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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Brachyanodon Crosse & Fischer, 1894: cladomics
     5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis = 5 (mean = 1.38, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 5, (mean = 3.33 median = 3.5)
proportion of cladograms that support basic monophyly = 0.33

Chong et al., 2008 [Publ 2668], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
41810714320
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana **   
  +--------------+                              
  |              +-- Brachyanodon_californiensis **
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4 **
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3 **
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2 **
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1 **
              +--+                              
                 +-- Brachyanodon_kennerlyi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
58776265223
  +----------------------- Anodonta_anatina           
  |                                                   
--+  +-------------------- Pseudanodonta              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2 **
     |  |                                             
     |  |           +----- Anodonta_cygnea            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata        
           |                                          
           |  +----------- Brachyanodon_impura **        
           |  |                                       
           +--+  +-------- Brachyanodon **               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1 **
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2 **
                    |                                 
                    |  +-- Brachyanodon_californiensis **
                    +--+                              
                       +-- Brachyanodon_oregonensis__1 **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4754243120
           +-- Anodonta_exulcerata        
  +--------+                              
  |        +-- Anodonta_cygnea            
  |                                       
--+        +-- Brachyanodon_californiensis **
  |  +-----+                              
  |  |     +-- Brachyanodon_nuttalliana **   
  |  |                                    
  +--+     +-- Brachyanodon_kennerlyi **     
     |  +--+                              
     |  |  +-- Brachyanodon_oregonensis **   
     +--+                                 
        |  +-- Anodonta_anatina           
        +--+                              
           +-- Utterbackiana

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
320031020023
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana                 
  |                              
  +-- Brachyanodon_kennerlyi **     
  |                              
  +-- Brachyanodon_nuttalliana **   
  |                              
  +-- Brachyanodon_oregonensis **   
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2202300--
  +-- Brachyanodon_oregonensis **
--+                           
  +-- Brachyanodon_nuttalliana **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
22021500--
  +-- Brachyanodon_californiensis **
--+                              
  +-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112332--
-- Brachyanodon_nuttalliana **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011842--
-- Brachyanodon_nuttalliana **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017532--
  +-------- Pseudanodonta           
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Brachyanodon_nuttalliana **
     +--+                           
        +-- Anodonta_cygnea

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
11012800--
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119932--
-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
11014032--
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1101400--
-- Brachyanodon_nuttalliana **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110114042--
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017532--
  +----- Pseudanodonta           
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Anodonta

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
11012500--
-- Brachyanodon_nuttalliana **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019542--
  +----- Anodonta                
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Pseudanodonta

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119732--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112132--
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Brachyanodon_nuttalliana **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
110110111--
-- Brachyanodon_oregonensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113732--
  +----- Pseudanodonta           
--+                              
  |  +-- Anodonta                
  +--+                           
     +-- Brachyanodon_nuttalliana **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
11011511--
-- Brachyanodon_kennerlyi **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
110110111--
  +-- Anodonta                
--+                           
  +-- Brachyanodon_oregonensis **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019542--
-- Brachyanodon_nuttalliana **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017542--
-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019142--
-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
11012700--
-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019142--
-- Brachyanodon_nuttalliana **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
11011511--
-- Brachyanodon_kennerlyi **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119932--
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113732--
  +-- Anodonta                
--+                           
  +-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017932--
  +----- Pseudanodonta           
  |                              
--+  +-- Anodonta_anatina        
  |  |                           
  +--+-- Anodonta_cygnea         
     |                           
     +-- Brachyanodon_nuttalliana **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
11015242--
-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015342--
-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
11011411--
-- Brachyanodon_kennerlyi **
 
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