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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Bivalvia | ... | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pilsbryoconcha Simpson, 1900: cladomics
     9 Recent valid species (9 total species)

Cladomics Summary

number of species = 9 (9 total)
analyzed in 77 cladograms in 48 publications
maximum number of terminals in an analysis = 9 (mean = 1.94, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 26
maximum number of included terminals = 9, (mean = 3.77 median = 2)
proportion of cladograms that support basic monophyly = 0.85


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Bolotov et al., 2023 [Publ 4312], fig. S2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
99810322770
  +-------------------------------- Songkhlanaia                   
  |                                                                
  |                             +-- Nyeinchanconcha                
--+  +--------------------------+                                  
  |  |                          +-- Sundadontina                   
  |  |                                                             
  +--+                       +----- Namkongnaia                    
     |  +--------------------+                                     
     |  |                    |  +-- Thaiconcha                     
     |  |                    +--+                                  
     +--+                       +-- Bineurus                       
        |                                                          
        |  +----------------------- Pseudodon                      
        |  |                                                       
        +--+  +-------------------- Pilsbryoconcha_carinifera **      
           |  |                                                    
           +--+                 +-- Pilsbryoconcha_linguaeformis__2 **
              |  +--------------+                                  
              |  |              +-- Pilsbryoconcha **                 
              +--+                                                 
                 |  +-------------- Pilsbryoconcha_linguaeformis__1 **
                 |  |                                              
                 +--+           +-- Pilsbryoconcha_schomburgki **     
                    |  +--------+                                  
                    |  |        +-- Pilsbryoconcha_exilis **          
                    +--+                                           
                       |  +-------- Pilsbryoconcha_hoikaab **         
                       +--+                                        
                          |  +----- Pilsbryoconcha_mekongiana **      
                          +--+                                     
                             |  +-- Pilsbryoconcha_acuta **           
                             +--+                                  
                                +-- Pilsbryoconcha_kittitati **       

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
7768433198
        +-------------- Songkhlanaia              
        |                                         
        |        +----- Nyeinchanconcha           
  +-----+  +-----+                                
  |     |  |     |  +-- Thaiconcha                
  |     |  |     +--+                             
  |     +--+        +-- Bineurus                  
  |        |                                      
  |        |  +-------- Sundadontina              
  |        +--+                                   
  |           |  +----- Isannaia                  
--+           +--+                                
  |              |  +-- Namkongnaia               
  |              +--+                             
  |                 +-- Pseudodon                 
  |                                               
  |  +----------------- Lannanaia                 
  |  |                                            
  |  |              +-- Pilsbryoconcha_exilis__2 **  
  +--+  +-----------+                             
     |  |           +-- Pilsbryoconcha_schomburgki **
     |  |                                         
     +--+           +-- Pilsbryoconcha_carinifera ** 
        |  +--------+                             
        |  |        +-- Pilsbryoconcha_exilis__1 **  
        +--+                                      
           |  +-------- Pilsbryoconcha_hoikaab **    
           +--+                                   
              |  +----- Pilsbryoconcha_mekongiana ** 
              +--+                                
                 |  +-- Pilsbryoconcha_acuta **      
                 +--+                             
                    +-- Pilsbryoconcha_kittitati **  

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
88683126710
     +-------------- Songkhlanaia                
     |                                           
  +--+           +-- Namkongnaia                 
  |  |  +--------+                               
  |  |  |        +-- Pseudodon                   
  |  +--+                                        
  |     |  +-------- Sundadontina                
  |     +--+                                     
  |        |  +----- Nyeinchanconcha             
  |        +--+                                  
--+           |  +-- Thaiconcha                  
  |           +--+                               
  |              +-- Bineurus                    
  |                                              
  |              +-- Pilsbryoconcha_exilis **       
  |  +-----------+                               
  |  |           +-- Pilsbryoconcha_schomburgki **  
  |  |                                           
  +--+           +-- Pilsbryoconcha_carinifera **   
     |  +--------+                               
     |  |        +-- Pilsbryoconcha_linguaeformis **
     +--+                                        
        |  +-------- Pilsbryoconcha_acuta **        
        +--+                                     
           |  +----- Pilsbryoconcha_kittitati **    
           +--+                                  
              |  +-- Pilsbryoconcha_mekongiana **   
              +--+                               
                 +-- Pilsbryoconcha_hoikaab **      

