharia_rugifera        
     |                             |        |                                 
     |                             |        |  +-- Mutela_rostrata            
     |  +--------------------------+        +--+                              
     |  |                          |           +-- Mutela_dubia               
     |  |                          |                                          
     |  |                          |  +----------- Etheria_elliptica          
     |  |                          +--+                                       
     |  |                             |  +-------- Mulleria_rivoli            
     |  |                             +--+                                    
     |  |                                |  +----- Monocondylaea_minuana      
     +--+                                +--+                                 
        |                                   |  +-- Anodontites_trigona        
        |                                   +--+                              
        |                                      +-- Anodontites_guanarensis    
        |                                                                     
        |                                      +-- Margaritifera_margaritifera
        |  +-----------------------------------+                              
        |  |                                   +-- Cumberlandia_monodonta     
        |  |                                                                  
        +--+  +----------------------------------- Coelatura_aegyptiaca       
           |  |                                                               
           |  |  +-------------------------------- Pseudomulleria_dalyi       
           |  |  |                                                            
           +--+  |                             +-- Unio_pictorum              
              |  |                    +--------+                              
              |  |                    |        +-- Unio_caffer                
              |  |  +-----------------+                                       
              +--+  |                 |  +-------- Lasmigona_costata          
                 |  |                 +--+                                    
                 |  |                    |  +----- Alasmidonta_marginata      
                 |  |                    +--+                                 
                 |  |                       |  +-- Strophitus_undulatus       
                 |  |                       +--+                              
                 +--+                          +-- Pyganodon_grandis          
                    |                                                         
                    |  +-------------------------- Lens_contradens            
                    |  |                                                      
                    |  |                    +----- Gonidea_angulata           
                    |  |  +-----------------+                                 
                    |  |  |                 |  +-- Pilsbryoconcha_exilis      
                    +--+  |                 +--+                              
                       |  |                    +-- Pseudodon_vondembuschianus 
                       |  |                                                   
                       |  |           +----------- Amblema_plicata            
                       |  |           |                                       
                       +--+           |     +----- Obliquaria_reflexa         
                          |  +--------+  +--+                                 
                          |  |        |  |  |  +-- Quadrula_verrucosa         
                          |  |        |  |  +--+                              
                          |  |        +--+     +-- Quadrula_quadrula          
                          |  |           |                                    
                          |  |           |  +----- Eurynia_dilatata           
                          +--+           +--+                                 
                             |              |  +-- Pleurobema_sintoxia        
                             |              +--+                              
                             |                 +-- Fusconaia_flava            
                             |                                                
                             |  +----------------- Ptychobranchus_fasciolaris 
                             +--+                                             
                                |  +-------------- Truncilla_truncata         
                                +--+                                          
                                   |  +----------- Ortmanniana_ligamentina    
                                   +--+                                       
                                      |  +-------- Lampsilis_cardium          
                                      +--+                                    
                                         |  +----- Cambarunio_iris            
                                         +--+                                 
                                            |  +-- Ligumia_recta              
                                            +--+                              
                                               +-- Epioblasma_triquetra       
Cladogram Figure 4
  +-------------------------------------------------- Neotrigonia_margaritacea   
  |                                                                              
  |                                            +----- Velesunio_ambiguus         
  |                                   +--------+                                 
  |                                   |        |  +-- Velesunio_angasi           
  |                                   |        +--+                              
  |  +--------------------------------+           +-- Lortiella_rugata           
  |  |                                |                                          
  |  |                                |  +----------- Echyridella_menziesii      
--+  |                                |  |                                       
  |  |                                +--+        +-- Hyridella_depressa         
  |  |                                   |  +-----+                              
  |  |                                   |  |     +-- Hyridella_australis        
  |  |                                   +--+                                    
  |  |                                      |  +----- Castalia                   
  |  |                                      +--+                                 
  |  |                                         |  +-- Diplodon_chilensis         
  |  |                                         +--+                              
  |  |                                            +-- Diplodon_gratus            
  +--+                                                                           
     |                                            +-- Etheria_elliptica          
     |                                   +--------+                              
     |                                   |        +-- Mulleria_rivoli            
     |                                   |                                       
     |                                   |     +----- Monocondylaea_minuana      
     |  +--------------------------------+  +--+                                 
     |  |                                |  |  |  +-- Anodontites_trigona        
     |  |                                |  |  +--+                              
     |  |                                |  |     +-- Anodontites_guanarensis    
     |  |                                +--+                                    
     |  |                                   |     +-- Mutela_rostrata            
     |  |                                   |  +--+                              
     +--+                                   |  |  +-- Mutela_dubia               
        |                                   +--+                                 
        |                                      |  +-- Chambardia_wahlbergi       
        |                                      +--+                              
        |                                         +-- Aspatharia_rugifera        
        |                                                                        
        |  +----------------------------------------- Coelatura_aegyptiaca       
        |  |                                                                     
        +--+  +-------------------------------------- Pseudomulleria_dalyi       
           |  |                                                                  
           |  |                                   +-- Margaritifera_margaritifera
           +--+  +--------------------------------+                              
              |  |                                +-- Cumberlandia_monodonta     
              |  |                                                               
              |  |                                +-- Pilsbryoconcha_exilis      
