The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina genus Arcidens Simpson, 1900: cladomics
2 Recent valid species (3 total species) Cladomics Summarynumber of species = 2 (3 total)
analyzed in 13 cladograms in 9 publications
maximum number of terminals in an analysis =
2 (mean = 1.62, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 8
maximum number of included terminals = 2, (mean =
2 median = 2)
proportion of cladograms that support basic monophyly
= 0.88
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 31 | 14 | -- | +----------------- Utterbackiana
+--+
| | +-------------- Alasminota
| +--+
| | +----------- Pseudodontoideus
| +--+
| | +-------- Lasmigona
| +--+
| | +----- Pterosyna
| +--+
| | +-- Arcidens_confragosus **
--+ +--+
| +-- Arcidens_wheeleri **
|
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
| +-----+
| | | +-- Ligodonta
| | +--+
| | +-- Alasmidonta
+--------+
| +-- Anodontoides
| +-----+
| | +-- Strophitus
+--+
| +----- Sulcularia
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Whelan et al., 2023 [Publ 4287], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 31 | 13 | -- | +-- Utterbackia
+--------------+
| +-- Pyganodon
|
| +----------- Utterbackiana
| |
| +--+ +-- Lasmigona
--+ | | +-----+
| | | | +-- Pseudodontoideus
| | +--+
| | | +----- Pterosyna
| | +--+
| | | +-- Arcidens_confragosus **
+--+ +--+
| +-- Arcidens_wheeleri **
|
| +----- Sulcularia
| +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+-----+
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Alasminota
+--+
+-- Alasmidonta
Whelan et al., 2023 [Publ 4287], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 25 | 11 | -- | +-------------------- Utterbackiana
|
| +-- Utterbackia
--+ +--------------+
| | +-- Pyganodon
| |
| | +-------- Alasmidonta
+--+ +-----+
| | | +----- Pseudodontoideus
| | +--+
| | | +-- Arcidens_confragosus **
+--+ +--+
| +-- Arcidens_wheeleri **
|
| +----------- Alasminota
| |
+--+ +-- Anodontoides
| +-----+
| | +-- Strophitus
+--+
| +----- Sulcularia
+--+
| +-- Platynaias
+--+
+-- Prolasmidonta
Whelan et al., 2023 [Publ 4287], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 25 | 23 | 10 | -- | +-------------------- Sulcularia
|
| +----------- Utterbackiana
| +-----+
--+ | | +-------- Alasminota
| | +--+
| | | +----- Pseudodontoideus
| | +--+
+--+ | +-- Arcidens_confragosus **
| +--+
| +-- Arcidens_wheeleri **
|
| +-------------- Alasmidonta
+--+
| +----------- Platynaias
+--+
| +-------- Strophitus
+--+
| +----- Pyganodon
+--+
| +-- Utterbackia
+--+
+-- Anodontoides
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 628 | 34 | 13 | -- | +----------- Pseudodontoideus
--+
| +-------- Lasmigona
+--+
| +----- Pterosyna
+--+
| +-- Arcidens_confragosus **
+--+
+-- Arcidens_wheeleri **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 20 | 0 | 2 | 103 | 14 | 9 | -- | +-- Alasmidonta_marginata
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata
|
+-- Beringiana
|
+-- Brachyanodon_kennerlyi
|
+-- Brachyanodon_nuttalliana
|
+-- Brachyanodon_oregonensis
|
+-- Arcidens_confragosus **
|
+-- Arcidens_wheeleri **
|
+-- Pterosyna
|
--+-- Platynaias_compressa
|
+-- Lasmigona
|
+-- Platynaias_decorata
|
+-- Platynaias_subviridis
|
+-- Pyganodon_grandis
|
+-- Pyganodon_lacustris
|
+-- Pseudodontoideus
|
+-- Strophitus
|
+-- Utterbackia_imbecillis
|
+-- Utterbackia_peninsularis__1
|
+-- Utterbackia_peninsularis__2
Inoue et al., 2014 [Publ 3635], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 12 | 8 | -- | +----------- Pseudodontoideus
+--+
| | +-------- Lasmigona
| +--+
| | +----- Pterosyna
| +--+
--+ | +-- Arcidens_confragosus **
| +--+
| +-- Arcidens_wheeleri **
|
| +-------- Strophitus
| |
+-----+ +-- Pyganodon
| +--+
| | +-- Utterbackia
+--+
| +-- Platynaias
+--+
+-- Alasmidonta
Inoue et al., 2014 [Publ 3635], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 5 | 2 | -- | +-------- Pyganodon_grandis
|
+-------- Pyganodon_cataracta
|
--+-------- Pyganodon_fragilis
|
+-------- Pyganodon
|
| +----- Platynaias
+--+
| +-- Arcidens_confragosus **
+--+
+-- Arcidens_wheeleri **
Hewitt et al., 2018 [Publ 3758], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 5 | 5 | -- | +-------- Pyganodon
|
| +-- Platynaias
--+-----+
| +-- Strophitus
|
| +----- Lasmigona
+--+
| +-- Arcidens_wheeleri **
+--+
+-- Pterosyna
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 8 | 8 | -- | +-------------- Strophitus
|
| +----- Pterosyna
--+ +-----+
| | | +-- Lasmigona
| | +--+
+--+ +-- Arcidens_confragosus **
|
| +-------- Utterbackia
| |
+--+-------- Alasmidonta
|
| +----- Pyganodon
+--+
| +-- Anodontoides
+--+
+-- Platynaias
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 0 | 0 | -- | -- Arcidens_confragosus **
Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- | -- Arcidens_confragosus **
Pieri et al., 2018 [Publ 3867], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | -- Arcidens_confragosus **
|