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
88683126710
              +----- Nyeinchanconcha             
        +-----+                                  
        |     |  +-- Thaiconcha                  
        |     +--+                               
  +-----+        +-- Bineurus                    
  |     |                                        
  |     |  +-------- Songkhlanaia                
  |     +--+                                     
  |        |  +----- Sundadontina                
  |        +--+                                  
  |           |  +-- Namkongnaia                 
--+           +--+                               
  |              +-- Pseudodon                   
  |                                              
  |              +-- Pilsbryoconcha_exilis **       
  |  +-----------+                               
  |  |           +-- Pilsbryoconcha_schomburgki **  
  |  |                                           
  +--+           +-- Pilsbryoconcha_carinifera **   
     |  +--------+                               
     |  |        +-- Pilsbryoconcha_linguaeformis **
     +--+                                        
        |  +-------- Pilsbryoconcha_kittitati **    
        +--+                                     
           |  +----- Pilsbryoconcha_acuta **        
           +--+                                  
              |  +-- Pilsbryoconcha_hoikaab **      
              +--+                               
                 +-- Pilsbryoconcha_mekongiana **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
886868766
                    +-- Nyeinchanconcha             
                 +--+                               
                 |  +-- Sundadontina                
  +--------------+                                  
  |              |  +-- Thaiconcha                  
  |              +--+                               
  |                 +-- Bineurus                    
--+                                                 
  |                 +-- Namkongnaia                 
  |  +--------------+                               
  |  |              +-- Pseudodon                   
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
886868766
                    +-- Nyeinchanconcha             
                 +--+                               
                 |  +-- Sundadontina                
  +--------------+                                  
  |              |  +-- Thaiconcha                  
  |              +--+                               
  |                 +-- Bineurus                    
--+                                                 
  |                 +-- Namkongnaia                 
  |  +--------------+                               
  |  |              +-- Pseudodon                   
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **    

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
77685128910
  +----------------------- Pseudodon                 
  |                                                  
  |              +-------- Sundadontina              
  |  +-----------+                                   
--+  |           |  +----- Nyeinchanconcha           
  |  |           +--+                                
  |  |              |  +-- Thaiconcha                
  |  |              +--+                             
  |  |                 +-- Bineurus                  
  +--+                                               
     |              +----- Namkongnaia               
     |  +-----------+                                
     |  |           |  +-- Lannanaia                 
     |  |           +--+                             
     |  |              +-- Songkhlanaia              
     +--+                                            
        |  +-------------- Isannaia                  
        |  |                                         
        |  |           +-- Pilsbryoconcha_exilis__2 **  
        +--+  +--------+                             
           |  |        +-- Pilsbryoconcha_schomburgki **
           |  |                                      
           +--+        +-- Pilsbryoconcha_exilis__1 **  
              |  +-----+                             
              |  |     +-- Pilsbryoconcha_carinifera ** 
              |  |                                   
              +--+     +-- Pilsbryoconcha_acuta **      
                 |  +--+                             
                 |  |  +-- Pilsbryoconcha_kittitati **  
                 +--+                                
                    |  +-- Pilsbryoconcha_hoikaab **    
                    +--+                             
                       +-- Pilsbryoconcha_mekongiana ** 

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
55467721712
                    +-- Pilsbryoconcha_schomburgki **  
           +--------+                               
           |        +-- Pilsbryoconcha_exilis__1 **    
  +--------+                                        
  |        |  +-------- Pilsbryoconcha **              
  |        +--+                                     
  |           |  +----- Pilsbryoconcha_linguaeformis **
  |           +--+                                  
--+              |  +-- Pilsbryoconcha_exilis__2 **    
  |              +--+                               
  |                 +-- Pilsbryoconcha_carinifera **   
  |                                                 
  |  +----------------- Songkhlanaia                
  +--+                                              
     |  +-------------- Namkongnaia                 
     +--+                                           
        |  +----------- Pseudodon                   
        +--+                                        
           |  +-------- Sundadontina                
           +--+                                     
              |  +----- Nyeinchanconcha             
              +--+                                  
                 |  +-- Thaiconcha                  
                 +--+                               
                    +-- Bineurus                    

