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Page last updated
Thu 7 August 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | ... | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Utterbackiana Frierson, 1927: cladomics
     5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 19 cladograms in 11 publications
maximum number of terminals in an analysis = 4 (mean = 2.32, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 10
maximum number of included terminals = 4, (mean = 3.5 median = 4)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
44243729120
              +-- Utterbackia                
  +-----------+                              
  |           +-- Pyganodon                  
  |                                          
  |        +----- Pressodonta                
  |     +--+                                 
  |     |  |  +-- Anodontoides               
  |     |  +--+                              
  |     |     +-- Strophitus                 
--+  +--+                                    
  |  |  |     +-- Prolasmidonta              
  |  |  |  +--+                              
  |  |  |  |  +-- Platynaias                 
  |  |  +--+                                 
  |  |     |  +-- Alasminota                 
  |  |     +--+                              
  |  |        +-- Alasmidonta                
  +--+                                       
     |        +-- Utterbackiana_couperiana **   
     |     +--+                              
     |     |  +-- Utterbackiana_heardi **       
     |  +--+                                 
     |  |  |  +-- Utterbackiana_suborbiculata **
     |  |  +--+                              
     |  |     +-- Utterbackiana_hartfieldorum **
     +--+                                    
        |     +-- Lasmigona_costata          
        |  +--+                              
        |  |  +-- Pseudodontoideus           
        +--+                                 
           |  +-- Lasmigona_complanata       
           +--+                              
              +-- Arcidens

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
44242423110
                    +-- Utterbackiana_heardi **       
                 +--+                              
                 |  +-- Utterbackiana_couperiana **   
  +--------------+                                 
  |              |  +-- Utterbackiana_hartfieldorum **
  |              +--+                              
  |                 +-- Utterbackiana_suborbiculata **
--+                                                
  |                 +-- Utterbackia                
  |  +--------------+                              
  |  |              +-- Pyganodon                  
  |  |                                             
  +--+           +----- Alasmidonta                
     |  +--------+                                 
     |  |        |  +-- Arcidens                   
     |  |        +--+                              
     +--+           +-- Pseudodontoideus           
        |                                          
        |  +----------- Alasminota                 
        |  |                                       
        +--+        +-- Anodontoides               
           |  +-----+                              
           |  |     +-- Strophitus                 
           +--+                                    
              |  +----- Pressodonta                
              +--+                                 
                 |  +-- Platynaias                 
                 +--+                              
                    +-- Prolasmidonta

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
4424121044
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_heardi **       
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
4424121042
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_suborbiculata **
  +--+                                 
     |  +-- Utterbackiana_heardi **       
     +--+                              
        +-- Utterbackiana_couperiana **

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
4424141370
                 +-- Utterbackia                
  +--------------+                              
  |              +-- Pyganodon                  
  |                                             
--+              +-- Anodontoides               
  |  +-----------+                              
  |  |           +-- Strophitus                 
  |  |                                          
  +--+           +-- Alasmidonta                
     |  +--------+                              
     |  |        +-- Alasminota                 
     +--+                                       
        |  +-------- Pseudodontoideus           
        |  |                                    
        +--+     +-- Utterbackiana_couperiana **   
           |  +--+                              
           |  |  +-- Utterbackiana_heardi **       
           +--+                                 
              |  +-- Utterbackiana_hartfieldorum **
              +--+                              
                 +-- Utterbackiana_suborbiculata **

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
44142321103
  +-------------------- Pressodonta                
  |                                                
  |     +-------------- Alasmidonta                
  |  +--+                                          
  |  |  |  +----------- Platynaias                 
--+  |  +--+                                       
  |  |     |  +-------- Strophitus                 
  |  |     +--+                                    
  |  |        |  +----- Pyganodon                  
  |  |        +--+                                 
  +--+           |  +-- Utterbackia                
     |           +--+                              
     |              +-- Anodontoides               
     |                                             
     |           +----- Alasminota                 
     |        +--+                                 
     |        |  |  +-- Arcidens                   
     |        |  +--+                              
     +--------+     +-- Pseudodontoideus           
              |                                    
              |  +----- Utterbackiana_couperiana **   
              |  |                                 
              +--+  +-- Utterbackiana_heardi **       
                 |  |                              
                 +--+-- Utterbackiana_hartfieldorum **
                    |                              
                    +-- Utterbackiana_suborbiculata **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
3324121044
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
315241210416
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana__1 **
  |                                 
--+----- Utterbackiana_couperiana__2 **
  |                                 
  +----- Utterbackiana_suborbiculata **
  |                                 
  |  +-- Lasmigona_costata          
  +--+                              
  |  +-- Utterbackiana_implicata **    
  |                                 
  |  +-- Lasmigona_complanata       
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
3324121044
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
22022544--
     +----- Utterbackia             
  +--+                              
  |  |  +-- Utterbackiana_couperiana **
  |  +--+                           
--+     +-- Utterbackiana **           
  |                                 
  |  +----- Strophitus              
  +--+                              
     |  +-- Lasmigona               
     +--+                           
        +-- Alasmidonta

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012744--
-- Utterbackiana_couperiana **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013344--
  +----- Utterbackia             
  |                              
--+----- Utterbackiana_couperiana **
  |                              
  |  +-- Alasmidonta             
  +--+                           
     +-- Lasmigona

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iTtTbiriSoTroTroSiD
1101300--
-- Utterbackiana_implicata **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
11013700--
-- Utterbackiana_suborbiculata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
11011111--
  +-- Utterbackiana_implicata **
--+                          
  +-- Pyganodon

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013344--
  +----------- Utterbackiana_couperiana **
--+                                    
  |  +-------- Utterbackia             
  +--+                                 
     |  +----- Strophitus              
     +--+                              
        |  +-- Alasmidonta             
        +--+                           
           +-- Lasmigona

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013286--
-- Utterbackiana_couperiana **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Utterbackiana_couperiana **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iTtTbiriSoTroTroSiD
1101300--
-- Utterbackiana_implicata **
 
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