              +--+  +-----------------------------+                              
                 |  |                             +-- Pseudodon_vondembuschianus 
                 |  |                                                            
                 |  |  +----------------------------- Lens_contradens            
                 +--+  |                                                         
                    |  |                          +-- Unio_pictorum              
                    |  |                 +--------+                              
                    |  |                 |        +-- Unio_caffer                
                    |  |  +--------------+                                       
                    +--+  |              |  +-------- Lasmigona_costata          
                       |  |              +--+                                    
                       |  |                 |  +----- Alasmidonta_marginata      
                       |  |                 +--+                                 
                       |  |                    |  +-- Strophitus_undulatus       
                       |  |                    +--+                              
                       +--+                       +-- Pyganodon_grandis          
                          |                                                      
                          |  +----------------------- Gonidea_angulata           
                          |  |                                                   
                          |  |                    +-- Eurynia_dilatata           
                          |  |                    |                              
                          |  |              +-----+-- Pleurobema_sintoxia        
                          +--+              |     |                              
                             |  +-----------+     +-- Fusconaia_flava            
                             |  |           |                                    
                             |  |           |  +----- Obliquaria_reflexa         
                             |  |           +--+                                 
                             +--+              |  +-- Quadrula_verrucosa         
                                |              +--+                              
                                |                 +-- Quadrula_quadrula          
                                |                                                
                                |  +----------------- Ptychobranchus_fasciolaris 
                                +--+                                             
                                   |  +-------------- Truncilla_truncata         
                                   +--+                                          
                                      |  +----------- Ortmanniana_ligamentina    
                                      +--+                                       
                                         |  +-------- Lampsilis_cardium          
                                         +--+                                    
                                            |  +----- Cambarunio_iris            
                                            +--+                                 
                                               |  +-- Ligumia_recta              
                                               +--+                              
                                                  +-- Epioblasma_triquetra       
Cladogram Figure 5
  +-------------------------------- Neotrigonia_margaritacea   
  |                                                            
  |  +----------------------------- Coelatura_aegyptiaca       
  |  |                                                         
  |  |                          +-- Margaritifera_margaritifera
  |  |     +--------------------+                              
  |  |     |                    +-- Cumberlandia_monodonta     
  |  |     |                                                   
--+  |     |                 +----- Velesunio_ambiguus         
  |  |     |        +--------+                                 
  |  |     |        |        |  +-- Velesunio_angasi           
  |  |     |        |        +--+                              
  |  |  +--+  +-----+           +-- Lortiella_rugata           
  |  |  |  |  |     |                                          
  |  |  |  |  |     |  +----------- Echyridella_menziesii      
  |  |  |  |  |     |  |                                       
  +--+  |  |  |     +--+        +-- Hyridella_depressa         
     |  |  |  |        |  +-----+                              
     |  |  |  |        |  |     +-- Hyridella_australis        
     |  |  |  |        +--+                                    
     |  |  +--+           |  +----- Castalia                   
     |  |     |           +--+                                 
     |  |     |              |  +-- Diplodon_chilensis         
     |  |     |              +--+                              
     |  |     |                 +-- Diplodon_gratus            
     |  |     |                                                
     |  |     |  +----------------- Pseudomulleria_dalyi       
     |  |     |  |                                             
     |  |     +--+  +-------------- Mulleria_rivoli            
     |  |        |  |                                          
     |  |        +--+  +----------- Etheria_elliptica          
     +--+           |  |                                       
        |           |  |     +----- Monocondylaea_minuana      
        |           +--+  +--+                                 
        |              |  |  |  +-- Anodontites_trigona        
        |              |  |  +--+                              
        |              |  |     +-- Anodontites_guanarensis    
        |              +--+                                    
        |                 |     +-- Mutela_rostrata            
        |                 |  +--+                              
        |                 |  |  +-- Mutela_dubia               
        |                 +--+                                 
        |                    |  +-- Chambardia_wahlbergi       
        |                    +--+                              
        |                       +-- Aspatharia_rugifera        
        |                                                      
        |                       +-- Pilsbryoconcha_exilis      
        |  +--------------------+                              
        |  |                    +-- Pseudodon_vondembuschianus 
        |  |                                                   
        |  |  +-------------------- Lens_contradens            
        +--+  |                                                
           |  |                 +-- Unio_pictorum              
           |  |        +--------+                              
           |  |        |        +-- Unio_caffer                
           +--+  +-----+                                       
              |  |     |  +-------- Lasmigona_costata          
              |  |     +--+                                    
              |  |        |  +----- Alasmidonta_marginata      
              |  |        +--+                                 
              |  |           |  +-- Strophitus_undulatus       
              +--+           +--+                              
                 |              +-- Pyganodon_grandis          
                 |                                             
                 |  +-------------- Gonidea_angulata           
                 |  |                                          
                 |  |        +----- Obliquaria_reflexa         
                 |  |  +-----+                                 
                 +--+  |     |  +-- Quadrula_verrucosa         
                    |  |     +--+                              
                    |  |        +-- Quadrula_quadrula          
                    |  |                                       
                    +--+  +-------- Amblema_plicata            
                       |  |                                    
                       |  |  +----- Eurynia_dilatata           
                       |  +--+                                 
                       |  |  |  +-- Pleurobema_sintoxia        
                       +--+  +--+                              
                          |     +-- Fusconaia_flava            
                          |                                    
                          |  +----- Truncilla_truncata         
                          |  |                                 
                          |  +----- Ptychobranchus_fasciolaris 
                          +--+                                 
                             |  +-- Ortmanniana_ligamentina    
                             |  |                              
                             |  +-- Cambarunio_iris            
                             +--+                              
                                +-- Ligumia_recta              
                                |                              
                                +-- Epioblasma_triquetra       
 
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