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3313331569
           +----- Pilsbryoconcha_exilis **       
  +--------+                                  
  |        |  +-- Pilsbryoconcha_linguaeformis **
  |        +--+                               
  |           +-- Pilsbryoconcha **              
  |                                           
  |           +-- Nyeinchanconcha             
--+        +--+                               
  |        |  +-- Sundadontina_cumingii       
  |  +-----+                                  
  |  |     |  +-- Thaiconcha                  
  |  |     +--+                               
  |  |        +-- Bineurus                    
  +--+                                        
     |        +-- Namkongnaia                 
     |  +-----+                               
     |  |     +-- Pseudodon                   
     +--+                                     
        |  +----- Sundadontina_tumida         
        +--+                                  
           |  +-- Sundadontina_brandti        
           +--+                               
              +-- Sundadontina_taskaevi       

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3313331569
        +----- Pilsbryoconcha_exilis **       
  +-----+                                  
  |     |  +-- Pilsbryoconcha_linguaeformis **
  |     +--+                               
  |        +-- Pilsbryoconcha **              
--+                                        
  |     +----- Nyeinchanconcha             
  |  +--+                                  
  |  |  |  +-- Thaiconcha                  
  |  |  +--+                               
  +--+     +-- Bineurus                    
     |                                     
     |  +----- Sundadontina                
     +--+                                  
        |  +-- Namkongnaia                 
        +--+                               
           +-- Pseudodon                   

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2204137105--
     +-------- Pilsbryoconcha_exilis__1 ** 
  +--+                                  
  |  |  +----- Pilsbryoconcha_exilis__3 ** 
  |  +--+                               
  |     |  +-- Pilsbryoconcha_exilis__2 ** 
  |     +--+                            
--+        +-- Pilsbryoconcha_carinifera **
  |                                     
  |        +-- Sundadontina_tumida      
  |  +-----+                            
  |  |     +-- Pseudodon                
  |  |                                  
  +--+     +-- Nyeinchanconcha          
     |  +--+                            
     |  |  +-- Sundadontina_cumingii    
     +--+                               
        |  +-- Thaiconcha               
        +--+                            
           +-- Bineurus                 

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
230289119
  +----- Pilsbryoconcha_exilis **    
--+                               
  |  +-- Pseudodon                
  +--+                            
     +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2204137115--
  +-------- Pilsbryoconcha_exilis__3 ** 
--+                                  
  |  +----- Pilsbryoconcha_exilis__2 ** 
  +--+                               
     |  +-- Pilsbryoconcha_exilis__1 ** 
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2203197105--
  +-------- Nyeinchanconcha          
  |                                  
  +-------- Thaiconcha               
  |                                  
  +-------- Sundadontina_cumingii    
  |                                  
  +-------- Sundadontina_tumida__1   
--+                                  
  +-------- Sundadontina_tumida__2   
  |                                  
  +-------- Pseudodon                
  |                                  
  +-------- Bineurus                 
  |                                  
  |  +----- Pilsbryoconcha_exilis__2 ** 
  +--+                               
     |  +-- Pilsbryoconcha_carinifera **
     +--+                            
        +-- Pilsbryoconcha_exilis__1 ** 

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2204197105--
     +-------- Pilsbryoconcha_exilis__3 ** 
  +--+                                  
  |  |  +----- Pilsbryoconcha_exilis__2 ** 
  |  +--+                               
  |     |  +-- Pilsbryoconcha_exilis__1 ** 
  |     +--+                            
--+        +-- Pilsbryoconcha_carinifera **
  |                                     
  |        +-- Pseudodon                
  |  +-----+                            
  |  |     +-- Sundadontina             
  +--+                                  
     |  +----- Nyeinchanconcha          
     +--+                               
        |  +-- Thaiconcha               
        +--+                            
           +-- Bineurus                 

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
220243125--
           +-- Nyeinchanconcha          
  +--------+                            
  |        +-- Bineurus                 
  |                                     
--+        +-- Thaiconcha               
  |  +-----+                            
  |  |     +-- Sundadontina             
  +--+                                  
     |  +----- Pseudodon                
     +--+                               
        |  +-- Pilsbryoconcha_carinifera **
        +--+                            
           +-- Pilsbryoconcha_exilis **    

