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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Gonideini

genus Inversidens Haas, 1911: cladomics
     3 Recent valid species (3 total species)

Cladomics Summary

number of species = 3 (3 total)
analyzed in 24 cladograms in 12 publications
maximum number of terminals in an analysis = 2 (mean = 1.29, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 2, (mean = 2 median = 2)
proportion of cladograms that support basic monophyly = 1

Dai et al., 2023 [Publ 4285], fig. 3-BI

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
220243129--
  +-- Inversidens_brandtii **    
--+                           
  +-- Inversidens_rentianensis **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
22022266--
           +-- Ptychorhynchus          
  +--------+                           
  |        +-- Parvasolenaia           
  |                                    
--+        +-- Inversidens_brandtii **    
  |  +-----+                           
  |  |     +-- Inversidens_rentianensis **
  +--+                                 
     |  +----- Gonidea                 
     +--+                              
        |  +-- Microcondylaea          
        +--+                           
           +-- Leguminaia

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
220243129--
  +-- Inversidens_brandtii **    
--+                           
  +-- Inversidens_rentianensis **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
22023577--
  +-- Inversidens_rentianensis **
--+                           
  +-- Inversidens_brandtii **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
22023577--
                 +-- Inversidens_brandtii **    
  +--------------+                           
  |              +-- Inversidens_rentianensis **
--+                                          
  |  +-------------- Sinosolenaia            
  |  |                                       
  +--+           +-- Gonidea                 
     |  +--------+                           
     |  |        +-- Microcondylaea          
     +--+                                    
        |  +-------- Koreosolenaia           
        +--+                                 
           |  +----- Parvasolenaia           
           +--+                              
              |  +-- Postolata               
              +--+                           
                 +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
22023577--
  +-- Inversidens_brandtii **    
--+                           
  +-- Inversidens_rentianensis **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
22023577--
  +-- Inversidens_brandtii **    
--+                           
  +-- Inversidens_rentianensis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019076--
        +----- Gonidea             
  +-----+                          
  |     |  +-- Leguminaia          
  |     +--+                       
--+        +-- Microcondylaea      
  |                                
  |  +-------- Inversidens_brandtii **
  +--+                             
     |  +----- Sinosolenaia        
     +--+                          
        |  +-- Parvasolenaia       
        +--+                       
           +-- Ptychorhynchus

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016722--
-- Inversidens_brandtii **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
11011196--
  +-------------- Sinosolenaia_carinata   
  |                                       
  |        +----- Inversidens_rentianensis **
--+  +-----+                              
  |  |     |  +-- Gonidea                 
  |  |     +--+                           
  +--+        +-- Leguminaia              
     |                                    
     |  +-------- Sinosolenaia_oleivora   
     +--+                                 
        |  +----- Postolata               
        +--+                              
           |  +-- Obovalis                
           +--+                           
              +-- Ptychorhynchus

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011411--
-- Inversidens **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019076--
        +----- Sinosolenaia        
  +-----+                          
  |     |  +-- Ptychorhynchus      
  |     +--+                       
--+        +-- Parvasolenaia       
  |                                
  |  +-------- Inversidens_brandtii **
  +--+                             
     |  +----- Gonidea             
     +--+                          
        |  +-- Microcondylaea      
        +--+                       
           +-- Leguminaia

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016722--
-- Inversidens_brandtii **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013233--
  +-- Inversidens_brandtii **
--+                       
  +-- Sinosolenaia

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016722--
  +----- Inversidens_brandtii **
--+                          
  |  +-- Obovalis            
  +--+                       
     +-- Ptychorhynchus

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1101911--
-- Inversidens_brandtii **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013333--
  +-------- Sinosolenaia        
  |                             
--+  +----- Inversidens_brandtii **
  |  |                          
  +--+  +-- Obovalis_omiensis__1
     |  |                       
     +--+-- Obovalis_omiensis__2
        |                       
        +-- Ptychorhynchus

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012011--
-- Inversidens_brandtii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019064--
  +-------- Inversidens_rentianensis **
  |                                 
--+     +-- Parvasolenaia           
  |  +--+                           
  |  |  +-- Ptychorhynchus          
  +--+                              
     |  +-- Microcondylaea          
     +--+                           
        +-- Sinosolenaia

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011111--
-- Inversidens_brandtii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019064--
  +-------- Inversidens_rentianensis **
  |                                 
--+     +-- Parvasolenaia           
  |  +--+                           
  |  |  +-- Ptychorhynchus          
  +--+                              
     |  +-- Microcondylaea          
     +--+                           
        +-- Sinosolenaia

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
110127117--
  +----------------- Inversidens_rentianensis **
  |                                          
--+              +-- Microcondylaea          
  |  +-----------+                           
  |  |           +-- Sinosolenaia            
  +--+                                       
     |  +----------- Parvasolenaia           
     +--+                                    
        |  +-------- Koreosolenaia           
        +--+                                 
           |  +----- Ptychorhynchus          
           +--+                              
              |  +-- Postolata               
              +--+                           
                 +-- Cosmopseudodon

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1101120118--
     +-------- Sinosolenaia        
  +--+                             
  |  |  +----- Gonidea             
  |  +--+                          
  |     |  +-- Leguminaia          
--+     +--+                       
  |        +-- Microcondylaea      
  |                                
  |  +-------- Inversidens_brandtii **
  |  |                             
  +--+     +-- Obovalis            
     |  +--+                       
     |  |  +-- Ptychorhynchus      
     +--+                          
        |  +-- Parvasolenaia       
        +--+                       
           +-- Koreosolenaia

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011411--
-- Inversidens **
 
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