The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina genus Lasmigona Rafinesque, 1831: cladomics 1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total) analyzed in 63 cladograms in 39 publications maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 32 | 14 | -- |
+-------------- Utterbackiana
+--+
| | +----------- Alasminota
| +--+
| | +-------- Pseudodontoideus
| +--+
| | +----- Lasmigona_costata **
| +--+
| | +-- Pterosyna
| +--+
--+ +-- Arcidens
|
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
| +-----+
| | | +-- Ligodonta
| | +--+
| | +-- Alasmidonta
+-----+
| +-- Anodontoides
| +-----+
| | +-- Strophitus
+--+
| +----- Sulcularia
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Boyer et al., 2011 [Publ 2718], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 5 | 5 | -- |
+----- Alasmidonta
|
+----- Pyganodon
|
--+ +-- Pterosyna
+--+
| +-- Lasmigona_costata **
|
| +-- Platynaias
+--+
+-- Strophitus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 4 | 3 | -- |
+-- Alasmidonta
+--+
| +-- Lasmigona_costata **
--+
| +-- Strophitus
+--+
+-- Pyganodon
Wu et al., 2023 [Publ 4296], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 2 | -- |
+----- Lasmigona_costata **
--+
| +-- Strophitus
+--+
+-- Pyganodon
Zanatta et al., 2007 [Publ 2698], fig. 1B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 13 | 5 | -- |
+-- Lasmigona_costata **
|
--+-- Platynaias
|
+-- Pterosyna
Lydeard et al., 1996 [Publ 2656], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 5 | 4 | -- |
+-- Utterbackia
+-----+
| +-- Utterbackiana
--+
| +----- Strophitus
+--+
| +-- Lasmigona_costata **
+--+
+-- Alasmidonta
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 35 | 13 | -- |
+-------- Pseudodontoideus
--+
| +----- Lasmigona_costata **
+--+
| +-- Pterosyna
+--+
+-- Arcidens
Chen et al., 2025 [Publ 4487], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Campbell et al., 2005 [Publ 2660], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 3 | 3 | -- |
+-- Lasmigona_costata **
--+
+-- Pseudodontoideus
Boyer et al., 2011 [Publ 2718], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 8 | 8 | -- |
+-------------- Strophitus
|
| +----- Pterosyna
--+ +-----+
| | | +-- Lasmigona_costata **
| | +--+
+--+ +-- Arcidens
|
| +-------- Utterbackia
| |
+--+-------- Alasmidonta
|
| +----- Pyganodon
+--+
| +-- Anodontoides
+--+
+-- Platynaias
Amaro et al., 2017 [Publ 3791], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 2 | 2 | -- |
Krebs et al., 2003 [Publ 2691], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 8 | 7 | -- |
+-- Pterosyna
--+
+-- Lasmigona_costata **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 4 | 4 | -- |
+----- Pyganodon
|
+----- Simpsonaias
--+
+----- Strophitus
|
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Liu et al., 2023 [Publ 4310], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 2 | 2 | -- |
Hoeh, 1990 [Publ 2717], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 12 | 5 | -- |
+-- Pterosyna
|
--+-- Platynaias
|
+-- Lasmigona_costata **
Lydeard et al., 1996 [Publ 2656], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 4 | 4 | -- |
+----- Strophitus
--+
| +-- Lasmigona_costata **
+--+
+-- Alasmidonta
Dai et al., 2025 [Publ 4440], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Bolotov et al., 2018 [Publ 3826], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 81 | 4 | 4 | -- |
+----------- Pyganodon
--+
| +-------- Simpsonaias
+--+
| +----- Strophitus
+--+
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
King et al., 1999 [Publ 2699], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 8 | 5 | -- |
+----------- Pyganodon
|
--+ +-- Lasmigona_costata **
| +-----+
| | +-- Pterosyna
+--+
| +----- Strophitus
+--+
| +-- Prolasmidonta
+--+
+-- Platynaias
Huang et al., 2002 [Publ 2657], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----------- Utterbackiana
--+
| +-------- Utterbackia
+--+
| +----- Strophitus
+--+
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Hoeh et al., 2009 [Publ 2716], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 3 | 3 | -- |
+-------- Lasmigona_costata **
--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Pyganodon
Bolotov et al., 2017 [Publ 3761], fig. S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 4 | 4 | -- |
+----- Pyganodon
|
+----- Simpsonaias
--+
+----- Strophitus
|
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Amaro et al., 2017 [Publ 3791], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 4 | 3 | -- |
+-- Lasmigona_costata **
+--+
| +-- Platynaias
--+
| +-- Pyganodon
+--+
+-- Utterbackia
Dai et al., 2025 [Publ 4440], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 15 | 9 | -- |
Bolotov et al., 2017 [Publ 3761], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 4 | 4 | -- |
+-- Lasmigona_costata **
+-----+
| +-- Alasmidonta
--+
| +----- Simpsonaias
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Hoeh, 1990 [Publ 2717], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 12 | 5 | -- |
+-- Lasmigona_costata **
--+
+-- Utterbackiana
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 2 | 1 | -- |
+-- Lasmigona_costata **
--+
+-- Platynaias
Hoeh, 1990 [Publ 2717], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 12 | 5 | -- |
+-- Pterosyna
|
--+-- Platynaias
|
+-- Lasmigona_costata **
Huang et al., 2002 [Publ 2657], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----- Utterbackia
|
--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Whelan et al., 2011 [Publ 3380], fig. 1-MP | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Pyganodon
