MUSSELpdb | valid gen. Ensidens cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Ensidens Frierson, 1911: cladomics
6 Recent valid species (6 total species)

Cladomics Summary

number of species = 6 (6 total)
analyzed in 47 cladograms in 33 publications
maximum number of terminals in an analysis = 6 (mean = 2.74, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 6, (mean = 4.9 median = 5)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Konopleva et al., 2019 [Publ 4041], fig. 3

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
6	12	0	7	28	6	2	0

  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_dugasti **     
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Hyriopsis_bialata    
  |                        
  +-- Hyriopsis_myersiana  
  |                        
--+-- Hyriopsis_desowitzi  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai     
  |                        
  +-- Ensidens_jaculus__1 **  
  |                        
  +-- Ensidens_jaculus__2 **  
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	625	11	2	11

                 +-- Ensidens_ingallsianus **  
           +-----+                          
           |     +-- Ensidens_telus **         
           |                                
  +--------+     +-- Ensidens_dugasti **       
  |        |  +--+                          
  |        |  |  +-- Ensidens_sagittarius **   
  |        +--+                             
  |           |  +-- Ensidens_jaculus **       
  |           +--+                          
--+              +-- Ensidens_spiculus **      
  |                                         
  |           +----- Rectidens              
  |  +--------+                             
  |  |        |  +-- Hyriopsis_delaportei   
  |  |        +--+                          
  |  |           +-- Hyriopsis_desowitzi    
  +--+                                      
     |  +----------- Hyriopsis_panhai       
     |  |                                   
     |  |        +-- Hyriopsis_bialata      
     +--+  +-----+                          
        |  |     +-- Hyriopsis_khoratensis  
        |  |                                
        +--+     +-- Hyriopsis_kratiensis   
           |  +--+                          
           |  |  +-- Hyriopsis_phuphaniensis
           +--+                             
              |  +-- Hyriopsis_myersiana    
              +--+                          
                 +-- Hyriopsis_sakhonensis

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	76	10	2	11

                       +-- Ensidens_ingallsianus **  
                 +-----+                          
                 |     +-- Ensidens_telus **         
                 |                                
  +--------------+     +-- Ensidens_jaculus **       
  |              |  +--+                          
  |              |  |  +-- Ensidens_spiculus **      
  |              +--+                             
  |                 |  +-- Ensidens_sagittarius **   
--+                 +--+                          
  |                    +-- Ensidens_dugasti **       
  |                                               
  |                    +-- Hyriopsis_desowitzi    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis
                    +--+                          
                       +-- Hyriopsis_kratiensis

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	111	6	2	11

     +----------- Hyriopsis_desowitzi  
     |                                 
  +--+  +-------- Rectidens            
  |  |  |                              
  |  +--+     +-- Hyriopsis_khoratensis
  |     |  +--+                        
  |     |  |  +-- Hyriopsis_bialata    
  |     +--+                           
  |        |  +-- Hyriopsis_myersiana  
--+        +--+                        
  |           +-- Hyriopsis_panhai     
  |                                    
  |           +-- Ensidens_telus **       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus **
  |     |                              
  +-----+     +-- Ensidens_jaculus **     
        |  +--+                        
        |  |  +-- Ensidens_spiculus **    
        +--+                           
           |  +-- Ensidens_dugasti **     
           +--+                        
              +-- Ensidens_sagittarius **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	105	6	2	11

     +-------------- Hyriopsis_desowitzi  
  +--+                                    
  |  |  +----------- Rectidens            
  |  +--+                                 
  |     |  +-------- Hyriopsis_panhai     
  |     +--+                              
  |        |  +----- Hyriopsis_myersiana  
  |        +--+                           
--+           |  +-- Hyriopsis_khoratensis
  |           +--+                        
  |              +-- Hyriopsis_bialata    
  |                                       
  |              +-- Ensidens_telus **       
  |        +-----+                        
  |        |     +-- Ensidens_ingallsianus **
  |        |                              
  +--------+     +-- Ensidens_dugasti **     
           |  +--+                        
           |  |  +-- Ensidens_sagittarius ** 
           +--+                           
              |  +-- Ensidens_spiculus **    
              +--+                        
                 +-- Ensidens_jaculus **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	23	7	2	11

              +-- Rectidens            
  +-----------+                        
  |           +-- Hyriopsis            
  |                                    
--+           +-- Ensidens_ingallsianus **
  |  +--------+                        
  |  |        +-- Ensidens_telus **       
  +--+                                 
     |  +-------- Ensidens_spiculus **    
     +--+                              
        |  +----- Ensidens_dugasti **     
        +--+                           
           |  +-- Ensidens_jaculus **     
           +--+                        
              +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	1	1	1	11

  +-------------- Rectidens            
  |                                    
--+           +-- Ensidens_telus **       
  |  +--------+                        
  |  |        +-- Ensidens_ingallsianus **
  +--+                                 
     |  +-------- Ensidens_dugasti **     
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	119	6	2	11

           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_jaculus **     
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_sagittarius ** 
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_myersiana  
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_khoratensis

