The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Gonideini genus Inversidens Haas, 1911: cladomics
5 Recent valid species (5 total species) Cladomics Summarynumber of species = 5 (5 total)
analyzed in 29 cladograms in 15 publications
maximum number of terminals in an analysis =
3 (mean = 1.38, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 10
maximum number of included terminals = 3, (mean =
2.1 median = 2)
proportion of cladograms that support basic monophyly
= 1
Wu et al., 2026 [Publ 4811], fig. 3 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 1 | 1 | 0 | +-------- Gonidea
--+
| +----- Inversidens_trisulcata **
+--+
| +-- Inversidens_brandtii **
+--+
+-- Inversidens_rentianensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 629 | 19 | 12 | -- | +-------------- Sinosolenaia_carinata
|
| +----- Gonidea
--+ +-----+
| | | +-- Microcondylaea
| | +--+
| | +-- Leguminaia
+--+
| +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
+--+
| +----- Sinosolenaia_recognita
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Dai et al., 2023 [Publ 4285], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 12 | 9 | -- | +-- Inversidens_brandtii **
--+
+-- Inversidens_rentianensis **
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 22 | 6 | 6 | -- | +-- Ptychorhynchus
+--------+
| +-- Parvasolenaia
|
--+ +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
Dai et al., 2023 [Publ 4285], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 12 | 9 | -- | +-- Inversidens_brandtii **
--+
+-- Inversidens_rentianensis **
Wu et al., 2024 [Publ 4350], fig. 6-7 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 7 | 7 | -- | +-- Inversidens_brandtii **
+--------------+
| +-- Inversidens_rentianensis **
--+
| +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2024 [Publ 4350], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 7 | 7 | -- | +-- Inversidens_brandtii **
--+
+-- Inversidens_rentianensis **
Wu et al., 2024 [Publ 4350], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 7 | 7 | -- | +-- Inversidens_brandtii **
--+
+-- Inversidens_rentianensis **
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 93 | 11 | 7 | -- | +-- Inversidens_rentianensis **
+-----------------+
| +-- Inversidens_trisulcata **
--+
| +----------------- Sinosolenaia
+--+
| +-------------- Parvasolenaia
+--+
| +----------- Pseudopostolata
+--+
| +-------- Koreosolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Postolata
Wu et al., 2024 [Publ 4350], fig. 5c iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 7 | 7 | -- | +-- Inversidens_rentianensis **
--+
+-- Inversidens_brandtii **
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 3 | 3 | -- | +-- Inversidens_brandtii **
--+
+-- Sinosolenaia
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 6 | 5 | -- | +----------- Inversidens_rentianensis **
|
--+ +-- Microcondylaea
| +-----+
| | +-- Sinosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 140 | 10 | 7 | -- | +-------------------- Inversidens_rentianensis **
--+
| +----------------- Microcondylaea
+--+
| +-------------- Sinosolenaia
+--+
| +----------- Koreosolenaia
+--+
| +-------- Parvasolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Postolata
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- | +----- Inversidens_brandtii **
--+
| +-- Obovalis
+--+
+-- Ptychorhynchus
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 11 | 8 | -- | +-------- Sinosolenaia
+--+
| | +----- Gonidea
| +--+
| | +-- Leguminaia
--+ +--+
| +-- Microcondylaea
|
| +-------- Inversidens_brandtii **
| |
+--+ +-- Obovalis
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 6 | 4 | -- | +-------- Inversidens_rentianensis **
|
--+ +-- Parvasolenaia
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Microcondylaea
+--+
+-- Sinosolenaia
Liu et al., 2024 [Publ 4380], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 9 | 6 | -- | +-------------- Sinosolenaia_carinata
|
| +----- Inversidens_rentianensis **
--+ +-----+
| | | +-- Gonidea
| | +--+
+--+ +-- Leguminaia
|
| +-------- Sinosolenaia_oleivora
+--+
| +----- Postolata
+--+
| +-- Obovalis
+--+
+-- Ptychorhynchus
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- | -- Inversidens_brandtii **
Zhang et al., 2024 [Publ 4397], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 10 | 7 | -- | +----------------- Inversidens_rentianensis **
|
--+ +-- Microcondylaea
| +-----------+
| | +-- Sinosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Koreosolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Postolata
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 7 | 6 | -- | +----- Sinosolenaia
+-----+
| | +-- Ptychorhynchus
| +--+
--+ +-- Parvasolenaia
|
| +-------- Inversidens_brandtii **
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 6 | 4 | -- | +-------- Inversidens_rentianensis **
|
--+ +-- Parvasolenaia
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Microcondylaea
+--+
+-- Sinosolenaia
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 7 | 6 | -- | +----- Gonidea
+-----+
| | +-- Leguminaia
| +--+
--+ +-- Microcondylaea
|
| +-------- Inversidens_brandtii **
+--+
| +----- Sinosolenaia
+--+
| +-- Parvasolenaia
+--+
+-- Ptychorhynchus
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | -- Inversidens_brandtii **
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 1 | 1 | -- | -- Inversidens_brandtii **
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 3 | 3 | -- | +-------- Sinosolenaia
|
--+ +----- Inversidens_brandtii **
| |
+--+ +-- Obovalis_omiensis__1
| |
+--+-- Obovalis_omiensis__2
|
+-- Ptychorhynchus
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- | -- Inversidens_brandtii **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | -- Inversidens_brandtii **
|