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
2802506214
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
2203116155--
  +----------------- Bineurus                
  |                                          
--+  +-------------- Nyeinchanconcha         
  |  |                                       
  +--+           +-- Pseudodon               
     |  +--------+                           
     |  |        +-- Sundadontina            
     +--+                                    
        |  +-------- Thaiconcha              
        +--+                                 
           |  +----- Pilsbryoconcha_exilis__2 **
           +--+                              
              |  +-- Pilsbryoconcha_exilis__1 **
              +--+                           
                 +-- Pilsbryoconcha **          

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2204123105--
  +-------------- Bineurus                 
  |                                        
  |           +-- Nyeinchanconcha          
  +-----------+                            
  |           +-- Sundadontina_cumingii    
--+                                        
  |           +-- Sundadontina_tumida      
  +-----------+                            
  |           +-- Pseudodon                
  |                                        
  |  +----------- Thaiconcha               
  +--+                                     
     |  +-------- Pilsbryoconcha_exilis__3 ** 
     +--+                                  
        |  +----- Pilsbryoconcha_exilis__2 ** 
        +--+                               
           |  +-- Pilsbryoconcha_exilis__1 ** 
           +--+                            
              +-- Pilsbryoconcha_carinifera **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2204195105--
     +-------- Pilsbryoconcha_exilis__3 ** 
  +--+                                  
  |  |  +----- Pilsbryoconcha_exilis__2 ** 
  |  +--+                               
  |     |  +-- Pilsbryoconcha_carinifera **
  |     +--+                            
--+        +-- Pilsbryoconcha_exilis__1 ** 
  |                                     
  |        +-- Sundadontina_tumida      
  |  +-----+                            
  |  |     +-- Pseudodon                
  |  |                                  
  +--+     +-- Nyeinchanconcha          
     |  +--+                            
     |  |  +-- Sundadontina_cumingii    
     +--+                               
        |  +-- Thaiconcha               
        +--+                            
           +-- Bineurus                 

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
220243125--
        +-- Nyeinchanconcha          
  +-----+                            
  |     +-- Bineurus                 
  |                                  
  |     +-- Thaiconcha               
--+-----+                            
  |     +-- Sundadontina             
  |                                  
  |  +----- Pseudodon                
  +--+                               
     |  +-- Pilsbryoconcha_carinifera **
     +--+                            
        +-- Pilsbryoconcha_exilis **    

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2802816214
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
2203110155--
  +----------------- Bineurus                
  |                                          
--+  +-------------- Nyeinchanconcha         
  |  |                                       
  +--+           +-- Pseudodon               
     |  +--------+                           
     |  |        +-- Sundadontina            
     +--+                                    
        |  +-------- Thaiconcha              
        +--+                                 
           |  +----- Pilsbryoconcha_exilis__2 **
           +--+                              
              |  +-- Pilsbryoconcha_exilis__1 **
              +--+                           
                 +-- Pilsbryoconcha **          

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
22027833--
        +-- Pilsbryoconcha_exilis **
  +-----+                        
  |     +-- Pilsbryoconcha_acuta ** 
--+                              
  |  +----- Pseudodon            
  +--+                           
     |  +-- Bineurus             
     +--+                        
        +-- Nyeinchanconcha      

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
220211944--
        +-- Pilsbryoconcha_exilis **    
  +-----+                            
  |     +-- Pilsbryoconcha_carinifera **
--+                                  
  |  +----- Pseudodon                
  |  |                               
  +--+  +-- Nyeinchanconcha          
     |  |                            
     +--+-- Bineurus                 
        |                            
        +-- Sundadontina             

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
230289119
  +----- Pilsbryoconcha_exilis **    
--+                               
  |  +-- Pseudodon                
  +--+                            
     +-- Pilsbryoconcha_carinifera **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012211--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014311--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
110146105--
  +-------------- Pilsbryoconcha_exilis **
--+                                    
  |  +----------- Pseudodon            
  +--+                                 
     |  +-------- Sundadontina         
     +--+                              
        |  +----- Nyeinchanconcha      
        +--+                           
           |  +-- Bineurus             
           +--+                        
              +-- Thaiconcha           

