MUSSELpdb | valid gen. Utterbackiana cladomics

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Page last updated
Thu 3 September 2025

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genus Utterbackiana Frierson, 1927: cladomics
5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 22 cladograms in 14 publications
maximum number of terminals in an analysis = 5 (mean = 2.5, median = 2.5)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 13
maximum number of included terminals = 5, (mean = 3.54 median = 4)
proportion of cladograms that support basic monophyly = 0.92

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	626	31	13	0

        +-------- Utterbackiana_implicata **    
        |                                    
  +-----+     +-- Utterbackiana_couperiana **   
  |     |  +--+                              
  |     |  |  +-- Utterbackiana_heardi **       
  |     +--+                                 
  |        |  +-- Utterbackiana_hartfieldorum **
--+        +--+                              
  |           +-- Utterbackiana_suborbiculata **
  |                                          
  |  +----------- Simpsonaias                
  |  |                                       
  |  |        +-- Alasminota                 
  +--+  +-----+                              
     |  |     +-- Alasmidonta                
     |  |                                    
     +--+     +-- Prolasmidonta              
        |  +--+                              
        |  |  +-- Platynaias                 
        +--+                                 
           |  +-- Anodontoides               
           +--+                              
              +-- Strophitus

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	32	29	14	1

                 +-- Utterbackiana_heardi **       
              +--+                              
              |  +-- Utterbackiana_couperiana **   
     +--------+                                 
     |        |  +-- Utterbackiana_hartfieldorum **
     |        +--+                              
  +--+           +-- Utterbackiana_suborbiculata **
  |  |                                          
  |  |  +----------- Alasminota                 
  |  +--+                                       
  |     |  +-------- Pseudodontoideus           
  |     +--+                                    
  |        |  +----- Lasmigona                  
  |        +--+                                 
  |           |  +-- Pterosyna                  
--+           +--+                              
  |              +-- Arcidens                   
  |                                             
  |              +-- Prolasmidonta              
  |           +--+                              
  |           |  +-- Platynaias                 
  |     +-----+                                 
  |     |     |  +-- Ligodonta                  
  |     |     +--+                              
  |     |        +-- Alasmidonta                
  +-----+                                       
        |        +-- Anodontoides               
        |  +-----+                              
        |  |     +-- Strophitus                 
        +--+                                    
           |  +----- Sulcularia                 
           +--+                                 
              |  +-- Pyganodon                  
              +--+                              
                 +-- Utterbackia

Whelan et al., 2023 [Publ 4287], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	37	29	13	1

              +-- Utterbackia                
  +-----------+                              
  |           +-- Pyganodon                  
  |                                          
  |        +----- Sulcularia                 
  |     +--+                                 
  |     |  |  +-- Anodontoides               
  |     |  +--+                              
  |     |     +-- Strophitus                 
--+  +--+                                    
  |  |  |     +-- Prolasmidonta              
  |  |  |  +--+                              
  |  |  |  |  +-- Platynaias                 
  |  |  +--+                                 
  |  |     |  +-- Alasminota                 
  |  |     +--+                              
  |  |        +-- Alasmidonta                
  +--+                                       
     |        +-- Utterbackiana_couperiana **   
     |     +--+                              
     |     |  +-- Utterbackiana_heardi **       
     |  +--+                                 
     |  |  |  +-- Utterbackiana_suborbiculata **
     |  |  +--+                              
     |  |     +-- Utterbackiana_hartfieldorum **
     +--+                                    
        |     +-- Lasmigona                  
        |  +--+                              
        |  |  +-- Pseudodontoideus           
        +--+                                 
           |  +-- Pterosyna                  
           +--+                              
              +-- Arcidens

Smith et al., 2018 [Publ 3790], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	14	13	7	1

                 +-- Utterbackia                
  +--------------+                              
  |              +-- Pyganodon                  
  |                                             
--+              +-- Anodontoides               
  |  +-----------+                              
  |  |           +-- Strophitus                 
  |  |                                          
  +--+           +-- Alasmidonta                
     |  +--------+                              
     |  |        +-- Alasminota                 
     +--+                                       
        |  +-------- Pseudodontoideus           
        |  |                                    
        +--+     +-- Utterbackiana_couperiana **   
           |  +--+                              
           |  |  +-- Utterbackiana_heardi **       
           +--+                                 
              |  +-- Utterbackiana_hartfieldorum **
              +--+                              
                 +-- Utterbackiana_suborbiculata **

