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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Anemina Haas, 1969: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 83 cladograms in 56 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Wu et al., 2025 [Publ 4442], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110134327--
  +----------------- Cristaria                  
  |                                             
  |           +----- Beringiana                 
--+  +--------+                                 
  |  |        |  +-- Acudonta                   
  |  |        +--+                              
  |  |           +-- Sinanodonta                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus     
        |                                       
        |  +-------- Pletholophus_honglinhensis 
        +--+                                    
           |  +----- Anemina_arcaeformis **        
           +--+                                 
              |  +-- Amuranodonta               
              +--+                              
                 +-- Buldowskia

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014832--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014455--
  +-- Anemina_arcaeformis **
--+                      
  +-- Amuranodonta

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014332--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014455--
  +-- Anemina_arcaeformis **
--+                      
  +-- Amuranodonta

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019032--
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014622--
  +----- Sinanodonta        
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Cristaria

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013522--
  +----- Cristaria          
--+                         
  |  +-- Sinanodonta        
  +--+                      
     +-- Anemina_arcaeformis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina_arcaeformis **
--+                      
  +-- Buldowskia

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013932--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015222--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
1101511--
  +-- Sinanodonta        
--+                      
  +-- Anemina_arcaeformis **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015252--
  +-- Anemina_arcaeformis **
  |                      
--+-- Cristaria          
  |                      
  +-- Sinanodonta

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
11014031--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11013955--
  +----------- Cristaria          
  |                               
--+        +-- Anemina_arcaeformis **
  |  +-----+                      
  |  |     +-- Amuranodonta       
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Sinanodonta        
        +--+                      
           +-- Beringiana

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014132--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014532--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11013955--
  +-- Anemina_arcaeformis **
--+                      
  +-- Amuranodonta

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015253--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
1101971--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
11013322--
-- Anemina_arcaeformis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119883--
-- Anemina_arcaeformis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017553--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina_arcaeformis **
--+                      
  +-- Buldowskia

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119883--
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011221--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015532--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012822--
  +----- Cristaria          
--+                         
  |  +-- Sinanodonta        
  +--+                      
     +-- Anemina_arcaeformis **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015333--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina_arcaeformis **
--+                      
  +-- Buldowskia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019453--
     +-- Anemina_arcaeformis **
  +--+                      
  |  +-- Cristaria          
--+                         
  |  +-- Beringiana         
  +--+                      
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013395--
-- Anemina_arcaeformis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019053--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Anemina_arcaeformis **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
11011264--
  +----------- Cristaria          
--+                               
  |  +-------- Anemina_arcaeformis **
  +--+                            
     |  +----- Beringiana         
     +--+                         
        |  +-- Mimo               
        +--+                      
           +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119683--
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
110127267--
  +-------------- Pletholophus       
  |                                  
  |        +----- Beringiana         
--+  +-----+                         
  |  |     |  +-- Mimo               
  |  |     +--+                      
  +--+        +-- Sinanodonta        
     |                               
     |  +-------- Cristaria          
     +--+                            
        |  +----- Anemina_arcaeformis **
        +--+                         
           |  +-- Amuranodonta       
           +--+                      
              +-- Buldowskia

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
11011321--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Kondakov et al., 2018 [Publ 3914], fig. 5

iTtTbiriSoTroTroSiD
1101771--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
110130266--
        +----- Amuranodonta       
  +-----+                         
  |     |  +-- Anemina_arcaeformis **
  |     +--+                      
--+        +-- Buldowskia         
  |                               
  |  +-------- Cristaria          
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
11011832--
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
110155125--
     +----- Anemina_arcaeformis **
  +--+                         
  |  |  +-- Amuranodonta       
  |  +--+                      
--+     +-- Buldowskia         
  |                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017943--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
11014272--
-- Anemina_arcaeformis **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012022--
-- Anemina_arcaeformis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
110130266--
        +----- Amuranodonta       
  +-----+                         
  |     |  +-- Anemina_arcaeformis **
  |     +--+                      
--+        +-- Buldowskia         
  |                               
  |  +-------- Cristaria          
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
1101522--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013132--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012000--
-- Anemina_arcaeformis **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014722--
  +----- Sinanodonta        
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Cristaria

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1101120236--
  +-------- Cristaria          
  |                            
  |  +----- Anemina_arcaeformis **
  +--+                         
--+  |  +-- Amuranodonta       
  |  +--+                      
  |     +-- Buldowskia         
  |                            
  |  +----- Pletholophus       
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
11014472--
-- Anemina_arcaeformis **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
110130266--
  +----------- Cristaria          
  |                               
  |     +----- Amuranodonta       
--+  +--+                         
  |  |  |  +-- Anemina_arcaeformis **
  |  |  +--+                      
  +--+     +-- Buldowskia         
     |                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019032--
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012122--
-- Anemina_arcaeformis **

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
1101971--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Bauer et al., 2021 [Publ 4365], fig. 2

iTtTbiriSoTroTroSiD
1101771--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015122--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Anemina_arcaeformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iTtTbiriSoTroTroSiD
11011083--
  +-------- Sinanodonta        
--+                            
  |  +----- Amuranodonta       
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Buldowskia

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012000--
-- Anemina_arcaeformis **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
11011193--
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019053--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014332--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
110116147--
  +----------- Cristaria          
  |                               
  |     +----- Amuranodonta       
--+  +--+                         
  |  |  |  +-- Anemina_arcaeformis **
  |  |  +--+                      
  |  |     +-- Buldowskia         
  +--+                            
     |     +-- Beringiana         
     |  +--+                      
     |  |  +-- Pletholophus       
     +--+                         
        |  +-- Acudonta           
        +--+                      
           +-- Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019453--
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013585--
     +----- Amuranodonta       
  +--+                         
  |  |  +-- Anemina_arcaeformis **
  |  +--+                      
--+     +-- Buldowskia         
  |                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015232--
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011432--
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta
 
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