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013511--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
11012322--
  +-- Pseudodon                
--+                            
  +-- Pilsbryoconcha_carinifera **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
11013000--
-- Pilsbryoconcha_carinifera **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015211--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
11013653--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Bineurus             
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Pseudodon            

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011811--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017211--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011711--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
11013900--
-- Pilsbryoconcha_exilis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015633--
  +-------- Bineurus             
--+                              
  |  +----- Pilsbryoconcha_exilis **
  +--+                           
     |  +-- Pseudodon            
     +--+                        
        +-- Sundadontina         

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11013054--
  +-------- Sundadontina             
  |                                  
--+     +-- Nyeinchanconcha          
  |  +--+                            
  |  |  +-- Bineurus                 
  +--+                               
     |  +-- Pilsbryoconcha_carinifera **
     +--+                            
        +-- Pseudodon                

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015311--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019411--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014511--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016711--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014611--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017211--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019011--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
11013653--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Pseudodon            
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Bineurus             

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
11013753--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Pseudodon            
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Bineurus             

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
110112106--
  +----------- Bineurus             
  |                                 
  |     +----- Namkongnaia          
--+  +--+                           
  |  |  |  +-- Sundadontina         
  |  |  +--+                        
  +--+     +-- Nyeinchanconcha      
     |                              
     |  +----- Pseudodon            
     +--+                           
        |  +-- Pilsbryoconcha_exilis **
        +--+                        
           +-- Thaiconcha           

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014574--
        +-- Pilsbryoconcha_exilis **
  +-----+                        
  |     +-- Pseudodon            
--+                              
  |  +----- Sundadontina         
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Bineurus             

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017211--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014511--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1101800--
-- Pilsbryoconcha_exilis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015633--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Bineurus             
  +--+                           
     |  +-- Pseudodon            
     +--+                        
        +-- Sundadontina         

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
120226186--
        +----- Pilsbryoconcha_exilis__2 **
  +-----+                              
  |     |  +-- Pilsbryoconcha_exilis__1 **
  |     +--+                           
  |        +-- Namkongnaia             
--+                                    
  |        +-- Pseudodon               
  |  +-----+                           
  |  |     +-- Sundadontina            
  +--+                                 
     |  +----- Nyeinchanconcha         
     +--+                              
        |  +-- Thaiconcha              
        +--+                           
           +-- Bineurus                

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013954--
  +----------- Pilsbryoconcha_carinifera **
--+                                     
  |  +-------- Pseudodon                
  +--+                                  
     |  +----- Sundadontina             
     +--+                               
        |  +-- Nyeinchanconcha          
        +--+                            
           +-- Bineurus                 

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
110147105--
  +-------------- Pilsbryoconcha_exilis **
--+                                    
  |  +----------- Pseudodon            
  +--+                                 
     |  +-------- Sundadontina         
     +--+                              
        |  +----- Nyeinchanconcha      
        +--+                           
           |  +-- Bineurus             
           +--+                        
              +-- Thaiconcha           

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011811--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013164--
  +----------- Pilsbryoconcha_exilis **
--+                                 
  |  +-------- Pseudodon            
  +--+                              
     |  +----- Sundadontina         
     +--+                           
        |  +-- Thaiconcha           
        +--+                        
           +-- Bineurus             

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018711--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015111--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014611--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Namkongnaia          
--+                        
  +-- Pilsbryoconcha_exilis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014711--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
11013653--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Pseudodon            
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Bineurus             

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019011--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
11013653--
  +-------- Pilsbryoconcha_exilis **
  |                              
--+  +----- Bineurus_anodontinum 
  |  |                           
  +--+----- Bineurus_loeiensis   
     |                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Pseudodon            

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
11012711--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014311--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
11013653--
  +-------- Pilsbryoconcha_exilis **
--+                              
  |  +----- Pseudodon            
  +--+                           
     |  +-- Thaiconcha           
     +--+                        
        +-- Bineurus             

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019411--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013711--
  +-- Pseudodon            
--+                        
  +-- Pilsbryoconcha_exilis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015211--
  +-- Pilsbryoconcha_exilis **
--+                        
  +-- Pseudodon            
 
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