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	46	6	2	11

              +-- Ensidens_ingallsianus **
        +-----+                        
        |     +-- Ensidens_telus **       
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_jaculus **     
--+        +--+                        
  |           +-- Ensidens_spiculus **    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Hyriopsis_khoratensis
           +--+                        
              +-- Hyriopsis_bialata

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	195	6	2	11

  +-------------- Hyriopsis_desowitzi  
  |                                    
  +-------------- Hyriopsis_myersiana  
  |                                    
  +-------------- Hyriopsis_panhai     
  |                                    
--+           +-- Hyriopsis_bialata    
  +-----------+                        
  |           +-- Hyriopsis_khoratensis
  |                                    
  |  +----------- Rectidens            
  |  |                                 
  +--+        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	193	6	2	11

              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
--+        +--+                        
  |           +-- Ensidens_jaculus **     
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis
        +--+                           
           |  +-- Hyriopsis_bialata    
           +--+                        
              +-- Hyriopsis_myersiana

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	133	6	2	11

              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
  |        +--+                        
--+           +-- Ensidens_jaculus **     
  |                                    
  |           +-- Hyriopsis_bialata    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis
  +--+                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi

Muanta et al., 2019 [Publ 3921], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	4	6	4	4	1	12

  +-------------- Hyriopsis              
  |                                      
--+           +-- Ensidens_ingallsianus **  
  |  +--------+                          
  |  |        +-- Ensidens_telus **         
  +--+                                   
     |  +-------- Ensidens_sagittarius__2 **
     +--+                                
        |  +----- Ensidens_jaculus **       
        +--+                             
           |  +-- Ensidens_sagittarius__1 **
           +--+                          
              +-- Ensidens_dugasti **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	8	0	5	77	3	1	5

  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_jaculus **     
  |                        
  +-- Hyriopsis_desowitzi  
--+                        
  +-- Hyriopsis_myersiana  
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	195	6	2	11

  +----------- Hyriopsis_desowitzi  
  |                                 
  +----------- Rectidens            
  |                                 
  |        +-- Hyriopsis_bialata    
  |        |                        
  |        +-- Hyriopsis_myersiana  
--+--------+                        
  |        +-- Hyriopsis_panhai     
  |        |                        
  |        +-- Hyriopsis_khoratensis
  |                                 
  |        +-- Ensidens_ingallsianus **
  |  +-----+                        
  |  |     +-- Ensidens_telus **       
  +--+                              
     |  +----- Ensidens_sagittarius ** 
     +--+                           
        |  +-- Ensidens_spiculus **    
        +--+                        
           +-- Ensidens_jaculus **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	133	6	2	11

           +----- Hyriopsis_panhai     
           |                           
  +--------+----- Hyriopsis_khoratensis
  |        |                           
  |        |  +-- Hyriopsis_myersiana  
  |        +--+                        
--+           +-- Hyriopsis_bialata    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  |  +----------- Rectidens            
  +--+                                 
     |        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	26	6	2	11

           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_sagittarius ** 
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_jaculus **     
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_myersiana

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	1	1	11

  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	88	1	1	11

  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	77	4	2	10

  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens            
  |                           
  |  +-- Hyriopsis_khoratensis
--+--+                        
  |  +-- Hyriopsis_myersiana  
  |                           
  |  +-- Ensidens_ingallsianus **
  |  |                        
  +--+-- Ensidens_telus **       
     |                        
     +-- Ensidens_jaculus **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	19	9	2	--

                 +-- Ensidens_ingallsianus **  
  +--------------+                          
  |              +-- Ensidens_jaculus **       
  |                                         
--+              +-- Hyriopsis_desowitzi    
  |  +-----------+                          
  |  |           +-- Hyriopsis_delaportei   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai       
           |                                
           |  +----- Hyriopsis_khoratensis  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis
              +--+                          
                 +-- Hyriopsis_sakhonensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	34	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	3	2	--

  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	1	1	--

  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Rectidens            
  +--+                        
     +-- Hyriopsis

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	6	2	2	--

  +-- Ensidens **    
  |               
  +-- Hyriopsis__1
--+               
  +-- Hyriopsis__2
  |               
  +-- Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	1	1	--

  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	0	0	--

-- Ensidens_telus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	2	2	--

  +-- Ensidens_ingallsianus **
  |                        
--+-- Hyriopsis            
  |                        
  +-- Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	1	1	--

  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	1	1	--

  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	1	1	--

-- Ensidens_sagittarius **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

  +-------- Ensidens_sagittarius **
--+                             
  |  +----- Hyriopsis_bialata   
  +--+                          
     |  +-- Hyriopsis_desowitzi 
     +--+                       
        +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	1	1	--

  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	4	2	--

  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Ensidens_ingallsianus **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Ensidens_sagittarius **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

-- Ensidens_sagittarius **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	58	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Zieritz et al., 2024 [Publ 4381], fig. 6A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	2	1	--

  +-- Ensidens_ingallsianus **
--+                        
  +-- Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	3	2	--

-- Ensidens_sagittarius **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	2	--

  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

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