--+
+-- Lasmigona_costata **
Graf & Cummings, 2006 [Publ 1968], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 3 | 3 | -- |
+-------- Lasmigona_costata **
--+
| +----- Alasmidonta
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Krebs, 2004 [Publ 2692], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- |
+-- Lasmigona_costata **
--+
+-- Pyganodon
Huang et al., 2019 [Publ 3843], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 6 | 5 | -- |
Zanatta et al., 2007 [Publ 2698], fig. 1A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 13 | 5 | -- |
+-- Platynaias
|
--+-- Lasmigona_costata **
|
+-- Pterosyna
King et al., 1999 [Publ 2699], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 5 | 4 | -- |
+-- Pterosyna
+--+
| +-- Lasmigona_costata **
--+
| +-- Prolasmidonta
| |
+--+-- Strophitus
|
+-- Platynaias
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 3 | 3 | -- |
+----- Lasmigona_costata **
|
--+ +-- Strophitus
| |
+--+-- Alasmidonta
|
+-- Pyganodon
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 4 | 3 | -- |
+-- Alasmidonta
+--+
| +-- Lasmigona_costata **
--+
| +-- Strophitus
+--+
+-- Pyganodon
Krebs, 2004 [Publ 2692], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 1 | 1 | -- |
+-- Lasmigona_costata **
--+
+-- Pyganodon
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Simpsonaias
--+
| +----- Strophitus
+--+
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Huang et al., 2019 [Publ 3843], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 3 | 2 | -- |
+----- Lasmigona_costata **
--+
| +-- Pyganodon
+--+
+-- Utterbackia
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 4 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Simpsonaias
--+
| +----- Strophitus
+--+
| +-- Alasmidonta
+--+
+-- Lasmigona_costata **
Wu et al., 2021 [Publ 4366], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 2 | 2 | -- |
Liu et al., 2023 [Publ 4310], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 2 | 2 | -- |
+----- Lasmigona_costata **
--+
| +-- Strophitus
+--+
+-- Pyganodon
Chen et al., 2023 [Publ 4302], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Huang et al., 2019 [Publ 3843], fig. S3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 6 | 5 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Inoue et al., 2014 [Publ 3635], fig. 3A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 13 | 8 | -- |
+-------- Pseudodontoideus
+--+
| | +----- Lasmigona_costata **
| +--+
| | +-- Pterosyna
--+ +--+
| +-- Arcidens
|
| +-------- Strophitus
| |
+--+ +-- Pyganodon
| +--+
| | +-- Utterbackia
+--+
| +-- Platynaias
+--+
+-- Alasmidonta
Metzger et al., 2018 [Publ 3952], fig. 1C | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
Whelan et al., 2023 [Publ 4287], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 32 | 13 | -- |
+-- Utterbackia
+-----------+
| +-- Pyganodon
|
| +-------- Utterbackiana
| |
--+ +--+ +-- Lasmigona_costata **
| | | +--+
| | | | +-- Pseudodontoideus
| | +--+
| | | +-- Pterosyna
| | +--+
+--+ +-- Arcidens
|
| +----- Sulcularia
| +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+--+
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Alasminota
+--+
+-- Alasmidonta
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 4 | -- |
+-- Alasmidonta
+-----+
| +-- Lasmigona_costata **
--+
| +----- Pyganodon
+--+
| +-- Strophitus
+--+
+-- Simpsonaias
Bolotov et al., 2017 [Publ 3764], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 123 | 4 | 4 | -- |
Hoeh et al., 2009 [Publ 2716], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 3 | 3 | -- |
+-------- Lasmigona_costata **
--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Pyganodon
Chen et al., 2025 [Publ 4487], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Hoeh et al., 2009 [Publ 2716], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 3 | 3 | -- |
+-------- Lasmigona_costata **
--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Pyganodon
Wu et al., 2021 [Publ 4366], fig. 2_I | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 2 | 2 | -- |
Huang et al., 2019 [Publ 3843], fig. S4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 6 | 5 | -- |
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 1 | 1 | -- |
+-- Pyganodon
--+
+-- Lasmigona_costata **
Chen et al., 2023 [Publ 4302], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta
--+
+-- Lasmigona_costata **
Chase et al., 2018 [Publ 3847], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 10 | 6 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
|
--+ +-- Lasmigona_costata **
| +-----+
| | +-- Pterosyna
+--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Platynaias
Breton et al., 2011 [Publ 3376], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 10 | 6 | -- |
+-- Pterosyna
+--------+
| +-- Lasmigona_costata **
--+
| +-------- Strophitus
| |
+--+ +-- Platynaias
| +--+
| | +-- Alasmidonta
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Whelan et al., 2011 [Publ 3380], fig. 1-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Pyganodon
--+
+-- Lasmigona_costata **
Hewitt et al., 2018 [Publ 3758], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 5 | 5 | -- |
+-------- Pyganodon
|
| +-- Platynaias
--+-----+
| +-- Strophitus
|
| +----- Lasmigona_costata **
+--+
| +-- Arcidens
+--+
+-- Pterosyna
Campbell et al., 2005 [Publ 2660], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 3 | 3 | -- |
+----- Pyganodon
--+
| +-- Lasmigona_costata **
+--+
+-- Pseudodontoideus
|