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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Brachyanodon Crosse & Fischer, 1894: cladomics
     5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis = 5 (mean = 1.38, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 5, (mean = 3.33 median = 3.5)
proportion of cladograms that support basic monophyly = 0.33

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Chong et al., 2008 [Publ 2668], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
41810714320
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana **   
  +--------------+                              
  |              +-- Brachyanodon_californiensis **
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4 **
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3 **
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2 **
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1 **
              +--+                              
                 +-- Brachyanodon_kennerlyi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
58776275223
  +----------------------- Anodonta_anatina           
  |                                                   
--+  +-------------------- Pseudanodonta              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2 **
     |  |                                             
     |  |           +----- Anodonta_cygnea            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata        
           |                                          
           |  +----------- Brachyanodon_impura **        
           |  |                                       
           +--+  +-------- Brachyanodon **               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1 **
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2 **
                    |                                 
                    |  +-- Brachyanodon_californiensis **
                    +--+                              
                       +-- Brachyanodon_oregonensis__1 **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4754243120
           +-- Anodonta_exulcerata        
  +--------+                              
  |        +-- Anodonta_cygnea            
  |                                       
--+        +-- Brachyanodon_californiensis **
  |  +-----+                              
  |  |     +-- Brachyanodon_nuttalliana **   
  |  |                                    
  +--+     +-- Brachyanodon_kennerlyi **     
     |  +--+                              
     |  |  +-- Brachyanodon_oregonensis **   
     +--+                                 
        |  +-- Anodonta_anatina           
        +--+                              
           +-- Utterbackiana

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
320031020023
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana                 
  |                              
  +-- Brachyanodon_kennerlyi **     
  |                              
  +-- Brachyanodon_nuttalliana **   
  |                              
  +-- Brachyanodon_oregonensis **   
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2202300--
  +-- Brachyanodon_oregonensis **
--+                           
  +-- Brachyanodon_nuttalliana **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
22021500--
  +-- Brachyanodon_californiensis **
--+                              
  +-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015242--
-- Brachyanodon_nuttalliana **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
11015142--
-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113832--
  +----- Pseudanodonta           
--+                              
  |  +-- Anodonta                
  +--+                           
     +-- Brachyanodon_nuttalliana **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
110110111--
  +-- Anodonta                
--+                           
  +-- Brachyanodon_oregonensis **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
11011511--
-- Brachyanodon_kennerlyi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112132--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Brachyanodon_nuttalliana **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017442--
-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
11012700--
-- Brachyanodon_nuttalliana **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019442--
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119832--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019042--
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110120032--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11018032--
  +----- Pseudanodonta           
  |                              
--+  +-- Anodonta_anatina        
  |  |                           
  +--+-- Anodonta_cygnea         
     |                           
     +-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112432--
-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
11011511--
-- Brachyanodon_kennerlyi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110113942--
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011842--
-- Brachyanodon_nuttalliana **

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1101400--
-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
11012800--
-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019042--
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110120032--
-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113832--
  +-- Anodonta                
--+                           
  +-- Brachyanodon_nuttalliana **

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
11014032--
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017532--
  +----- Pseudanodonta           
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Anodonta

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
110110111--
-- Brachyanodon_oregonensis **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
11012500--
-- Brachyanodon_nuttalliana **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019442--
  +----- Anodonta                
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Pseudanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017532--
  +-------- Pseudanodonta           
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Brachyanodon_nuttalliana **
     +--+                           
        +-- Anodonta_cygnea
 
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