Zanatta et al., 2007 [Publ 2698], fig. 1A

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	12	10	5	1

  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_suborbiculata **
  +--+                                 
     |  +-- Utterbackiana_heardi **       
     +--+                              
        +-- Utterbackiana_couperiana **

Whelan et al., 2023 [Publ 4287], fig. 3

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	24	23	11	1

                    +-- Utterbackiana_heardi **       
                 +--+                              
                 |  +-- Utterbackiana_couperiana **   
  +--------------+                                 
  |              |  +-- Utterbackiana_hartfieldorum **
  |              +--+                              
  |                 +-- Utterbackiana_suborbiculata **
--+                                                
  |                 +-- Utterbackia                
  |  +--------------+                              
  |  |              +-- Pyganodon                  
  |  |                                             
  +--+           +----- Alasmidonta                
     |  +--------+                                 
     |  |        |  +-- Arcidens                   
     |  |        +--+                              
     +--+           +-- Pseudodontoideus           
        |                                          
        |  +----------- Alasminota                 
        |  |                                       
        +--+        +-- Anodontoides               
           |  +-----+                              
           |  |     +-- Strophitus                 
           +--+                                    
              |  +----- Sulcularia                 
              +--+                                 
                 |  +-- Platynaias                 
                 +--+                              
                    +-- Prolasmidonta

Zanatta et al., 2007 [Publ 2698], fig. 1B

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	12	10	5	3

  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_heardi **       
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana **

Whelan et al., 2023 [Publ 4287], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	1	4	23	21	10	4

  +-------------------- Sulcularia                 
  |                                                
  |     +-------------- Alasmidonta                
  |  +--+                                          
  |  |  |  +----------- Platynaias                 
--+  |  +--+                                       
  |  |     |  +-------- Strophitus                 
  |  |     +--+                                    
  |  |        |  +----- Pyganodon                  
  |  |        +--+                                 
  +--+           |  +-- Utterbackia                
     |           +--+                              
     |              +-- Anodontoides               
     |                                             
     |           +----- Alasminota                 
     |        +--+                                 
     |        |  |  +-- Arcidens                   
     |        |  +--+                              
     +--------+     +-- Pseudodontoideus           
              |                                    
              |  +----- Utterbackiana_couperiana **   
              |  |                                 
              +--+  +-- Utterbackiana_heardi **       
                 |  |                              
                 +--+-- Utterbackiana_hartfieldorum **
                    |                              
                    +-- Utterbackiana_suborbiculata **

Hoeh, 1990 [Publ 2717], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	2	4	12	10	5	3

  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Hoeh, 1990 [Publ 2717], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	2	4	12	10	5	3

  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Hoeh, 1990 [Publ 2717], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	15	2	4	12	10	5	16

  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta                   
  |                                 
  +----- Brachyanodon               
  |                                 
  +----- Utterbackiana_couperiana__1 **
  |                                 
--+----- Utterbackiana_couperiana__2 **
  |                                 
  +----- Utterbackiana_suborbiculata **
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata **    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	26	0	0	--

  +-- Utterbackiana_couperiana **   
--+                              
  +-- Utterbackiana_hartfieldorum **

Lydeard et al., 1996 [Publ 2656], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	25	4	4	--

     +----- Utterbackia             
  +--+                              
  |  |  +-- Utterbackiana_couperiana **
  |  +--+                           
--+     +-- Utterbackiana **           
  |                                 
  |  +----- Strophitus              
  +--+                              
     |  +-- Lasmigona               
     +--+                           
        +-- Alasmidonta

Pfeiffer et al., 2016 [Publ 3660], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Utterbackiana_suborbiculata **

Turner et al., 2000 [Publ 2722], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Utterbackiana_couperiana **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	0	0	--

-- Utterbackiana_implicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	11	1	1	--

  +-- Utterbackiana_implicata **
--+                          
  +-- Pyganodon

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	4	4	--

  +----------- Utterbackiana_couperiana **
--+                                    
  |  +-------- Utterbackia             
  +--+                                 
     |  +----- Strophitus              
     +--+                              
        |  +-- Alasmidonta             
        +--+                           
           +-- Lasmigona

Krebs et al., 2003 [Publ 2691], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	8	7	--

-- Utterbackiana_couperiana **

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	4	4	--

  +----- Utterbackia             
  |                              
--+----- Utterbackiana_couperiana **
  |                              
  |  +-- Alasmidonta             
  +--+                           
     +-- Lasmigona

Lydeard et al., 1996 [Publ 2656], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	4	4	--

-- Utterbackiana_couperiana **

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	0	0	--

-- Utterbackiana_implicata **

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