The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina genus Brachyanodon Crosse & Fischer, 1894: cladomics
5 Recent valid species (5 total species) Cladomics Summarynumber of species = 5 (5 total)
analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis =
5 (mean = 1.38, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 6
maximum number of included terminals = 5, (mean =
3.33 median = 3.5)
proportion of cladograms that support basic monophyly
= 0.33
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Chong et al., 2008 [Publ 2668], fig. 3 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 18 | 10 | 7 | 14 | 3 | 2 | 0 | +----------------- Anodonta_cygnea
|
+----------------- Pyganodon
|
+----------------- Alasminota
|
+----------------- Anodonta_anatina
|
+----------------- Unio
|
| +-- Brachyanodon_nuttalliana **
+--------------+
| +-- Brachyanodon_californiensis **
|
--+ +-- Beringiana
+--------------+
| +-- Sinanodonta
|
| +-- Pseudodontoideus
| |
+--------------+-- Alasmidonta
| |
| +-- Pterosyna
|
| +-------------- Pseudanodonta
| |
| | +----- Strophitus
| +--------+
+--+ | +-- Prolasmidonta
| +--+
| +-- Platynaias
|
| +----------- Brachyanodon_oregonensis__4 **
+--+
| +-------- Brachyanodon_oregonensis__3 **
+--+
| +----- Brachyanodon_oregonensis__2 **
+--+
| +-- Brachyanodon_oregonensis__1 **
+--+
+-- Brachyanodon_kennerlyi **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 8 | 7 | 7 | 627 | 5 | 2 | 23 | +----------------------- Anodonta_anatina
|
--+ +-------------------- Pseudanodonta
| |
+--+ +----------------- Brachyanodon_oregonensis__2 **
| |
| | +----- Anodonta_cygnea
+--+ +--------+
| | | +-- Anodonta_seddonae
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----------- Brachyanodon_impura **
| |
+--+ +-------- Brachyanodon **
| |
+--+ +----- Brachyanodon_nuttalliana__1 **
| |
+--+----- Brachyanodon_nuttalliana__2 **
|
| +-- Brachyanodon_californiensis **
+--+
+-- Brachyanodon_oregonensis__1 **
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 7 | 5 | 4 | 24 | 3 | 1 | 20 | +-- Anodonta_exulcerata
+--------+
| +-- Anodonta_cygnea
|
--+ +-- Brachyanodon_californiensis **
| +-----+
| | +-- Brachyanodon_nuttalliana **
| |
+--+ +-- Brachyanodon_kennerlyi **
| +--+
| | +-- Brachyanodon_oregonensis **
+--+
| +-- Anodonta_anatina
+--+
+-- Utterbackiana
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 20 | 0 | 3 | 102 | 0 | 0 | 23 | +-- Alasmidonta_marginata
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata
|
+-- Beringiana
|
+-- Brachyanodon_kennerlyi **
|
+-- Brachyanodon_nuttalliana **
|
+-- Brachyanodon_oregonensis **
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Pterosyna
|
--+-- Platynaias_compressa
|
+-- Lasmigona
|
+-- Platynaias_decorata
|
+-- Platynaias_subviridis
|
+-- Pyganodon_grandis
|
+-- Pyganodon_lacustris
|
+-- Pseudodontoideus
|
+-- Strophitus
|
+-- Utterbackia_imbecillis
|
+-- Utterbackia_peninsularis__1
|
+-- Utterbackia_peninsularis__2
Wu et al., 2026 [Publ 4811], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 0 | 0 | -- | +-- Brachyanodon_californiensis **
--+
+-- Brachyanodon_nuttalliana **
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 3 | 0 | 0 | -- | +-- Brachyanodon_oregonensis **
--+
+-- Brachyanodon_nuttalliana **
Zanatta et al., 2007 [Publ 2698], fig. 1B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | -- Brachyanodon_kennerlyi **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- | +-- Anodonta
--+
+-- Brachyanodon_oregonensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 3 | 2 | -- | +----- Anodonta_cygnea
--+
| +-- Anodonta_anatina
+--+
+-- Brachyanodon_nuttalliana **
Hoeh, 1990 [Publ 2717], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | -- Brachyanodon_kennerlyi **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 140 | 4 | 2 | -- | +-- Brachyanodon_nuttalliana **
--+
+-- Pseudanodonta
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 4 | 2 | -- | +----- Anodonta
--+
| +-- Brachyanodon_nuttalliana **
+--+
+-- Pseudanodonta
Cyr et al., 2007 [Publ 2745], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | -- Brachyanodon_nuttalliana **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- | +-- Anodonta
--+
+-- Brachyanodon_nuttalliana **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | -- Brachyanodon_nuttalliana **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- | +----- Pseudanodonta
--+
| +-- Anodonta
+--+
+-- Brachyanodon_nuttalliana **
Hoeh, 1990 [Publ 2717], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | -- Brachyanodon_kennerlyi **
Hoeh, 1990 [Publ 2717], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | -- Brachyanodon_kennerlyi **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 3 | 2 | -- | +----- Anodonta_anatina
--+
| +-- Anodonta_cygnea
+--+
+-- Brachyanodon_nuttalliana **
Zanatta et al., 2007 [Publ 2698], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | -- Brachyanodon_kennerlyi **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 0 | 0 | -- | -- Brachyanodon_nuttalliana **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- | -- Brachyanodon_nuttalliana **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 3 | 2 | -- | +----- Pseudanodonta
--+
| +-- Brachyanodon_nuttalliana **
+--+
+-- Anodonta
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 3 | 2 | -- | +----- Pseudanodonta
|
--+ +-- Anodonta_anatina
| |
+--+-- Anodonta_cygnea
|
+-- Brachyanodon_nuttalliana **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 3 | 2 | -- | +----- Anodonta_anatina
--+
| +-- Anodonta_cygnea
+--+
+-- Brachyanodon_nuttalliana **
Whelan et al., 2023 [Publ 4287], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 3 | 2 | -- | +-- Brachyanodon_nuttalliana **
--+
+-- Pseudanodonta
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 2 | -- | -- Brachyanodon_nuttalliana **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 3 | 2 | -- | +-------- Pseudanodonta
--+
| +----- Anodonta_anatina
+--+
| +-- Brachyanodon_nuttalliana **
+--+
+-- Anodonta_cygnea
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 123 | 3 | 2 | -- | -- Brachyanodon_nuttalliana **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 3 | 2 | -- | -- Brachyanodon_nuttalliana **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- | -- Brachyanodon_oregonensis **
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